253 lines
39 KiB
XML
253 lines
39 KiB
XML
<document id="29E11803728C6B8949988A4D71D9B4A1" ID-CLB-Dataset="4967" ID-DOI="10.11646/zootaxa.4984.1.7" ID-GBIF-Dataset="9ccf3e9f-c4cf-4202-b39c-4a87c9763f81" ID-ISSN="1175-5326" ID-Zenodo-Dep="4928596" ID-ZooBank="604672CE-8468-4A69-93A3-D8024DB42327" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="admin" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1623435240056" checkinUser="plazi" docAuthor="Snegovaya, Nataly Yu., Cokendolpher, James C. & Zamani, Alireza" docDate="2021" docId="6B0C87BCFFC0C64A03D07E055AC35219" docLanguage="en" docName="zootaxa.4984.1.7.pdf" docOrigin="Zootaxa 4984 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phalangium martensi Snegovaya & Cokendolpher & Zamani 2021, spec. nov." docType="treatment" docVersion="9" lastPageNumber="77" masterDocId="9735FFC4FFC2C64E0347792E5E0E574C" masterDocTitle="Further studies on harvestmen (Arachnida: Opiliones) from Iran, with the descriptions of two new species" masterLastPageNumber="86" masterPageNumber="73" pageNumber="75" updateTime="1720380746336" updateUser="admin" zenodo-license-document="CLOSED">
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<mods:title id="7D6DDF8BF6560AC67AF09070457F2E4F">Further studies on harvestmen (Arachnida: Opiliones) from Iran, with the descriptions of two new species</mods:title>
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<mods:name id="0F5EC1C5071C12A2BB10B73B60F61179" type="personal">
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<mods:namePart id="FB995E5FD3E5674267C2C07DFFE40EAD">Snegovaya, Nataly Yu.</mods:namePart>
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<mods:affiliation id="3E20666EC36F09DDF90A935F1F58DA02">Institute of Zoology, Azerbaijan Academy of Sciences, pr. 1128, kv. 504, Baku AZ 1073, Azerbaijan.</mods:affiliation>
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<mods:name id="C162AA197FC332AB7167B88FF307B90A" type="personal">
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<mods:roleTerm id="41FB407D5B1FCCA236489715F7E76F1C">Author</mods:roleTerm>
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<mods:namePart id="0F286EE8CB8C819A488C381697F70689">Cokendolpher, James C.</mods:namePart>
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<mods:affiliation id="39442BF35B57D58EE9EE813003765FA7">Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, Texas 79415, USA.</mods:affiliation>
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<mods:namePart id="3A4CC7230E5DA12CF0C7CE0D3566B591">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="F2E81A4F7BA5B0D66BD6334352A41C3B">Zoological Museum, Biodiversity Unit, FI- 20014, University of Turku, Finland.</mods:affiliation>
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<mods:typeOfResource id="B1F7848FFB8D9145CADD3BD348485E55">text</mods:typeOfResource>
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<mods:title id="2F01E32FE2F7A3FEE424A8A9768F9CA3">Zootaxa</mods:title>
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<mods:date id="7F1E103BC49232D087BAB2D10351C593">2021</mods:date>
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<mods:number id="9AE4995AB49E8042B7C479DEABD2EAC5">2021-06-10</mods:number>
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<treatment id="6B0C87BCFFC0C64A03D07E055AC35219" ID-DOI="http://doi.org/10.5281/zenodo.5196942" ID-GBIF-Taxon="184181576" ID-Zenodo-Dep="5196942" LSID="urn:lsid:plazi:treatment:6B0C87BCFFC0C64A03D07E055AC35219" httpUri="http://treatment.plazi.org/id/6B0C87BCFFC0C64A03D07E055AC35219" lastPageId="4" lastPageNumber="77" pageId="2" pageNumber="75">
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<subSubSection id="ABBF6521FFC0C64C03D07E055C135009" box="[151,541,1835,1862]" pageId="2" pageNumber="75" type="nomenclature">
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<paragraph id="E31A36AAFFC0C64C03D07E055C135009" blockId="2.[151,541,1835,1897]" box="[151,541,1835,1862]" pageId="2" pageNumber="75">
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<heading id="B85281C6FFC0C64C03D07E055C135009" bold="true" box="[151,541,1835,1862]" fontSize="11" level="1" pageId="2" pageNumber="75" reason="1">
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<emphasis id="D1D1EAB8FFC0C64C03D07E055C135009" bold="true" box="[151,541,1835,1862]" pageId="2" pageNumber="75">
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<taxonomicName id="24A54D29FFC0C64C03D07E055F905009" ID-CoL="9X5HB" authority="Snegovaya & Cokendolpher & Zamani, 2021" authorityName="Snegovaya & Cokendolpher & Zamani" authorityYear="2021" box="[151,414,1835,1862]" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="75" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
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<emphasis id="D1D1EAB8FFC0C64C03D07E055F905009" bold="true" box="[151,414,1835,1862]" italics="true" pageId="2" pageNumber="75">Phalangium martensi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="CAE257C3FFC0C64C02E17E055C135009" box="[422,541,1835,1861]" pageId="2" pageNumber="75" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="ABBF6521FFC0C64C03D07E615F1D5025" box="[151,275,1871,1897]" pageId="2" pageNumber="75" type="description">
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<paragraph id="E31A36AAFFC0C64C03D07E615F1D5025" blockId="2.[151,541,1835,1897]" box="[151,275,1871,1897]" pageId="2" pageNumber="75">
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<figureCitation id="7B9E2A2FFFC0C64C03D07E615F1D5025" box="[151,275,1871,1897]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="2" pageNumber="75">Figs 2A–K</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="ABBF6521FFC0C64C03D07EB95F2750B5" pageId="2" pageNumber="75" type="materials_examined">
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<paragraph id="E31A36AAFFC0C64C03D07EB95F2750B5" blockId="2.[151,1436,1943,2041]" pageId="2" pageNumber="75">
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<emphasis id="D1D1EAB8FFC0C64C03D07EB95F3250FD" bold="true" box="[151,316,1943,1969]" pageId="2" pageNumber="75">Type material</emphasis>
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.
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<materialsCitation id="53CD3CF7FFC0C64C020F7EB95D855099" ID-GBIF-Occurrence="3334349305" collectingDate="2019-01" collectionCode="ZMMU" collectorName="Alireza Zamani" country="Iran" elevation="549" latitude="32.6" location="Khuzestan Province" longLatPrecision="1209" longitude="48.466667" pageId="2" pageNumber="75" specimenCount="1" stateProvince="Khuzestan" typeStatus="holotype">
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<emphasis id="D1D1EAB8FFC0C64C020F7EB95D3050FD" bold="true" box="[328,830,1943,1970]" pageId="2" pageNumber="75">
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<collectingCountry id="9BB2763AFFC0C64C020F7EB95F9A50FD" box="[328,404,1943,1969]" name="Iran" pageId="2" pageNumber="75">IRAN</collectingCountry>
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:
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<emphasis id="D1D1EAB8FFC0C64C02D97EB95C8A50FD" bold="true" box="[414,644,1943,1970]" italics="true" pageId="2" pageNumber="75">
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<collectingRegion id="2161F848FFC0C64C02D97EB95C8A50FD" box="[414,644,1943,1970]" country="Iran" name="Khuzestan" pageId="2" pageNumber="75">Khuzestan Province</collectingRegion>
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</emphasis>
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:
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<typeStatus id="3C1E8808FFC0C64C01D57EB95CF150FD" box="[658,767,1943,1969]" pageId="2" pageNumber="75" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(
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<collectionCode id="85B4AE6FFFC0C64C000C7EB95DA650FD" box="[843,936,1943,1969]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="75" type="Museum">ZMMU</collectionCode>
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), Dezful, Qaleh Shadab (Shahiun), Bazargah,
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<geoCoordinate id="8691506DFFC0C64C03D07E955EF45099" box="[151,250,1979,2005]" degrees="32" direction="north" minutes="36" orientation="latitude" pageId="2" pageNumber="75" precision="925" value="32.6">32°36’N</geoCoordinate>
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,
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<geoCoordinate id="8691506DFFC0C64C02407E955F695099" box="[263,359,1979,2005]" degrees="48" direction="east" minutes="28" orientation="longitude" pageId="2" pageNumber="75" precision="925" value="48.466667">48°28’E</geoCoordinate>
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,
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<elevation id="6888D199FFC0C64C02337E955FF45099" box="[372,506,1979,2006]" metricMagnitude="2" metricUnit="m" metricValue="5.49" pageId="2" pageNumber="75" unit="m" value="549.0">
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<quantity id="245D9B4FFFC0C64C02337E955FB2509A" box="[372,444,1979,2006]" metricMagnitude="2" metricUnit="m" metricValue="5.49" pageId="2" pageNumber="75" unit="m" value="549.0">549 m</quantity>
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a.s.l.
