treatments-xml/data/6B/BD/82/6BBD828793815E9E0CB61A1DB82C1F28.xml
2024-06-21 12:39:24 +02:00

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<document id="ABCC177BEF25B555289A74EF8CC650AF" ENCODING="UTF-8" ID-GBIF-Dataset="ab80b869-a307-4dab-a688-d654f3868af9" ModsDocAuthor="Eguchi, K." ModsDocID="22171" checkinTime="1243372496133" checkinUser="christiana" docAuthor="Eguchi, K." docDate="2008" docId="6BBD828793815E9E0CB61A1DB82C1F28" docLanguage="en" docName="22171" docOrigin="Zootaxa 1902" docSource="http://hdl.handle.net/10199/19085" docTitle="Pheidole smythiesii Forel" docType="treatment" docVersion="10" lastPageNumber="90" masterDocId="05AA04380484BD1D646413E1E2F24622" masterDocTitle="A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae)." masterLastPageNumber="118" masterPageNumber="1" pageNumber="87" updateTime="1701311757962" updateUser="plazi">
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<mods:title id="B475DFE1BF9BC2F4D70C1DDB03F6E236">A revision of Northern Vietnamese species of the ant genus Pheidole (Insecta: Hymenoptera: Formicidae: Myrmicinae).</mods:title>
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<mods:namePart id="0C0EA2700F93B1896E1EEF85ED323EB7">Eguchi, K.</mods:namePart>
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<treatment id="6BBD828793815E9E0CB61A1DB82C1F28" ID-DOI="http://doi.org/10.5281/zenodo.6230498" ID-GBIF-Taxon="100133935" ID-Zenodo-Dep="6230498" LSID="urn:lsid:plazi:treatment:6BBD828793815E9E0CB61A1DB82C1F28" httpUri="http://treatment.plazi.org/id/6BBD828793815E9E0CB61A1DB82C1F28" lastPageNumber="90" pageNumber="87">
<subSubSection id="E39C6E1066F2F1022762A2260A0423D1" type="nomenclature">
<paragraph id="F8CA923D9132E68255D9760EA26EAF88" pageNumber="87">
<taxonomicName id="D5B4D4F3B80FC77C506DAD8B72937D0D" ID-CoL="4FYLN" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="smythiesii">Pheidole smythiesii Forel</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="2DD6DE0468D7E8844E8025B123B1BB5A" type="description">
<paragraph id="B1F9D0E9D5FA5839CB8B242F6B0186F9" pageNumber="87">Figs. 24a-g</paragraph>
</subSubSection>
<subSubSection id="1E583DD3C53573ABAE096855F2EA47DC" type="reference_group">
<paragraph id="6D4788C4323ACFCA2FB137F40227B313" pageNumber="87">
<taxonomicName id="4776ABCE0111FC531E2DF3B38D1952BB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142586" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole (Ceratopheidole) smythiesii Forel" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="smythiesii" subGenus="Ceratopheidole">Pheidole (Ceratopheidole) smythiesii Forel</taxonomicName>
, 1902: 165, 185. Syntypes: 2 majors, 3 minors &amp; 3 males, &quot;Assam (Smythies) LXVII 8, 17 et 11.&quot; [Assam, India], MHNG, examined; 1 major, &quot;Assam (Smythies) LXVII 10&quot;, MHNG, examined; 1 queen, &quot;Assam Smythies XCV 4&quot;, MHNG, examined.
</paragraph>
</subSubSection>
<subSubSection id="C2210F966BB8BE364C453A6C3A89692D" type="discussion">
<paragraph id="92FD74768993E2168A652BA00A0A0827" pageNumber="87">
<taxonomicName id="E43566C3B516BA0EE204E0EA9A4A5070" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="smythiesii">Pheidole smythiesii Forel</taxonomicName>
. Bolton 1995: 330 (revived combination in
<taxonomicName id="25F5C36066A45A2FDD435547E9B4CE18" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
), Bui &amp; Eguchi 2003: 9 (checklist), Eguchi 2003: 335-336 (redescription of male), Eguchi, Bui et al. 2005: 90 (checklist).
