365 lines
42 KiB
XML
365 lines
42 KiB
XML
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<mods:title id="C087F91E1742E1F14FA88F0CA11D7AA7">A new framework of the evolution of the ctenodactylids (Mammalia: Rodentia) in Asia: new species and phylogenetic status of distylomyins</mods:title>
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<mods:namePart id="0179617AE7592EA4FB5832985B9E69A9">Oliver, Adriana</mods:namePart>
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<mods:affiliation id="D1BB28BAAA98DEA804BAAD2D39C3FF5C">Department of Paleobiology, National Museum of Natural Sciences-Spanish National Research Council (MNCN-CSIC), C / José Gutiérrez Abascal 2, 28006 Madrid, Spain & Association Mujeres con los Pies en la Tierra, C / Jose Antonio Novais, 12, 28040 Madrid, Spain</mods:affiliation>
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<mods:namePart id="DAC1231C4267A9369C80BAF79196A5F5">Carro-Rodríguez, Patricia M.</mods:namePart>
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<mods:affiliation id="2EB8C437A8CA2DE3B951C87EB45655CB">Association Mujeres con los Pies en la Tierra, C / Jose Antonio Novais, 12, 28040 Madrid, Spain & Department of Geodynamics, Stratigraphy and Palaeontology (GEODESPAL), Universidad Complutense de Madrid, C / Jose Antonio Novais, 12, 28040, Madrid, Spain & Sedimentary Geology and Environmental Change, Geosciences Institute (CSIC, UCM), C / del Dr Severo Ochoa, 7, 28040, Madrid, Spain</mods:affiliation>
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<mods:affiliation id="2BE49249517714DAA4E73A8E5C34FE22">Geological-Paleontological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:date id="84EFE12C9F47713B2858BA413EF889F6">2023</mods:date>
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<taxonomicName id="24D5C2A9CA1D8930FED6FD17F77CFD24" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[276,518,704,731]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="taatsinius" status="sp. nov.">
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<emphasis id="D1A16538CA1D8930FED6FD17F77CFD24" bold="true" box="[276,518,704,731]" italics="true" pageId="7" pageNumber="640">Prodistylomys taatsinius</emphasis>
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<figureCitation id="7BEEA5AFCA1D8930FFB3FD30F5B8FD00" box="[113,194,743,767]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStartId-0="8.[131,196,1206,1230]" captionStartId-1="9.[113,178,930,954]" captionTargetBox-0="[139,1467,152,1169]" captionTargetBox-1="[121,1450,152,895]" captionTargetId-0="figure-294@8.[128,1478,144,1178]" captionTargetId-1="figure-110@9.[114,1458,144,903]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Figure 4. Prodistylomys taatsinius sp. nov. from the type locality of LOG-A/1, in Luugar Khudag, Valley of Lakes (Mongolia): (A) LOG-A/1 (NHMW 2012/0048/0019) M1/2 left; (B) LOG-A/1 (NHMW 2012/0048/0022) M3 right; (C) LOG-A/1 (NHMW 2012/0048/0003) m1 right; (D) LOG-A/1 (NHMW 2012/0048/001) from left to right: p4-m1-m2 right, holotype. All teeth are at the same magnification. Leftside specimens are underlined. Scale bar: 1 mm." captionText-1="Figure 5. Upper and lower cheek teeth of Prodistylomys taatsinius sp. nov. from the Valley of Lakes (Mongolia). From Unkheltseg:(A) UNCH-A/3 (NHMW 2012/0049/0003) P4 left; (B) UNCH-A/3 (NHMW 2012/0049/0005) M1/2 left; (C) UNCH-A/3 (NHMW 2012/0049/0001) m2 right; From Huch Teeg: (D) RHN-A/12 (NHMW 2012/0050/0001) DP4 right; (E) RHN-A/12 (NHMW 2012/0050/0003) M1/2 right; (F) RHN-A/12 (NHMW 2012/0050/0006) p4 right; (G) RHN-A/12 (NHMW 2012/0050/0002) m3 right. All teeth are at the same magnification.Left-side specimens are underlined.Scale bar:1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10469784" figureDoi-1="http://doi.org/10.5281/zenodo.10469786" httpUri-0="https://zenodo.org/record/10469784/files/figure.png" httpUri-1="https://zenodo.org/record/10469786/files/figure.png" pageId="7" pageNumber="640">Figs 4-5</figureCitation>
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; Supporting Information Model S1-2
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</paragraph>
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<emphasis id="D1A16538CA1D8930FFB3FCF6F42BFCC7" box="[113,337,801,825]" italics="true" pageId="7" pageNumber="640">Zoobank registration:</emphasis>
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urn:lsid:zoobank.org:pub:
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.