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</elevation>
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,
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<collectorName id="4E50537CFFC0C64C01417E955CB95099" box="[518,695,1979,2005]" pageId="2" pageNumber="75">Alireza Zamani</collectorName>
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leg.
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<date id="971B106AFFC0C64C01B77E925D855099" box="[752,907,1979,2006]" pageId="2" pageNumber="75" value="2019-01">
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<collectingDate id="875FE982FFC0C64C01B77E925D855099" box="[752,907,1979,2006]" pageId="2" pageNumber="75" value="2019-01">January 2019</collectingDate>
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</date>
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||
</materialsCitation>
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||
.
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<materialsCitation id="53CD3CF7FFC0C64C00D17E955F2B50B5" ID-GBIF-Occurrence="3334349325" collectingDate="2019-01" collectionCode="RCNS" collectorName="Alireza Zamani" country="Iran" elevation="549" latitude="32.6" location="Khuzestan Province" longLatPrecision="1209" longitude="48.466667" pageId="2" pageNumber="75" specimenCount="1" specimenCount-male="1" stateProvince="Khuzestan" typeStatus="paratype">
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<emphasis id="D1D1EAB8FFC0C64C00D17E955A045099" bold="true" box="[918,1034,1979,2005]" pageId="2" pageNumber="75">
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<typeStatus id="3C1E8808FFC0C64C00D17E955A0B5099" box="[918,1029,1979,2005]" pageId="2" pageNumber="75" type="paratype">Paratype</typeStatus>
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.
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</emphasis>
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<specimenCount id="F5A3FD23FFC0C64C07557E955A515099" box="[1042,1119,1979,2005]" pageId="2" pageNumber="75" type="male">1 male</specimenCount>
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(
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<collectionCode id="85B4AE6FFFC0C64C07287E955AB05099" box="[1135,1214,1979,2005]" pageId="2" pageNumber="75">RCNS</collectionCode>
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), with same data as for holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<caption id="B7DA6622FFC1C64D03D07EB95A9150BB" ID-DOI="http://doi.org/10.5281/zenodo.4928600" ID-Zenodo-Dep="4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="3" pageNumber="76" startId="3.[151,264,1943,1967]" targetBox="[237,1350,181,1905]" targetPageId="3">
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<paragraph id="E31A36AAFFC1C64D03D07EB95A9150BB" blockId="3.[151,1417,1943,2040]" pageId="3" pageNumber="76">
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<emphasis id="D1D1EAB8FFC1C64D03D07EB95F5A50E3" bold="true" box="[151,340,1943,1967]" pageId="3" pageNumber="76">FIGURES 2A–K.</emphasis>
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<taxonomicName id="24A54D29FFC1C64D021D7EB65C3B50E3" authority="Snegovaya & Cokendolpher & Zamani, 2021" authorityName="Snegovaya & Cokendolpher & Zamani" authorityYear="2021" box="[346,565,1944,1968]" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
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<emphasis id="D1D1EAB8FFC1C64D021D7EB65C3B50E3" box="[346,565,1944,1968]" italics="true" pageId="3" pageNumber="76">Phalangium martensi</emphasis>
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</taxonomicName>
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<emphasis id="D1D1EAB8FFC1C64D017C7EB65CAC50FC" bold="true" box="[571,674,1944,1968]" pageId="3" pageNumber="76">
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<taxonomicNameLabel id="CAE257C3FFC1C64D017C7EB65CAC50FC" box="[571,674,1944,1968]" pageId="3" pageNumber="76" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm
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</paragraph>
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</caption>
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||
<subSubSection id="ABBF6521FFC6C64A038079B95D4F5799" pageId="4" pageNumber="77" type="etymology">
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<paragraph id="E31A36AAFFC6C64A038079B95D4F5799" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="77">
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<emphasis id="D1D1EAB8FFC6C64A038079B95F4657FD" bold="true" box="[199,328,151,177]" pageId="4" pageNumber="77">Etymology</emphasis>
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. The specific name “
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<taxonomicName id="24A54D29FFC6C64A017079B95C9757FC" authorityName="Snegovaya & Cokendolpher & Zamani" authorityYear="2021" box="[567,665,151,176]" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="martensi">
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<emphasis id="D1D1EAB8FFC6C64A017079B95C9757FC" box="[567,665,151,176]" italics="true" pageId="4" pageNumber="77">martensi</emphasis>
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</taxonomicName>
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” is the genitive of the family name of our colleague Prof. Dr Jochen Martens, one of the world’s leading authorities on
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<taxonomicName id="24A54D29FFC6C64A018A79955D305799" authorityName="Sundevall" authorityYear="1833" box="[717,830,187,213]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="77" phylum="Arthropoda" rank="order">Opiliones</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="ABBF6521FFC6C64A038079F05D1C55BD" pageId="4" pageNumber="77" type="diagnosis">
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<paragraph id="E31A36AAFFC6C64A038079F05DB15629" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="77">
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<emphasis id="D1D1EAB8FFC6C64A038079F05F3457B4" bold="true" box="[199,314,222,248]" pageId="4" pageNumber="77">Diagnosis</emphasis>
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. The new species has long legs, all of about the same thickness, with longitudinal rows of fairly large denticles (
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<figureCitation id="7B9E2A2FFFC6C64A024B782D5FBC5651" box="[268,434,259,285]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2A–B, D</figureCitation>
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). Pedipalp slender, long, all segments only with setae (
|
||
<figureCitation id="7B9E2A2FFFC6C64A076D782D5AA85651" box="[1066,1190,259,285]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2E–F</figureCitation>
|
||
); chelicera with large rigid dorsal horn (
|
||
<figureCitation id="7B9E2A2FFFC6C64A021878095C2D560D" box="[351,547,295,321]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2A–B, G–H</figureCitation>
|
||
). The penis has a thin truncus with a wide base, tapering sharply from 1/3 of the penis length (
|
||
<figureCitation id="7B9E2A2FFFC6C64A026878655F705629" box="[303,382,331,357]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Fig. 2I</figureCitation>
|
||
); the glans is triangular in lateral view (
|
||
<figureCitation id="7B9E2A2FFFC6C64A007D78655DBD5629" box="[826,947,331,357]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2J–K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E31A36AAFFC6C64A038078405D1C55BD" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="77">
|
||
<emphasis id="D1D1EAB8FFC6C64A038078405FDF56C5" bold="true" box="[199,465,366,393]" pageId="4" pageNumber="77">Comparative remarks</emphasis>
|
||
. Judging from the presence of a hard dorsal horn on the second segment of the chelicera, the new species is most similar to
|
||
<taxonomicName id="24A54D29FFC6C64A010878BD5D7756E1" authority="Audouin, 1826" authorityName="Audouin" authorityYear="1826" box="[591,889,403,429]" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="savignyi">
|
||
<emphasis id="D1D1EAB8FFC6C64A010878BD5CC656E0" box="[591,712,403,428]" italics="true" pageId="4" pageNumber="77">P. savignyi</emphasis>
|
||
<bibRefCitation id="87344B5BFFC6C64A019778BD5D7756E1" author="Audouin, V." box="[720,889,403,429]" pageId="4" pageNumber="77" pagination="99 - 186" refId="ref6277" refString="Audouin, V. (1826) Explication sommaire des planches d'Arachnides de l'Egypte et de la Syrie, publiees par Jules-Cesar Savigny, membre de l'Institut; offrant un expose des caracteres naturelles des genres, avec la distinction des especes. In: Jomard, E. F. (Ed.), Description de l'Egypte: ou recueil des observations et des recherches qui ont ete faites en Egypte pendant l'expedition de l'armee francaise, publie par les ordres de Sa Majeste l'Empereur Napoleon le Grand, 1 (4 eme partie). L'Imprimerie Royale, Paris, pp. 99 - 186." type="book chapter" year="1826">Audouin, 1826</bibRefCitation>
|
||
</taxonomicName>
|
||
and can be distinguished by the following characteristics: longer legs (Body Leg Index = 2.13) and different shape of femur I which is cylindrical (
|
||
<figureCitation id="7B9E2A2FFFC6C64A07B378995B7D569D" box="[1268,1395,439,465]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2A–B</figureCitation>
|
||
), in contrast to the fusiform shape in
|
||
<taxonomicName id="24A54D29FFC6C64A014078F55C8E56B8" authorityName="Audouin" authorityYear="1826" box="[519,640,475,500]" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="savignyi">
|
||
<emphasis id="D1D1EAB8FFC6C64A014078F55C8E56B8" box="[519,640,475,500]" italics="true" pageId="4" pageNumber="77">P. savignyi</emphasis>
|
||
</taxonomicName>
|
||
, and especially by not having the 1st pair of legs thickened; longer and more slender palps (
|
||
<figureCitation id="7B9E2A2FFFC6C64A023A78D15FF65555" box="[381,504,511,537]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2E–F</figureCitation>
|
||
); different structure of the penis (
|
||
<figureCitation id="7B9E2A2FFFC6C64A003378D15DE25555" box="[884,1004,511,537]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2I–K</figureCitation>
|
||
). The new species is also similar to
|
||
<taxonomicName id="24A54D29FFC6C64A06CF78D15CA55571" authority="Tchemeris & Snegovaya, 2010" authorityName="Tchemeris & Snegovaya" authorityYear="2010" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="kopetdaghensis">
|
||
<emphasis id="D1D1EAB8FFC6C64A06CF78D15F4A5571" italics="true" pageId="4" pageNumber="77">P. kopetdaghensis</emphasis>
|
||
<bibRefCitation id="87344B5BFFC6C64A020B7B0D5CA55571" author="Tchemeris, A. N. & Snegovaya, N. Y." box="[332,683,547,573]" pageId="4" pageNumber="77" pagination="67 - 72" refId="ref8967" refString="Tchemeris, A. N. & Snegovaya, N. Y. (2010) Three new species of Phalangiidae (Arachnida: Opiliones) from Turkmenistan. Acta Arachnologica, 59, 67 - 72. https: // doi. org / 10.2476 / asjaa. 59.67" type="journal article" year="2010">Tchemeris & Snegovaya, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
, but different in the following features: longitudinal rows of spiky denticles located on the legs (
|
||
<figureCitation id="7B9E2A2FFFC6C64A02A27B695C33552D" box="[485,573,583,609]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Fig. 2D</figureCitation>
|
||
) (vs. blunt denticles); femur I has a cylindrical shape, in contrast to the fusiform shape in
|
||
<taxonomicName id="24A54D29FFC6C64A02727B455FF255C9" authorityName="Tchemeris & Snegovaya" authorityYear="2010" box="[309,508,619,645]" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="kopetdaghensis">
|
||
<emphasis id="D1D1EAB8FFC6C64A02727B455FF255C9" box="[309,508,619,645]" italics="true" pageId="4" pageNumber="77">P. kopetdaghensis</emphasis>
|
||
</taxonomicName>
|
||
; a rigid dorsal horn on the second segment of the chelicera (unlike the softer horn in
|
||
<taxonomicName id="24A54D29FFC6C64A03D07BA15F6E55E5" authorityName="Tchemeris & Snegovaya" authorityYear="2010" box="[151,352,655,681]" class="Arachnida" family="Phalangiidae" genus="Phalangium" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="77" phylum="Arthropoda" rank="species" species="kopetdaghensis">
|
||
<emphasis id="D1D1EAB8FFC6C64A03D07BA15F6E55E5" box="[151,352,655,681]" italics="true" pageId="4" pageNumber="77">P. kopetdaghensis</emphasis>
|
||
</taxonomicName>
|
||
) and the structure of the penis (truncus with a wide base and sharply tapering to 1/3 of its length distally), and with thinner distal part, the so-called “spoon” of the penis [i.e. upper third of the distal part (
|
||
<figureCitation id="7B9E2A2FFFC6C64A06707B9D5B885581" box="[1335,1414,690,717]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Fig. 2I</figureCitation>
|
||
)], and by having longer and thinner pedipalps (
|
||
<figureCitation id="7B9E2A2FFFC6C64A01CC7BF95D0B55BD" box="[651,773,727,753]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2E–F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABBF6521FFC6C64A03807BD45DE6527D" pageId="4" pageNumber="77" type="description">
|
||
<paragraph id="E31A36AAFFC6C64A03807BD45B2953A5" blockId="4.[151,1437,151,1365]" pageId="4" pageNumber="77">
|
||
<emphasis id="D1D1EAB8FFC6C64A03807BD45F5F5458" bold="true" box="[199,337,762,788]" pageId="4" pageNumber="77">Description</emphasis>
|
||
.
|
||
<emphasis id="D1D1EAB8FFC6C64A02187BD55F925459" bold="true" box="[351,412,763,789]" pageId="4" pageNumber="77">Male</emphasis>
|
||
. Body length 4.7, width 3.0. Body medium-sized: coloration light brown, dorsally with dark brown saddle pattern. