treatments-xml/data/6B/77/85/6B7785212A5CFFCE2851C0F7FE30FAEB.xml

<document id="1A846BB776681A290AF667C410433732" ID-CLB-Dataset="27816" ID-DOI="10.5852/ejt.2019.508" ID-GBIF-Dataset="6e176ff1-c6ce-429c-8b36-9d32c3cdc5c9" ID-ISSN="2118-9773" ID-Zenodo-Dep="2612875" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1553714929206" checkinUser="plazi" docAuthor="Rahayu, Sri &amp; Rodda, Michele" docDate="2019" docId="6B7785212A5CFFCE2851C0F7FE30FAEB" docLanguage="en" docName="ejt-508_rahayu_rodda.pdf.imf" docOrigin="European Journal of Taxonomy 508" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Hoya sumatrana Rahayu &amp; Rodda 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="18" masterDocId="974EFD592A52FFDF2A5FC529FF8CFF8F" masterDocTitle="Hoya of Sumatra, an updated checklist, three new species, and a new subspecies" masterLastPageNumber="23" masterPageNumber="1" pageNumber="15" updateTime="1698715017593" updateUser="ExternalLinkService">
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<mods:title id="F4EB26F9CF2ABB63A46C5D1A5CEEBB7D">Hoya of Sumatra, an updated checklist, three new species, and a new subspecies</mods:title>
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<mods:namePart id="EF85D0BC74423A061F715540A81C00F5">Rahayu, Sri</mods:namePart>
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<mods:namePart id="D7D93E36B2C1AF3A379EC06C4ECA4414">Rodda, Michele</mods:namePart>
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<subSubSection id="ABC467BC2A5CFFD12851C0F7FC61F990" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="E36134372A5CFFD12851C0F7FBAAFA77" blockId="14.[526,1062,1502,1605]" box="[526,1062,1502,1528]" pageId="14" pageNumber="15">
<heading id="B829835B2A5CFFD12851C0F7FBAAFA77" box="[526,1062,1502,1528]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
<taxonomicName id="24DE4FB42A5CFFD12851C0F7FC48FA77" ID-CoL="7WNGZ" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[526,964,1502,1528]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A5CFFD12851C0F7FD55FA77" bold="true" box="[526,729,1502,1528]" italics="true" pageId="14" pageNumber="15">Hoya sumatrana</emphasis>
S.Rahayu &amp; Rodda
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<taxonomicNameLabel id="CA99555E2A5CFFD12994C0F7FBAAFA77" box="[971,1062,1502,1528]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="E36134372A5CFFD12818C32DFC61F990" blockId="14.[526,1062,1502,1605]" box="[583,1005,1540,1567]" pageId="14" pageNumber="15">urn:lsid:ipni.org:names:77195002-1</paragraph>
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<paragraph id="E36134372A5CFFD128B8C302FCC1F9CA" blockId="14.[526,1062,1502,1605]" box="[743,845,1579,1605]" pageId="14" pageNumber="15">
<figureCitation id="7BE528B22A5CFFD128B8C302FCC1F9CA" box="[743,845,1579,1605]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="15.[189,236,1915,1944]" captionStartId-1="16.[189,236,1947,1976]" captionTargetBox-0="[230,1358,265,1874]" captionTargetBox-1="[189,1398,958,1907]" captionTargetId-0="figure@15.[230,1358,263,1874]" captionTargetId-1="figure@16.[189,1398,958,1907]" captionTargetPageId-0="15" captionText-0="Fig. 3. Hoya sumatrana S.Rahayu &amp; Rodda sp. nov. A. Buds. B. Inflorescence, from underneath. C. Flower, from top. D. Corolla, from underneath, calyx and pedicel. E. Flower, side view. F. Corona, from underneath. G. Pedicel, calyx and ovaries. H. Pollinarium. Photos: M. Rodda." captionText-1="Fig. 4. Hoya sumatrana S.Rahayu &amp; Rodda sp. nov. A–B. Leaves from above. C–D. Leaves from from underneath. Photos: M. Rodda." httpUri-0="https://zenodo.org/record/2612881/files/figure.png" httpUri-1="https://zenodo.org/record/2612883/files/figure.png" pageId="14" pageNumber="15">Figs 3–4</figureCitation>
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<heading id="B829835B2A5CFFD12AE2C359FEC8F905" bold="true" box="[189,324,1648,1674]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="D1AAE8252A5CFFD12AE2C359FEC8F905" bold="true" box="[189,324,1648,1674]" pageId="14" pageNumber="15">Etymology</emphasis>
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The new species is named after the Indonesian island of 
<collectingRegion id="211AFAD52A5CFFD1290BC38BFC36F933" box="[852,954,1698,1724]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="15">Sumatra</collectingRegion>
.
</paragraph>
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<emphasis id="D1AAE8252A5CFFD12AE2C3CEFEE0F88F" bold="true" box="[189,364,1766,1793]" pageId="14" pageNumber="15">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A5CFFD12AB5C231FD0CF8F0" blockId="14.[189,1399,1816,1919]" pageId="14" pageNumber="15">
<materialsCitation id="53B63E6A2A5CFFD12AB5C231FD0CF8F0" ID-GBIF-Occurrence="2029330399" collectingDate="2013-09-20" collectionCode="BO" collectorName="S. Rahayu" country="Indonesia" location="Tambling" municipality="Taman Nasional Bukit Barisan Selatan" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="D1AAE8252A5CFFD12AB5C231FED0F8BD" bold="true" box="[234,348,1816,1842]" pageId="14" pageNumber="15">
<typeStatus id="3C658A952A5CFFD12AB5C231FED0F8BD" box="[234,348,1816,1842]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="9BC974A72A5CFFD12AE2C216FED7F8D6" box="[189,347,1855,1881]" name="Indonesia" pageId="14" pageNumber="15">INDONESIA</collectingCountry>
• 
<collectingRegion id="211AFAD52A5CFFD12B2BC216FE55F8D6" box="[372,473,1855,1881]" country="Indonesia" name="Sumatera Utara" pageId="14" pageNumber="15">Sumatra</collectingRegion>
, 
<collectingRegion id="211AFAD52A5CFFD12BBBC216FDDBF8D6" box="[484,599,1855,1881]" country="Indonesia" name="Lampung" pageId="14" pageNumber="15">Lampung</collectingRegion>
, 
<collectingMunicipality id="0305AE4D2A5CFFD1283CC216FBA7F8D6" box="[611,1067,1855,1881]" pageId="14" pageNumber="15">Taman Nasional Bukit Barisan Selatan</collectingMunicipality>
, 
<location id="E60162EC2A5CFFD12E69C216FB26F8D6" LSID="urn:lsid:plazi:treatment:6B7785212A5CFFCE2851C0F7FE30FAEB:E60162EC2A5CFFD12E69C216FB26F8D6" box="[1078,1194,1855,1881]" country="Indonesia" municipality="Taman Nasional Bukit Barisan Selatan" name="Tambling" pageId="14" pageNumber="15" stateProvince="Sumatera Utara">Tambling</location>
; 
<date id="976012F72A5CFFD12EE9C216FADFF8D6" box="[1206,1363,1855,1881]" pageId="14" pageNumber="15" value="2013-09-20">
<collectingDate id="8724EB1F2A5CFFD12EE9C216FADFF8D6" box="[1206,1363,1855,1881]" pageId="14" pageNumber="15" value="2013-09-20">20 Sep. 2013</collectingDate>
</date>
; 
<collectorName id="4E2B51E12A5CFFD12F00C216FE9BF8F0" pageId="14" pageNumber="15">S. Rahayu</collectorName>
861 leg.; lowland forests; 
<collectionCode id="85CFACF22A5CFFD12810C24CFDF7F8F0" box="[591,635,1893,1919]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15186" name="Herbarium Bogoriense" pageId="14" pageNumber="15" type="Herbarium">BO</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="E36134372A5CFFD12AB5C283FEC9F84B" blockId="14.[189,812,1962,2027]" box="[234,325,1962,1988]" pageId="14" pageNumber="15">
<emphasis id="D1AAE8252A5CFFD12AB5C283FEC9F84B" bold="true" box="[234,325,1962,1988]" pageId="14" pageNumber="15">
<typeStatus id="3C658A952A5CFFD12AB5C283FEC9F84B" box="[234,325,1962,1988]" pageId="14" pageNumber="15" type="isotype">Isotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="E36134372A5CFFD12AE2C2F8FCA0F864" blockId="14.[189,812,1962,2027]" box="[189,812,2001,2027]" pageId="14" pageNumber="15">
<materialsCitation id="53B63E6A2A5CFFD12AE2C2F8FCABF864" ID-GBIF-Occurrence="2029333402" box="[189,807,2001,2027]" collectingDate="2013-09-20" collectionCode="SING" collectorName="S. Rahayu" country="Indonesia" location="Same" municipality="Taman Nasional Bukit Barisan Selatan" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Sumatera Utara">
<collectingCountry id="9BC974A72A5CFFD12AE2C2F8FED7F864" box="[189,347,2001,2027]" name="Indonesia" pageId="14" pageNumber="15">INDONESIA</collectingCountry>
• 
<location id="E60162EC2A5CFFD12B2BC2F8FE39F864" LSID="urn:lsid:plazi:treatment:6B7785212A5CFFCE2851C0F7FE30FAEB:E60162EC2A5CFFD12B2BC2F8FE39F864" box="[372,437,2001,2027]" country="Indonesia" name="Same" pageId="14" pageNumber="15">Same</location>
data as for the holotype; 
<collectionCode id="85CFACF22A5CFFD12880C2F8FCABF864" box="[735,807,2001,2027]" country="Singapore" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15640" lsid="urn:lsid:biocol.org:col:15640" name="Singapore Botanic Gardens" pageId="14" pageNumber="15" type="Herbarium">SING</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="B7A164BF2A5DFFD02AE2C252FAFBF86A" httpUri="https://zenodo.org/record/2612881/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,236,1915,1944]" targetBox="[230,1358,265,1874]" targetPageId="15">
<paragraph id="E36134372A5DFFD02AE2C252FAFBF86A" blockId="15.[189,1399,1915,2021]" pageId="15" pageNumber="16">
<emphasis id="D1AAE8252A5DFFD02AE2C252FE9DF817" bold="true" box="[189,273,1915,1944]" pageId="15" pageNumber="16">Fig. 3.</emphasis>
<taxonomicName id="24DE4FB42A5DFFD02B47C255FD5BF818" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[280,727,1916,1944]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A5DFFD02B47C255FE61F817" box="[280,493,1916,1944]" italics="true" pageId="15" pageNumber="16">Hoya sumatrana</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A5DFFD02882C255FCB2F817" box="[733,830,1916,1944]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="D1AAE8252A5DFFD0291AC252FCD0F817" bold="true" box="[837,860,1915,1944]" pageId="15" pageNumber="16">A</emphasis>
. Buds. 
<emphasis id="D1AAE8252A5DFFD029E2C255FC5EF817" bold="true" box="[957,978,1916,1944]" pageId="15" pageNumber="16">B</emphasis>
. Inflorescence, from underneath. 
<emphasis id="D1AAE8252A5DFFD02AE2C28BFF58F830" bold="true" box="[189,212,1954,1983]" pageId="15" pageNumber="16">C</emphasis>
. Flower, from top. 
<emphasis id="D1AAE8252A5DFFD02B85C28BFE7DF831" bold="true" box="[474,497,1954,1982]" pageId="15" pageNumber="16">D</emphasis>
. Corolla, from underneath, calyx and pedicel. 
<emphasis id="D1AAE8252A5DFFD02E01C28BFBFFF831" bold="true" box="[1118,1139,1954,1982]" pageId="15" pageNumber="16">E</emphasis>
. Flower, side view. 
<emphasis id="D1AAE8252A5DFFD02AE2C2E0FF5DF86A" bold="true" box="[189,209,1993,2021]" pageId="15" pageNumber="16">F</emphasis>
. Corona, from underneath. 
<emphasis id="D1AAE8252A5DFFD0286DC2E1FDC7F86A" bold="true" box="[562,587,1992,2021]" pageId="15" pageNumber="16">G</emphasis>
. Pedicel, calyx and ovaries. 
<emphasis id="D1AAE8252A5DFFD029E7C2E0FC5DF86A" bold="true" box="[952,977,1993,2021]" pageId="15" pageNumber="16">H</emphasis>
. Pollinarium. Photos: M. Rodda.
</paragraph>
</caption>
<subSubSection id="ABC467BC2A42FFCE2AE2C422FBD4FD83" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="E36134372A42FFCF2AE2C422FEC2FEAA" blockId="16.[189,334,267,293]" box="[189,334,267,293]" pageId="16" pageNumber="17">
<heading id="B829835B2A42FFCF2AE2C422FEC2FEAA" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="D1AAE8252A42FFCF2AE2C422FEC2FEAA" bold="true" box="[189,334,267,293]" pageId="16" pageNumber="17">Description</emphasis>
</heading>
</paragraph>
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Epiphytic climber, with white or yellowish latex in all vegetative parts. Roots adventitious below the nodes and along the internodes. Stems 
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in diameter, climbing, dull green, pubescent; older stems grey, glabrescent; internodes 2–5(10) cm. Leaf blades 3–7(15) × 
<quantity id="242699D22A42FFCF29F3C4A8FBBDFE13" box="[940,1073,385,412]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="16" pageNumber="17" unit="cm" value="4.5" valueMax="5.5" valueMin="3.5">3.5–5.5 cm</quantity>
, stiff and succulent, oblong, ovate or elliptic, base round to attenuate, apex acute, mid green on adaxial surface with sparse grey spots, pale green on abaxial surface, glabrous, or very sparsely pubescent when young, venation pinnate, inconspicuous, basal colleter ca 1 × 
<quantity id="242699D22A42FFCF2831C4C4FD41FD87" box="[622,717,493,520]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="17" unit="mm" value="1.5">1.5 mm</quantity>
, one at each lamina base, broadly triangular, pale brown; petioles 10–20 × 
<quantity id="242699D22A42FFCF2BD8C738FE67FDA3" box="[391,491,529,556]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="16" pageNumber="17" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
, terete, dark brown, sparsely pubescent when young. Inflorescences 
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in diameter, of 10–15 flowers, sciadioidal, flat; peduncles 15–35 × 
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, extra-axillary, positively geotropic, 1 at each node, producing flowers over an extended period of time, terete, dark red, sparsely pubescent. Flowers with terete pedicels 5–15 × 
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, dark red at the base, fading to pale pink towards the calyx, sparsely papillose. Calyx lobes narrowly triangular, spaced 
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apart, apex acute or rounded, 1.4–2 × 
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, reddish, papillose outside, inside glabrous, sparsely ciliate; basal colleters 0.16–0.18 × 
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, one at each calyx sinus, ovoid. Corolla 
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in diameter when flattened, rotate; tube ca 
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long, cream, glabrous outside, densely pubescent inside; lobes triangular, valvate in bud, with recurved edges and revolute tip, 5–6 × 
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, glabrous outside, densely pubescent inside with a glabrous tip ca 
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long. Corona staminal 
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high, 
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in diameter, red; lobes ca 2.2 × 
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, attached at the back of the anthers, held at ca 60° angle, ovoid, slightly carinate above, inner process bilobed, outer process rounded, with a narrow basal revolute margin. Anthers ca 0.5 × 
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, ovate, with a linear apical round membranaceous appendage to 
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long. Pollinia 280–320 × 100–120 μm, oblong, with a round base and obliquely truncate; pellucid margin all along the outer edge; caudicles 100–120 × 70 μm, broad, almost transparent; corpusculum 90–110 × 40–60 μm, oblong; style-head 5-angled in cross section, with five spreading lobes alternating with the stamens, style-head apex 
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long, 
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broad at the base, conical, with apex acute; ovary 1.5–2 × ca 
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at the base, conical, with apex acute. Fruit and seed not observed.
</paragraph>
</subSubSection>
<caption id="B7A164BF2A42FFCF2AE2C2B2FD1EF850" httpUri="https://zenodo.org/record/2612883/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,236,1947,1976]" targetBox="[189,1398,958,1907]" targetPageId="16">
<paragraph id="E36134372A42FFCF2AE2C2B2FD1EF850" blockId="16.[189,1398,1947,2015]" pageId="16" pageNumber="17">
<emphasis id="D1AAE8252A42FFCF2AE2C2B2FE9DF837" bold="true" box="[189,273,1947,1976]" pageId="16" pageNumber="17">Fig. 4.</emphasis>
<taxonomicName id="24DE4FB42A42FFCF2B48C2B4FD5FF838" authority="S.Rahayu &amp; Rodda" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[279,723,1949,1977]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A42FFCF2B48C2B4FE60F836" box="[279,492,1949,1977]" italics="true" pageId="16" pageNumber="17">Hoya sumatrana</emphasis>
S.Rahayu &amp; Rodda
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A42FFCF2886C2B5FCB6F837" box="[729,826,1948,1976]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="D1AAE8252A42FFCF291EC2B2FCF7F837" bold="true" box="[833,891,1947,1976]" pageId="16" pageNumber="17">A–B</emphasis>
. Leaves from above. 
<emphasis id="D1AAE8252A42FFCF2ED4C2B2FB44F837" bold="true" box="[1163,1224,1947,1976]" pageId="16" pageNumber="17">C–D</emphasis>
. Leaves from from underneath. Photos: M. Rodda.
</paragraph>
</caption>
<subSubSection id="ABC467BC2A43FFCE2AE2C71EFBC3FD26" pageId="17" pageNumber="18" type="distribution">
<paragraph id="E36134372A43FFCE2AE2C71EFE62FDDE" blockId="17.[189,494,567,593]" box="[189,494,567,593]" pageId="17" pageNumber="18">
<heading id="B829835B2A43FFCE2AE2C71EFE62FDDE" bold="true" box="[189,494,567,593]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="D1AAE8252A43FFCE2AE2C71EFE62FDDE" bold="true" box="[189,494,567,593]" pageId="17" pageNumber="18">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A43FFCE2AE2C743FBC3FD26" blockId="17.[189,1399,617,682]" pageId="17" pageNumber="18">
<taxonomicName id="24DE4FB42A43FFCE2AE2C743FE0FFD0C" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[189,387,618,643]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A43FFCE2AE2C743FE0FFD0C" box="[189,387,618,643]" italics="true" pageId="17" pageNumber="18">Hoya sumatrana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A43FFCE2BD3C740FE64FD0C" box="[396,488,617,643]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is only known from the 
<typeStatus id="3C658A952A43FFCE294CC740FCCAFD0C" box="[787,838,617,643]" pageId="17" pageNumber="18">type</typeStatus>
locality in in 
<collectingRegion id="211AFAD52A43FFCE29B0C740FBEEFD0C" box="[1007,1122,617,643]" country="Indonesia" name="Lampung" pageId="17" pageNumber="18">Lampung</collectingRegion>
, 
<collectingRegion id="211AFAD52A43FFCE2E2FC740FB59FD0C" box="[1136,1237,617,643]" country="Indonesia" name="Sumatera Utara" pageId="17" pageNumber="18">Sumatra</collectingRegion>
, 
<collectingCountry id="9BC974A72A43FFCE2EBDC740FAD4FD0C" box="[1250,1368,617,643]" name="Indonesia" pageId="17" pageNumber="18">Indonesia</collectingCountry>
. It was collected in a coastal swamp forest, where it was growing as an epiphyte.
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A43FFCE2AE2C7FDFE8FFC3D" pageId="17" pageNumber="18" type="conservation">
<paragraph id="E36134372A43FFCE2AE2C7FDFE39FD61" blockId="17.[189,437,724,750]" box="[189,437,724,750]" pageId="17" pageNumber="18">
<heading id="B829835B2A43FFCE2AE2C7FDFE39FD61" bold="true" box="[189,437,724,750]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="D1AAE8252A43FFCE2AE2C7FDFE39FD61" bold="true" box="[189,437,724,750]" pageId="17" pageNumber="18">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="E36134372A43FFCE2AE2C62FFEEEFCE2" blockId="17.[189,1398,774,877]" pageId="17" pageNumber="18">
Known only from a single collection and lacking information on the distribution area, the population size and the possible threats to the habitat, 
<taxonomicName id="24DE4FB42A43FFCE28E8C604FCD5FCC9" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[695,857,813,838]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A43FFCE28E8C604FCD5FCC9" box="[695,857,813,838]" italics="true" pageId="17" pageNumber="18">H. sumatrana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A43FFCE293EC604FC30FCC8" box="[865,956,813,839]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is considered as Data Deficient (DD) (
<bibRefCitation id="874F49C62A43FFCE2A9AC67AFEDAFCE2" author="IUCN &amp; Cambridge, UK" box="[197,342,851,877]" pageId="17" pageNumber="18" refId="ref8900" refString="IUCN 2012. IUCN Red List Categories and Criteria (version 3.1, 2 nd edition). IUCN, Gland, Switzerland and Cambridge, UK." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
<paragraph id="E36134372A43FFCE2AE2C6B1FE8FFC3D" blockId="17.[189,259,920,946]" box="[189,259,920,946]" pageId="17" pageNumber="18">
<heading id="B829835B2A43FFCE2AE2C6B1FE8FFC3D" bold="true" box="[189,259,920,946]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="D1AAE8252A43FFCE2AE2C6B1FE8FFC3D" bold="true" box="[189,259,920,946]" pageId="17" pageNumber="18">Notes</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ABC467BC2A43FFCE2AE2C6E3FE30FAEB" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="E36134372A43FFCE2AE2C6E3FE30FAEB" blockId="17.[189,1399,969,1380]" pageId="17" pageNumber="18">
<taxonomicName id="24DE4FB42A43FFCE2AE2C6E3FEF3FC6C" ID-CoL="7WNGZ" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[189,383,970,995]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A43FFCE2AE2C6E3FEF3FC6C" box="[189,383,970,995]" italics="true" pageId="17" pageNumber="18">Hoya sumatrana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A43FFCE2BDBC6E3FE51FC6B" box="[388,477,970,996]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is a new species in 
<emphasis id="D1AAE8252A43FFCE28E6C6E3FD5AFC6C" box="[697,726,970,995]" italics="true" pageId="17" pageNumber="18">H.</emphasis>
sect. 
<emphasis id="D1AAE8252A43FFCE294BC6E0FC20FC6C" box="[788,940,969,995]" italics="true" pageId="17" pageNumber="18">Peltostemma</emphasis>
<bibRefCitation id="874F49C62A43FFCE29EEC6E0FB21FC6C" author="Schlechter R." box="[945,1197,969,995]" pageId="17" pageNumber="18" pagination="1 - 18" refId="ref10043" refString="Schlechter R. 1916. Neue Asclepiadaceen von Sumatra und Celebes. Beihefte zum Botanischen Centralblatt 34 (2): 1 - 18." type="journal article" year="1916">Schlechter (1916: 14)</bibRefCitation>
, characterized by upright corona lobes, long linear anther appendages extending well above the style-head and pollinaria with well developed caudicle wings. This section includes six species, two of which, 
<taxonomicName id="24DE4FB42A43FFCE2E98C13EFEE2FBD8" ID-CoL="6MF68" authority="Kidyoo (2013)" authorityName="Kidyoo" authorityYear="2013" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="soidaoensis">
<emphasis id="D1AAE8252A43FFCE2E98C13EFAFBFBBF" box="[1223,1399,1046,1072]" italics="true" pageId="17" pageNumber="18">H. soidaoensis</emphasis>
<bibRefCitation id="874F49C62A43FFCE2AE2C115FEE2FBD8" author="Kidyoo M." box="[189,366,1084,1111]" pageId="17" pageNumber="18" pagination="45 - 50" refId="ref8993" refString="Kidyoo M. 2013. Hoya soidaoensis (Apocynaceae: Asclepiadoideae), a new species from southeastern Thailand. Phytotaxa 105 (2): 45 - 50. https: // doi. org / 10.11646 / phytotaxa. 105.2.3" type="journal article" year="2013">Kidyoo (2013)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="24DE4FB42A43FFCE2BF1C114FCA0FBD8" ID-CoL="3MRYN" authority="Kidyoo (2016)" authorityName="Kidyoo" authorityYear="2016" box="[430,812,1084,1111]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="phuwuaensis">
<emphasis id="D1AAE8252A43FFCE2BF1C114FDE2FBD8" box="[430,622,1085,1111]" italics="true" pageId="17" pageNumber="18">H. phuwuaensis</emphasis>
<bibRefCitation id="874F49C62A43FFCE2827C115FCA0FBD8" author="Kidyoo M." box="[632,812,1084,1111]" pageId="17" pageNumber="18" pagination="218 - 224" refId="ref9035" refString="Kidyoo M. 2016. Hoya phuwuaensis (Apocynaceae: Asclepiadoideae), a new species from Northeastern Thailand. Phytotaxa 282 (3): 218 - 224. https: // doi. org / 10.11646 / phytotaxa. 282.3.5" type="journal article" year="2016">Kidyoo (2016)</bibRefCitation>
</taxonomicName>
, have been recently published. 
<taxonomicName id="24DE4FB42A43FFCE2EF1C114FAFAFBD9" ID-CoL="7WNGZ" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[1198,1398,1085,1110]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A43FFCE2EF1C114FAFAFBD9" box="[1198,1398,1085,1110]" italics="true" pageId="17" pageNumber="18">Hoya sumatrana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A43FFCE2AE2C14DFE95FBF1" box="[189,281,1124,1150]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from all other members of this section because its flowers are much smaller; for example, the corolla of 
<taxonomicName id="24DE4FB42A43FFCE2BA7C1A3FD15FB2C" ID-CoL="7WNGZ" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[504,665,1162,1187]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A43FFCE2BA7C1A3FD15FB2C" box="[504,665,1162,1187]" italics="true" pageId="17" pageNumber="18">H. sumatrana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A43FFCE28FFC1A3FD76FB2B" box="[672,762,1162,1188]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is 
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in diameter, while the corolla of the second smallest species, 
<taxonomicName id="24DE4FB42A43FFCE2BD6C198FCBCFB45" ID-CoL="3MRQ5" authority="Kerr (1940: tab. 3407)" authorityName="Kerr" authorityYear="1940" box="[393,816,1200,1226]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="flagellata">
<emphasis id="D1AAE8252A43FFCE2BD6C198FD90FB45" box="[393,540,1200,1226]" italics="true" pageId="17" pageNumber="18">H. flagellata</emphasis>
<bibRefCitation id="874F49C62A43FFCE287BC199FCBCFB45" author="Kerr A. F. G." box="[548,816,1200,1226]" pageId="17" pageNumber="18" pagination="3406 - 3407" refId="ref8967" refString="Kerr A. F. G. 1940. Hoya mitrata &amp; Hoya flagellata. Hooker's Icones Plantarum 35: tab. 3406 - 3407." type="book chapter" year="1940">Kerr (1940: tab. 3407)</bibRefCitation>
</taxonomicName>
, is&gt; 
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in diameter, as well as the corona of 
<taxonomicName id="24DE4FB42A43FFCE2AE2C1FEFE0EFB7F" ID-CoL="7WNGZ" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[189,386,1239,1264]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A43FFCE2AE2C1FEFE0EFB7F" box="[189,386,1239,1264]" italics="true" pageId="17" pageNumber="18">Hoya sumatrana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A43FFCE2BD5C1FEFE69FB7E" box="[394,485,1239,1265]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
is 
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in diameter, while that of 
<taxonomicName id="24DE4FB42A43FFCE2990C1FEFBEFFB7F" ID-CoL="3MRQ5" authorityName="Kerr" authorityYear="1940" box="[975,1123,1238,1264]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="flagellata">
<emphasis id="D1AAE8252A43FFCE2990C1FEFBEFFB7F" box="[975,1123,1238,1264]" italics="true" pageId="17" pageNumber="18">H. flagellata</emphasis>
</taxonomicName>
is&gt; 
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in diameter. Another useful character to separate 
<taxonomicName id="24DE4FB42A43FFCE2833C1D7FC81FA98" ID-CoL="7WNGZ" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[620,781,1278,1303]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A43FFCE2833C1D7FC81FA98" box="[620,781,1278,1303]" italics="true" pageId="17" pageNumber="18">H. sumatrana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A43FFCE294AC1D4FCFCFA98" box="[789,880,1277,1303]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
from all other members of the section is the inner corona lobe process, that is bilobed in 
<taxonomicName id="24DE4FB42A43FFCE2893C00DFCE2FAB2" ID-CoL="7WNGZ" authority="Rahayu &amp; Rodda, 2019" authorityName="Rahayu &amp; Rodda" authorityYear="2019" box="[716,878,1316,1341]" class="Magnoliopsida" family="Apocynaceae" genus="Hoya" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="sumatrana" status="sp. nov.">
<emphasis id="D1AAE8252A43FFCE2893C00DFCE2FAB2" box="[716,878,1316,1341]" italics="true" pageId="17" pageNumber="18">H. sumatrana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CA99555E2A43FFCE2928C00DFC59FAB1" box="[887,981,1316,1342]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, while the other species have acute or rounded processes.
</paragraph>
</subSubSection>
</treatment>
</document>