</paragraph>
</subSubSection>
<subSubSection id="7CBACDBA5781DBCD9BDBD25E0E7B595B" type="reference_group">
<paragraph id="65714D6A907FF5A761880E397F011953" pageNumber="87">
<taxonomicName id="F84E68F298BB09079F19785C8FB6BEF1" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:244660" class="Insecta" family="Formicidae" genus="Ceratopheidole" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Ceratopheidole smythiesii" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="smythiesii">Ceratopheidole smythiesii Forel</taxonomicName>
. Emery 1922: 113.
</paragraph>
<paragraph id="5C9B4C657B725F04507A55AAED7FE138" pageNumber="87">
<taxonomicName id="CA6D46BB4DD2D771A60410253A61EBAD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:142585" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole (Ceratopheidole) smythiesii var. bengalensis Forel" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="variety" species="smythiesii" subGenus="Ceratopheidole" variety="bengalensis">Pheidole (Ceratopheidole) smythiesii var. bengalensis Forel</taxonomicName>
, 1902: 186.
<taxonomicNameLabel id="773F65AFB106D08593A5EC49FDCDE333">Syn.n.</taxonomicNameLabel>
Syntype: 1 minor, Bengal, India, MHNG, examined.
</paragraph>
<paragraph id="2A81C9A3C312AB956E6ACF72F41EBBEF" pageNumber="87">
<taxonomicName id="4953E5C9462C2F5E33B0A9166F002B3B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="smythiesii">Pheidole smythiesii subsp. bengalensis Forel</taxonomicName>
. Bolton 1995: 318 (revived combination).
<taxonomicNameLabel id="8E7969A9837D54E64F17DBD1EBD822B2">Syn.n.</taxonomicNameLabel>
<taxonomicName id="E382D0F7AE634BEFF41E43444512614B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:244661" class="Insecta" family="Formicidae" genus="Ceratopheidole" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Ceratopheidole smythiesii var. bengalensis" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="variety" species="smythiesii" variety="bengalensis">Ceratopheidole smythiesii var. bengalensis Forel</taxonomicName>
. Emery 1922e: 113.
<taxonomicNameLabel id="101FD6BBCD1BC400607AB3CD2370C790">Syn.n.</taxonomicNameLabel>
</paragraph>
<paragraph id="34B548D56BE7C7009C1C110C94465BDC" pageNumber="87">
<taxonomicName id="451FF8CC9FA3B2668BF05844E9AB2BBE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33616" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole bhavanae Bingham" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="bhavanae">Pheidole bhavanae Bingham</taxonomicName>
, 1903: 228-229. Bolton 1995: 318 (revived combination in
<taxonomicName id="F8A1C6F51CD51BFB6C4B3B8A499B775F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
).
<taxonomicNameLabel id="E41504F06826AF3608102AC79117D407">Syn.n.</taxonomicNameLabel>
Syntypes: 1 major, &quot;Sikkim Darjiling Senchal 8000ft 4. 1900 Bingham&quot; [India; according to the original description Rogers collected the material], BMNH, examined; original description also included syntype minor(s) from the same locality, not examined.
</paragraph>
<paragraph id="B34AAC76655A50234189E796702C32B5" pageNumber="87">
<taxonomicName id="B6942779C9B710E8C1F300EEE3F9F7C4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:244662" class="Insecta" family="Formicidae" genus="Ceratopheidole" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName-HNS="Ceratopheidole bhavanae" order="Hymenoptera" pageNumber="87" phylum="Arthropoda" rank="species" species="bhavanae">Ceratopheidole bhavanae Bingham</taxonomicName>
. Emery 1922e: 113.
<taxonomicNameLabel id="3994A69F36DCC6D3E4BDD3A173A290AD">Syn.n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="7B3930B0E5C46169B69B94E36C3D5FCE" type="materials_examined">
<paragraph id="46C86879F422BAB3079DEA89DF98BE7B" pageNumber="87">Other material examined: S. China: Guizhou: Huajiang near Sha Dong, Guanling [L. Latella]; Guangxi: Mao Er Shan, 690 m alt. [Eg00-GNGX-040, -041, -042]. Vietnam: Vinh Phuc: Tam Dao N.P., 900-1240 m alt. [VN98-SKY-13; VN98-HO-011]; Ha Tay (misspelled as &quot;Ha Tai&quot;): Ba Vi N.P., 21°03'N, 105°22'E, 800 m alt. [Eg02-VN-052]; Nghe An: Pu Hoat [T.V. Bui], Pu Hoat, ca. 750 m alt., primary forest [T.V. Bui's bottles: C, E, G, H, I]. Thailand: Chiang Mai: Doi Suthep-Pui N.P. [Sk. Yamane], Doi Suthep-Pui N.P., 800-900 m alt., dry evergreen forest [Eg01-TH-067, -068, -070, -075, -077, -094]. India: Himachal Pradesh: Riwalsar, 1300 m alt. [H. Bharti].</paragraph>
</subSubSection>
<subSubSection id="2D0679FB4A64766A1284AB252DE08C87" type="description">
<paragraph id="9F23396AF27FDBF66E7C4B7A1C097F33" pageNumber="87">Worker measurements &amp; indices: Major (n=5). - HL 2.63-3.03 mm; HW 2.63-3.09 mm; CI 95-102; SL 1.34-1.46 mm; SI 45-53; FL 2.26-2.39 mm; FI 74-86.</paragraph>
<paragraph id="A6618D21AFA6826AE5DA2023F9F1CE0C" pageNumber="87">Minor (n=5). - HL 0.95-1.19 mm; HW 0.82-1.03 mm; CI 86-91; SL 1.13-1.33 mm; SI 124-140; FL 1.38-1.67 mm; FI 156-168.</paragraph>
<caption id="AFF26E4A9410A015D5FEED2B40E10576">
<paragraph id="6D63DDD69F87A9CB091E0B96383A693C" pageNumber="88">
<pageStartToken id="D00306B6E681D55884F011DD8E6F3026" pageNumber="88">
<pageStartToken id="9B965BE41EA90F322C98CD25A6ECD093" pageNumber="88">FIGURE</pageStartToken>
</pageStartToken>
24a-d,
<taxonomicName id="A4A1C50546264865DF0B1266E4B3D34D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="88" phylum="Arthropoda" rank="species" species="smythiesii">Pheidole smythiesii</taxonomicName>
, major [T.V. Bui's bottle I] - a, head in full-face view; b, head in lateral view; c, mesosoma and waist in dorsal view; d, mesosoma and waist in lateral view.
</paragraph>
</caption>
<caption id="9772DA55F2C45D6254D5A94CE931D4EB">
<paragraph id="15E2C86D5E0EDA6951D0DA3FED429025" pageNumber="89">
<pageStartToken id="4B44D7C35360BDDD9A5AE9298AB001AF" pageNumber="89">
<pageStartToken id="828F61C33A94634A9DD102431FD9603F" pageNumber="89">FIGURE</pageStartToken>
</pageStartToken>
24e-g,
<taxonomicName id="CB5EA76BB17BE620547AD746A4F0C7E6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="89" phylum="Arthropoda" rank="species" species="smythiesii">Pheidole smythiesii</taxonomicName>
, minor [T.V. Bui's bottle I] - e, head in full-face view; f, mesosoma and waist in dorsal view; g, mesosoma and waist in lateral view.
</paragraph>
</caption>
<paragraph id="329E64274CEFEE18DCD514B1AFDED463" pageNumber="89">Worker description</paragraph>
<paragraph id="C5C96B5490675A63F06480A186FEB16F" lastPageNumber="90" pageNumber="89">Major. - Head in lateral view usually not impressed on vertex; frons and vertex longitudinally-obliquely rugose; dorsal and dorsolateral faces of vertexal lobe rugoso-recticulate, rugoso-punctate or almost smooth; frontal carina and antennal scrobe absent; median longitudinal carina of clypeus weak or inconspicuous; median process of hypostoma low or inconspicuous, rarely with a concavity in the center; submedian processes low or relatively well developed (but rarely very low); lateral processes conspicuous (but sometimes small); antenna with a 4-segmented club; maximal diameter of eye almost as long as or a little longer than antennal segment X. Promesonotal dome in dorsal view smooth to shagreened, usually with transverse rugulae, in lateral view usually with a conspicuous prominence, but sometimes only with an inconspicuous mound on its posterior slope; humerus not produced; the dome narrower at the humeri than at the bottom. Petiole as long as or shorter than postpetiole (excluding its helcium); postpetiole massive. First gastral tergite shagreened to smooth.</paragraph>
<paragraph id="C5CF2F92260C2A09FA13AE15D1A5A608" pageNumber="90">Minor. - Dorsum of head largely smooth or at most dimly punctured; preoccipital carina conspicuous dorsally and laterally; median part of clypeus smooth and shining; median longitudinal carina present at least anteriorly (but sometimes very weak); antenna with a 4-segmented club; scape extending far beyond posterolateral margin of head; maximal diameter of eye shorter than (but rarely as long as) antennal segment X. Promesonotal dome smooth or shagreened, sometimes with weak rugulae, in lateral view with a low mound on its posterior slope; humerus in dorso-oblique view not raised; mesopleuron, metapleuron and lateral face of propodeum weakly punctured or almost smooth. Petiole much shorter than postpetiole (excluding helcium); postpetiole massive.</paragraph>
<paragraph id="8F655E28DF0593C156E58D84815FD54F" pageNumber="90">
Recognition: There are no distinct differences between the syntypes of
<taxonomicName id="8D607F37ABFE557DA9DA76AE5156776B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="smythiesii">P. smythiesii</taxonomicName>
and
<taxonomicName id="BE5D85E85FF84EB4BF20FAECC817E3EA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151568" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii bengalensis Forel" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="subSpecies" species="smythiesii" subSpecies="bengalensis">P. smythiesii bengalensis Forel</taxonomicName>
. In Forel's original description (Forel 1902), he mentioned that the head of the minor is longer in the latter than that in the former. The cephalic index is, however, not so different between them: CI 0.86 in the latter and CI 0.88-0.89 in the former. Thus
<taxonomicName id="BB3DF1E3BF7AC654719522648D219125" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:151568" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii bengalensis Forel" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="subSpecies" species="smythiesii" subSpecies="bengalensis">P. smythiesii bengalensis</taxonomicName>
is merely a local population of
<taxonomicName id="B3A277DC60E4D7B2B73C66211ED52158" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="smythiesii">P. smythiesii</taxonomicName>
. The differences between
<taxonomicName id="CFA83481913C3B0E0571AB08275B29D9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="smythiesii">P. smythiesii</taxonomicName>
and
<taxonomicName id="ABD3A6AD426E76AA61D01A9E157B614E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33616" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole bhavanae Bingham" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="bhavanae">P. bhavanae</taxonomicName>
shown by Bingham (1903) seem to be inadequate for separating them at the species level, and in the course of examination of the type material I also could not find definitive evidence which supports Bingham's recognition of
<taxonomicName id="E427EDFB47FDC577B4C7F69BDE6B220E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33616" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole bhavanae Bingham" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="bhavanae">P. bhavanae</taxonomicName>
from
<taxonomicName id="E37D0AAB672E405432E9C45B0D8F6D82" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="smythiesii">P. smythiesii</taxonomicName>
. Thus I conclude that they are conspecific.
</paragraph>
</subSubSection>
<subSubSection id="BB66E5A9AB9C8E4BCA13A7C6008B0C8E" type="discussion">
<paragraph id="8E1AD8ED515BE4EBFE4E3022A32631A0" pageNumber="90">
In Vietnam only two species,
<taxonomicName id="01FBC3B9A557715F86E4837D8471625A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34015" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole smythiesii Forel" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="smythiesii">Pheidole smythiesii</taxonomicName>
and
<taxonomicName id="1A23CF6ADD7063E4F2CE40EAD2D06C49" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33753" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole gatesi (Wheeler)" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="gatesi">P. gatesi</taxonomicName>
, have 4-segmented antennal clubs, and they are morphologically very similar to each other. Differences between the two species are given under the remarks of
<taxonomicName id="C2837A2F30D4BDF8F75BBB533B413B31" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33753" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole gatesi (Wheeler)" order="Hymenoptera" pageNumber="90" phylum="Arthropoda" rank="species" species="gatesi">P. gatesi</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B8CBA328AD9F3BFED0D0BBCD4788A879" type="distribution">
<paragraph id="B3973E1ABA6D47EC8F3E97D7046BA0FC" pageNumber="90">Distribution &amp; bionomics: Known from the Indian and Indo-Chinese subregions. This species usually inhabits woody habitats and sometimes occurs in open lands, and nests in the soil. According to Alfred &amp; Agarwal (1990) workers tend aphid colonies (Micromyzus kalimponginsis) on Hedychium coronarium (Zingiberaceae) in Shillong, India.</paragraph>
</subSubSection>
</treatment>
</document>