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<emphasis id="D1A16538CA1D8930FF4FFC88F582FC88" box="[141,248,863,887]" italics="true" pageId="7" pageNumber="640">Distylomys</emphasis>
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/
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<emphasis id="D1A16538CA1D8930FEC6FC88F4F1FC88" box="[260,395,863,887]" italics="true" pageId="7" pageNumber="640">Prodistylomys</emphasis>
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sp.
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(
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<bibRefCitation id="8744C4DBCA1D8930FE79FC88F7ABFC88" author="Hock V & Daxner-Hock G & Schmid HP" box="[443,721,863,887]" pageId="7" pageNumber="640" pagination="83 - 125" refId="ref16320" refString="Hock V, Daxner-Hock G, Schmid HP et al. Oligocene-Miocene sediments, fossils and basalts from the Valley of Lakes (Central Mongolia) - an integrated study. Mitteilungen der Oesterreichischen Geologischen Gesellschass 1999; 90: 83 - 125." type="journal article" year="1999">
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Höck
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<emphasis id="D1A16538CA1D8930FE3FFCB7F757FC88" box="[509,557,863,887]" italics="true" pageId="7" pageNumber="640">et al.</emphasis>
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1999: 118–119
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, fig. 21/1).
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<emphasis id="D1A16538CA1D8930FF55FC49F47EFC49" box="[151,260,926,950]" italics="true" pageId="7" pageNumber="640">Distylomys</emphasis>
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sp.
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(
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<treatmentCitation id="62749F3BCA1D8930FEFEFC49F78FFC49" author="Daxner-Hock G & Badamgarav D" box="[316,757,926,950]" page="16" pageId="7" pageNumber="640" year="2007">
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<bibRefCitation id="8744C4DBCA1D8930FEFEFC49F78FFC49" author="Daxner-Hock G & Badamgarav D" box="[316,757,926,950]" pageId="7" pageNumber="640" pagination="1 - 24" refId="ref15459" refString="Daxner-Hock G, Badamgarav D. Oligocene-Miocene vertebrates from the Valley of Lakes (Central Mongolia): morphology, phylogenetic and stratigraphic implications - 1. Geological and stratigraphic seuing. Annalen des Naturhistorischen Museums in Wien 201 2007; 108 A: 1 - 24." type="journal article" year="2007">Daxner-Höck and Badamgarav 2007: 16</bibRefCitation>
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, 18).
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<emphasis id="D1A16538CA1D8930FF50FC0AF463FC0A" box="[146,281,989,1013]" italics="true" pageId="7" pageNumber="640">Prodistylomys</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CA92D843CA1D8930FEDDFC0AF4F8FC0B" box="[287,386,989,1012]" pageId="7" pageNumber="640" rank="species">nov. spec.</taxonomicNameLabel>
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2 (
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<treatmentCitation id="62749F3BCA1D8930FE5DFC0AF7FFFC0A" author="Oliver A & Sanisidro O & Pelaez-Campomanes P" box="[415,645,989,1013]" page="61" pageId="7" pageNumber="640" year="2017">
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<bibRefCitation id="8744C4DBCA1D8930FE5DFC0AF7FFFC0A" author="Oliver A & Sanisidro O & Pelaez-Campomanes P" box="[415,645,989,1013]" pageId="7" pageNumber="640" pagination="372 - 9" refId="ref17898" refString="Oliver A, Sanisidro O, Pelaez-Campomanes P et al. Wear reconstruction of the teeth of Yindirtemys deflexus (Ctenodactylidae, Rodentia) and evolution of its dental pauern. Hist Biol 2017 a; 30: 372 - 9." type="journal article" year="2017">
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Oliver
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<emphasis id="D1A16538CA1D8930FE2AFC0AF76DFC0A" box="[488,535,989,1013]" italics="true" pageId="7" pageNumber="640">et al.</emphasis>
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2017a: 61
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).
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<paragraph id="E36AB92ACA1D8930FF4FFC2BF775FBCC" blockId="7.[113,763,801,1138]" pageId="7" pageNumber="640">
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<emphasis id="D1A16538CA1D8930FF4FFC2BF46EFBEB" box="[141,276,1020,1044]" italics="true" pageId="7" pageNumber="640">Prodistylomys</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CA92D843CA1D8930FEDFFC2BF4FFFBEC" box="[285,389,1020,1043]" pageId="7" pageNumber="640" rank="species">nov. spec.</taxonomicNameLabel>
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2 (
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<treatmentCitation id="62749F3BCA1D8930FE68FC2BF7CCFBEB" author="Daxner-Hock G & Badamgarav D & Barsbold D" box="[426,694,1020,1044]" page="121" pageId="7" pageNumber="640" year="2017">
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Daxner-Höck
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<emphasis id="D1A16538CA1D8930FD81FC2AF70FFBEB" box="[579,629,1020,1044]" italics="true" pageId="7" pageNumber="640">et al.</emphasis>
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2017
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a: 121, 127, figs 5, 47l,
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, 47n, 47o, tabs 5, 8).
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||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FF4FFBECF5BDFB8D" blockId="7.[113,763,801,1138]" pageId="7" pageNumber="640">
|
||
<treatmentCitationGroup id="C3C59E04CA1D8930FF4FFBECF5BDFB8D" pageId="7" pageNumber="640">
|
||
<taxonomicName id="24D5C2A9CA1D8930FF4FFBECF41BFBAD" baseAuthorityName="Oliver et al." baseAuthorityYear="2017" box="[141,353,1083,1107]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="tatsinii">
|
||
<emphasis id="D1A16538CA1D8930FF4FFBECF41BFBAD" box="[141,353,1083,1107]" italics="true" pageId="7" pageNumber="640">Prodistylomys tatsinii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<treatmentCitation id="62749F3BCA1D8930FEB3FBECF721FBAC" author="Oliver A & Sanisidro O & Baatarjav B" box="[369,603,1083,1107]" page="2" pageId="7" pageNumber="640" year="2017">
|
||
<bibRefCitation id="8744C4DBCA1D8930FEB3FBECF721FBAC" author="Oliver A & Sanisidro O & Baatarjav B" box="[369,603,1083,1107]" pageId="7" pageNumber="640" pagination="51 - 65" refId="ref17864" refString="Oliver A, Sanisidro O, Baatarjav B et al. Diversification rates in Ctenodactylidae (Rodentia, Mammalia) from Mongolia. Palaeobiodivers Palaeoenviron 2017 b; 97: 51 - 65." type="journal article" year="2017">
|
||
Oliver
|
||
<emphasis id="D1A16538CA1D8930FE7DFBECF488FBAC" box="[447,498,1083,1107]" italics="true" pageId="7" pageNumber="640">et al.</emphasis>
|
||
2017b: 2
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
, fig. 1A
|
||
<emphasis id="D1A16538CA1D8930FD7BFBECF5CCFB8D" italics="true" pageId="7" pageNumber="640">nomen nudum</emphasis>
|
||
).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABCFEAA1CA1D8930FFB3FB43F781FB54" box="[113,763,1171,1195]" pageId="7" pageNumber="640" type="etymology">
|
||
<paragraph id="E36AB92ACA1D8930FFB3FB43F781FB54" blockId="7.[113,763,1171,1195]" box="[113,763,1171,1195]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FFB3FB43F44BFB54" box="[113,305,1171,1195]" italics="true" pageId="7" pageNumber="640">Derivation of name:</emphasis>
|
||
Named for the region near the Taatsin River.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABCFEAA1CA1D8930FFB3FB1AF43BFA69" pageId="7" pageNumber="640" type="materials_examined">
|
||
<paragraph id="E36AB92ACA1D8930FFB3FB1AF483FAFB" blockId="7.[113,762,1229,1284]" pageId="7" pageNumber="640">
|
||
<materialsCitation id="53BDB377CA1D8930FFB3FB1AF483FAFB" collectionCode="NHMW" collectorName="Right & Model S" pageId="7" pageNumber="640" specimenCode="NHMW 2012" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="D1A16538CA1D8930FFB3FB1AF5ABFB1A" box="[113,209,1229,1253]" italics="true" pageId="7" pageNumber="640">
|
||
<typeStatus id="3C6E0788CA1D8930FFB3FB1AF5B6FB1A" box="[113,204,1229,1253]" pageId="7" pageNumber="640" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="B3731151CA1D8930FF19FB1AF409FB1A" box="[219,371,1229,1253]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="640" type="Museum">NHMW 2012</specimenCode>
|
||
/0048/0001,
|
||
<collectorName id="4E20DCFCCA1D8930FE3CFB1AF74DFB1A" box="[510,567,1229,1253]" pageId="7" pageNumber="640">Right</collectorName>
|
||
fragmentary mandible with p4-m2 (
|
||
<figureCitation id="7BEEA5AFCA1D8930FEF3FB3BF401FAFB" box="[305,379,1260,1284]" captionStart="Figure 4" captionStartId="8.[131,196,1206,1230]" captionTargetBox="[139,1467,152,1169]" captionTargetId="figure-294@8.[128,1478,144,1178]" captionTargetPageId="8" captionText="Figure 4. Prodistylomys taatsinius sp. nov. from the type locality of LOG-A/1, in Luugar Khudag, Valley of Lakes (Mongolia): (A) LOG-A/1 (NHMW 2012/0048/0019) M1/2 left; (B) LOG-A/1 (NHMW 2012/0048/0022) M3 right; (C) LOG-A/1 (NHMW 2012/0048/0003) m1 right; (D) LOG-A/1 (NHMW 2012/0048/001) from left to right: p4-m1-m2 right, holotype. All teeth are at the same magnification. Leftside specimens are underlined. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10469784" httpUri="https://zenodo.org/record/10469784/files/figure.png" pageId="7" pageNumber="640">Fig. 4D</figureCitation>
|
||
,
|
||
<collectorName id="4E20DCFCCA1D8930FE44FB3BF4A5FAFB" box="[390,479,1260,1284]" pageId="7" pageNumber="640">Model S</collectorName>
|
||
1).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FFB3FAF1F780FAC2" blockId="7.[113,762,1317,1341]" box="[113,762,1317,1341]" pageId="7" pageNumber="640">
|
||
<materialsCitation id="53BDB377CA1D8930FFB3FAF1F78CFAC2" box="[113,758,1317,1341]" collectionCode="NHMW" pageId="7" pageNumber="640" specimenCode="LOG-A/1" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="D1A16538CA1D8930FFB3FAF1F5B5FAC2" box="[113,207,1318,1341]" italics="true" pageId="7" pageNumber="640">
|
||
<typeStatus id="3C6E0788CA1D8930FFB3FAF1F5B3FAC2" box="[113,201,1318,1341]" pageId="7" pageNumber="640" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="B3731151CA1D8930FF15FAF2F438FAC2" box="[215,322,1317,1341]" collectionCode="LOG-A" pageId="7" pageNumber="640">LOG-A/1</specimenCode>
|
||
;
|
||
<collectionCode id="85C421EFCA1D8930FE88FAF1F4D8FAC2" box="[330,418,1318,1341]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="640" type="Museum">NHMW</collectionCode>
|
||
2012/0048/0002-24 (
|
||
<figureCitation id="7BEEA5AFCA1D8930FD4BFAF2F7C5FAC2" box="[649,703,1317,1341]" captionStart="Figure 4" captionStartId="8.[131,196,1206,1230]" captionTargetBox="[139,1467,152,1169]" captionTargetId="figure-294@8.[128,1478,144,1178]" captionTargetPageId="8" captionText="Figure 4. Prodistylomys taatsinius sp. nov. from the type locality of LOG-A/1, in Luugar Khudag, Valley of Lakes (Mongolia): (A) LOG-A/1 (NHMW 2012/0048/0019) M1/2 left; (B) LOG-A/1 (NHMW 2012/0048/0022) M3 right; (C) LOG-A/1 (NHMW 2012/0048/0003) m1 right; (D) LOG-A/1 (NHMW 2012/0048/001) from left to right: p4-m1-m2 right, holotype. All teeth are at the same magnification. Leftside specimens are underlined. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10469784" httpUri="https://zenodo.org/record/10469784/files/figure.png" pageId="7" pageNumber="640">Fig. 4</figureCitation>
|
||
A-C)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FFB3FA88F43BFA69" blockId="7.[113,762,1375,1430]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FFB3FA88F586FA88" box="[113,252,1375,1399]" italics="true" pageId="7" pageNumber="640">Type locality:</emphasis>
|
||
Luugar Khudag (
|
||
<collectionCode id="85C421EFCA1D8930FE05FA88F75CFA88" box="[455,550,1375,1399]" pageId="7" pageNumber="640">LOG-A</collectionCode>
|
||
/1), Valley of Lakes (Central Mongolia).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABCFEAA1CA1D8930FFB3FA6FF18CFF18" pageId="7" pageNumber="640" type="diagnosis">
|
||
<paragraph id="E36AB92ACA1D8930FFB3FA6FF476F993" blockId="7.[113,763,1463,1644]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FFB3FA6FF5ADFA30" box="[113,215,1464,1487]" italics="true" pageId="7" pageNumber="640">Diagnosis:</emphasis>
|
||
Small-sized species of
|
||
<taxonomicName id="24D5C2A9CA1D8930FE1DFA60F71CFA30" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[479,614,1463,1487]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="genus">
|
||
<emphasis id="D1A16538CA1D8930FE1DFA60F71CFA30" box="[479,614,1463,1487]" italics="true" pageId="7" pageNumber="640">Prodistylomys</emphasis>
|
||
</taxonomicName>
|
||
with slender lower molars. Lower p4 with metaflexid and posterolophid; subtriangular upper P4; upper M1 and M2 subsquare, and mesoflexus (lateral flexus) connects or almost connects to the hypoflexus (medial flexus); in the M3 the flexi are fused and form a fossette.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FFB3F95AF75BF91B" blockId="7.[113,763,1677,1983]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FFB3F95AF43DF95A" box="[113,327,1677,1701]" italics="true" pageId="7" pageNumber="640">Differential diagnosis:</emphasis>
|
||
Similar to
|
||
<taxonomicName id="24D5C2A9CA1D8930FE7EF959F764F95A" box="[444,542,1678,1701]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="wangae">
|
||
<emphasis id="D1A16538CA1D8930FE7EF959F764F95A" box="[444,542,1678,1701]" italics="true" pageId="7" pageNumber="640">P. wangae</emphasis>
|
||
</taxonomicName>
|
||
in size, but differs in having well-developed roots; M3 with a fossette; p4 with larger protoconid, metaflexid and posterolophid.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FF4FF93CF7E3F89E" blockId="7.[113,763,1677,1983]" pageId="7" pageNumber="640">
|
||
<taxonomicName id="24D5C2A9CA1D8930FF4FF93CF408F8FC" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[141,370,1771,1795]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="taatsinius" status="sp. nov.">
|
||
<emphasis id="D1A16538CA1D8930FF4FF93CF408F8FC" box="[141,370,1771,1795]" italics="true" pageId="7" pageNumber="640">Prodistylomys taatsinius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CA92D843CA1D8930FEB4F93BF4C5F8FC" box="[374,447,1772,1795]" pageId="7" pageNumber="640" rank="species">sp.nov.</taxonomicNameLabel>
|
||
has a similar size to
|
||
<taxonomicName id="24D5C2A9CA1D8930FDBDF93BF788F8FC" box="[639,754,1772,1795]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="mengensis">
|
||
<emphasis id="D1A16538CA1D8930FDBDF93BF788F8FC" box="[639,754,1772,1795]" italics="true" pageId="7" pageNumber="640">P.mengensis</emphasis>
|
||
</taxonomicName>
|
||
; however, differs in the simpler P4; DP4 with a deep furrow on lingual and labial walls; upper molars with lingual dentine isthmus; M3 with fossette; ectoflexid without cement in the m2.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FF4FF8BFF781F840" blockId="7.[113,763,1677,1983]" pageId="7" pageNumber="640">
|
||
<taxonomicName id="24D5C2A9CA1D8930FF4FF8BFF411F87F" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[141,363,1896,1920]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="taatsinius" status="sp. nov.">
|
||
<emphasis id="D1A16538CA1D8930FF4FF8BFF411F87F" box="[141,363,1896,1920]" italics="true" pageId="7" pageNumber="640">Prodistylomys taatsinius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CA92D843CA1D8930FEB0F8BEF4B9F87F" box="[370,451,1897,1920]" pageId="7" pageNumber="640" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="24D5C2A9CA1D8930FD87F8BEF715F87F" box="[581,623,1896,1920]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="lii">
|
||
<emphasis id="D1A16538CA1D8930FD87F8BEF715F87F" box="[581,623,1896,1920]" italics="true" pageId="7" pageNumber="640">P. lii</emphasis>
|
||
</taxonomicName>
|
||
by its smaller size, moreslenderlowerteeth, p4 withmetaflexidandposterolophid, drop-shaped trigonid of the m1, and different morphology of the m3.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FC87FF47F18CFF18" blockId="7.[810,1459,144,231]" pageId="7" pageNumber="640">
|
||
<taxonomicName id="24D5C2A9CA1D8930FC87FF47F155FF57" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[837,1071,144,168]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="taatsinius" status="sp. nov.">
|
||
<emphasis id="D1A16538CA1D8930FC87FF47F155FF57" box="[837,1071,144,168]" italics="true" pageId="7" pageNumber="640">Prodistylomys taatsinius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CA92D843CA1D8930FBF7FF47F1FBFF58" box="[1077,1153,144,167]" pageId="7" pageNumber="640" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="24D5C2A9CA1D8930FAC0FF46F0EEFF57" box="[1282,1428,145,168]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="7" pageNumber="640" phylum="Chordata" rank="species" species="xinjiangensis">
|
||
<emphasis id="D1A16538CA1D8930FAC0FF46F0EEFF57" box="[1282,1428,145,168]" italics="true" pageId="7" pageNumber="640">P. xinjiangensis</emphasis>
|
||
</taxonomicName>
|
||
by its larger size, lack of metafossettid, lack of metaflexid in the m1 and m2, and different morphology of the m3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABCFEAA1CA1D893FFCE8FEDFF7CBF980" lastPageId="8" lastPageNumber="641" pageId="7" pageNumber="640" type="description">
|
||
<paragraph id="E36AB92ACA1D8930FCE8FEDFF1D8FEA0" blockId="7.[810,1459,264,351]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8FEDFF697FEDF" box="[810,1005,264,288]" italics="true" pageId="7" pageNumber="640">Other fossil layers:</emphasis>
|
||
UNCH-A/3; NHMW 2012/0049/0001- 0005 (
|
||
<figureCitation id="7BEEA5AFCA1D8930FCB8FEF0F6C7FEC0" box="[890,957,295,319]" captionStart="Figure 5" captionStartId="9.[113,178,930,954]" captionTargetBox="[121,1450,152,895]" captionTargetId="figure-110@9.[114,1458,144,903]" captionTargetPageId="9" captionText="Figure 5. Upper and lower cheek teeth of Prodistylomys taatsinius sp. nov. from the Valley of Lakes (Mongolia). From Unkheltseg:(A) UNCH-A/3 (NHMW 2012/0049/0003) P4 left; (B) UNCH-A/3 (NHMW 2012/0049/0005) M1/2 left; (C) UNCH-A/3 (NHMW 2012/0049/0001) m2 right; From Huch Teeg: (D) RHN-A/12 (NHMW 2012/0050/0001) DP4 right; (E) RHN-A/12 (NHMW 2012/0050/0003) M1/2 right; (F) RHN-A/12 (NHMW 2012/0050/0006) p4 right; (G) RHN-A/12 (NHMW 2012/0050/0002) m3 right. All teeth are at the same magnification.Left-side specimens are underlined.Scale bar:1 mm." figureDoi="http://doi.org/10.5281/zenodo.10469786" httpUri="https://zenodo.org/record/10469786/files/figure.png" pageId="7" pageNumber="640">Fig. 5</figureCitation>
|
||
A-C, Model S2), and RHN-A/12; NHMW 2012/0050/0001-0006 (
|
||
<figureCitation id="7BEEA5AFCA1D8930FBE9FE91F118FEA1" box="[1067,1122,326,350]" captionStart="Figure 6" captionStartId="9.[113,178,1607,1631]" captionTargetBox="[119,1449,1129,1569]" captionTargetId="figure-192@9.[114,1458,1123,1579]" captionTargetPageId="9" captionText="Figure 6. Prodistylomys mongoliensis sp. nov. from the type locality of Hotuliin Teeg, Valley of Lakes, Mongolia:(A) holotype of Prodistylomys mongoliensis sp. nov. from Hotuliin Teeg (HTE-012), (NHMW 2012/0051/0001). Right m2; (B) HTE-012 (NHMW 2012/0051/0002) M3 left. Left-side specimens are underlined. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10469788" httpUri="https://zenodo.org/record/10469788/files/figure.png" pageId="7" pageNumber="640">Fig. 6</figureCitation>
|
||
D-G).
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCE8FE56F169FE68" blockId="7.[810,1043,384,408]" box="[810,1043,384,408]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8FE56F6C6FE67" box="[810,956,385,408]" italics="true" pageId="7" pageNumber="640">Measurements:</emphasis>
|
||
<tableCitation id="AE578C91CA1D8930FC00FE57F177FE67" box="[962,1037,384,408]" captionStart="Table 4" captionStartId="12.[111,166,444,469]" captionTargetPageId="12" captionText="Table 4. Descriptive statistics of the upper and lower cheek teeth of Prodistylomys including the new species. All measurements are given in millimetres.Length and width represent the maximum antero-posterior and bucco-lingual distances. Abbreviations: Elem., element; N., number of teeth measured; min, minimum; max, maximum. For abbreviations of assemblages see Table 2." pageId="7" pageNumber="640">Table 4</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCE8FE6EF009FE2E" blockId="7.[810,1395,441,465]" box="[810,1395,441,465]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8FE6EF690FE2E" box="[810,1002,441,465]" italics="true" pageId="7" pageNumber="640">Stratigraphic range:</emphasis>
|
||
Early Miocene, Mongolian biozone D.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCE8FE25F185FDF5" blockId="7.[810,1279,498,522]" box="[810,1279,498,522]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8FE25F697FDF5" box="[810,1005,498,522]" italics="true" pageId="7" pageNumber="640">Geographical range:</emphasis>
|
||
Valley of Lakes,
|
||
<collectingCountry id="9BC2F9BACA1D8930FB57FE25F180FDF5" box="[1173,1274,498,522]" name="Mongolia" pageId="7" pageNumber="640">Mongolia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCFBFDEEF0DEFDAD" blockId="7.[825,1444,568,594]" box="[825,1444,568,594]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCFBFDEEF0DEFDAD" box="[825,1444,568,594]" italics="true" pageId="7" pageNumber="640">Description of the new material flom Luugar Khudag (LOG-A/1)</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCE8FD88F6A3FC4F" blockId="7.[810,1460,606,944]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8FD88F636FD88" box="[810,844,607,631]" italics="true" pageId="7" pageNumber="640">p4:</emphasis>
|
||
It is always hypsodont and rooted, with two (3/4) or three roots (1/4). There is cement in the metaflexid. The trigonid is rhomboidal, with the metaconid and protoconid narrower than the anterolophid. The metaconid and the protoconid are blunt. The trigonid is always larger than the talonid. The talonid is blunt, with the entoconid and hypoconid wider than the posterolophid. The entoconid is sharp. The mesoflexid is V-shaped, open and deep. The ectoflexid is V-shaped, open in three (3/4) and closed in one (1/4); one of them is shallow, whereas the remaining three are deep. The metaflexid is absent (1/3), incipient (1/3), or present (1/3).
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCE8FC05F167FA9D" blockId="7.[810,1459,978,1378]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8FC05F604FC15" box="[810,894,978,1002]" italics="true" pageId="7" pageNumber="640">m1/m2:</emphasis>
|
||
The teeth are always bilophodont and hypsodont. The roots vary between two (12/14) and three (2/14). If there is cement, it is included in the most internal part of the mesoflexid and ectoflexid (2/14). The shape of the trigonid is drop shaped (10/13) or fusiform (3/13). The metaconid is sharp in nine out of
|
||
<specimenCount id="F5D372A3CA1D8930FC85FBB9F6AFFB79" box="[839,981,1134,1158]" count="10" pageId="7" pageNumber="640" type="generic">10 specimens</specimenCount>
|
||
and blunt in one. The protoconid is sharp in two (2/11) and blunt in nine (9/11). The trigonid is larger than the talonid in five (5/7) and smaller in two (2/7). The talonid is always rhomboidal. The entoconid is sharp. The hypoconid is sharp in seven out of eight and blunt in one. The mesoflexid is V-shaped and deep, open in nine (9/13) and closed in four (4/13). The ectoflexid is also V-shaped, open in 12 out of 14 and closed in two, and deep.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCE8FA53F0CEF9E6" blockId="7.[810,1460,1412,1561]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8FA53F628FA63" box="[810,850,1412,1436]" italics="true" pageId="7" pageNumber="640">m3:</emphasis>
|
||
There is only
|
||
<specimenCount id="F5D372A3CA1D8930FC2BFA53F100FA63" box="[1001,1146,1412,1436]" count="1" pageId="7" pageNumber="640" type="generic">one specimen</specimenCount>
|
||
. The tooth is bilophodont and hypsodont with two roots. The trigonid is subtringular. The metaconid is blunt, the protoconid is sharp. The talonid is larger than the trigonid. The talonid is rounded with a blunt entoconid and hypoconid. The flexids are fused and form a shallow fossette.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCE8F9ECF6D5F8B2" blockId="7.[810,1459,1595,1869]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8F9ECF61BF9AC" box="[810,865,1595,1619]" italics="true" pageId="7" pageNumber="640">DP4:</emphasis>
|
||
The description is based on the material from the locality Huch Teeg (
|
||
<specimenCode id="B3731151CA1D8930FC2BF98DF116F98D" box="[1001,1132,1626,1650]" collectionCode="RHN-A" pageId="7" pageNumber="640">RHN-A/12</specimenCode>
|
||
). There are
|
||
<specimenCount id="F5D372A3CA1D8930FB27F98CF0F8F98D" box="[1253,1410,1626,1650]" count="2" pageId="7" pageNumber="640" type="generic">two specimens</specimenCount>
|
||
, one of them has the posterior part broken. The tooth is rooted, with only one root, and low. The main cusps are well developed (paracone, metacone, protocone, hypocone). The anterocone is absent. The metaloph is connected to the central part of the long hypocone. The entoloph is connected to the protocone, metaloph, and hypocone. Extended hypocone and posteroloph.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA1D8930FCE8F8B8F14DF83A" blockId="7.[810,1459,1903,1989]" pageId="7" pageNumber="640">
|
||
<emphasis id="D1A16538CA1D8930FCE8F8B8F634F878" box="[810,846,1903,1927]" italics="true" pageId="7" pageNumber="640">P4:</emphasis>
|
||
There are only
|
||
<specimenCount id="F5D372A3CA1D8930FC35F8B8F1E9F878" box="[1015,1171,1903,1927]" count="2" pageId="7" pageNumber="640" type="generic">two specimens</specimenCount>
|
||
, one of them damaged. The teeth are simple and subtriangular. They are hypsodont and rooted with only one root.
|
||
</paragraph>
|
||
<caption id="B7AAE9A2CA12893FFF41FB61F73CFADD" ID-DOI="http://doi.org/10.5281/zenodo.10469784" ID-Zenodo-Dep="10469784" httpUri="https://zenodo.org/record/10469784/files/figure.png" pageId="8" pageNumber="641" startId="8.[131,196,1206,1230]" targetBox="[139,1467,152,1169]" targetPageId="8" targetType="figure">
|
||
<paragraph id="E36AB92ACA12893FFF41FB61F73CFADD" blockId="8.[130,1475,1206,1314]" pageId="8" pageNumber="641">
|
||
<emphasis id="D1A16538CA12893FFF41FB61F767FB31" bold="true" box="[131,541,1206,1230]" pageId="8" pageNumber="641">
|
||
Figure 4.
|
||
<taxonomicName id="24D5C2A9CA12893FFF1DFB61F4B4FB31" authority="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck, 2023" authorityName="Oliver & Carro-Rodríguez & López-Guerrero & Daxner-Höck" authorityYear="2023" box="[223,462,1206,1230]" class="Mammalia" family="Ctenodactylidae" genus="Prodistylomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="8" pageNumber="641" phylum="Chordata" rank="species" species="taatsinius" status="sp. nov.">
|
||
<emphasis id="D1A16538CA12893FFF1DFB61F4B4FB31" bold="true" box="[223,462,1206,1230]" italics="true" pageId="8" pageNumber="641">Prodistylomys taatsinius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="CA92D843CA12893FFE10FB61F767FB31" box="[466,541,1206,1230]" pageId="8" pageNumber="641" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from the type locality of LOG-A/1, in Luugar Khudag, Valley of Lakes (Mongolia): (A) LOG-A/1 (NHMW 2012/0048/0019) M1/2 left; (B) LOG-A/1 (NHMW 2012/0048/0022) M3 right; (C) LOG-A/1 (NHMW 2012/0048/0003) m1 right; (D) LOG-A/1 (NHMW 2012/0048/001) from left to right: p4-m1-m2 right, holotype. All teeth are at the same magnification. Leftside specimens are underlined. Scale bar: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E36AB92ACA12893FFF43FA85F7BCFA18" blockId="8.[129,778,1362,1511]" pageId="8" pageNumber="641">
|
||
<emphasis id="D1A16538CA12893FFF43FA85F5A3FA95" box="[129,217,1362,1386]" italics="true" pageId="8" pageNumber="641">M1/M2:</emphasis>
|
||
Hypsodont and rooted teeth always with two roots. In three out of
|
||
<specimenCount id="F5D372A3CA12893FFF37FAA6F4F0FA76" box="[245,394,1393,1417]" count="4" pageId="8" pageNumber="641" type="generic">four specimens</specimenCount>
|
||
there is cement in the flexus.The trigon is kidney shaped and larger than the talon.The talon varies in shape, i.e. kidney shaped (1/3) or subelliptical (2/3) but always narrower than the trigon. The mesoflexus connects to the hypoflexus.
|
||
</paragraph>
|
||
<paragraph id="E36AB92ACA12893FFF43F9DEF7CBF980" blockId="8.[129,778,1545,1663]" pageId="8" pageNumber="641">
|
||
<emphasis id="D1A16538CA12893FFF43F9DEF5D6F9DE" box="[129,172,1545,1569]" italics="true" pageId="8" pageNumber="641">M3:</emphasis>
|
||
There is only
|
||
<specimenCount id="F5D372A3CA12893FFEF5F9DEF4B9F9DE" box="[311,451,1545,1569]" count="1" pageId="8" pageNumber="641" type="generic">one specimen</specimenCount>
|
||
broken lingually. It is hypsodont and with two roots fused. It has cement in the fossette. Both the trigon and the talon are kidney shaped. The talon is narrower than the trigon. The flexus is fused and forms a fossette.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABCFEAA1CA12893FFF43F977F752F908" pageId="8" pageNumber="641" type="discussion">
|
||
<paragraph id="E36AB92ACA12893FFF43F977F752F908" blockId="8.[129,778,1696,1783]" pageId="8" pageNumber="641">
|
||
<emphasis id="D1A16538CA12893FFF43F977F5A4F947" box="[129,222,1696,1720]" italics="true" pageId="8" pageNumber="641">Remarks:</emphasis>
|
||
This species occurs in the region of Taatsin Gol, in the localities of Luugar Khudag (LOG-A/1), Huch Teeg (RHN-A/12), and in Unkheltseg (UNCH-A/3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |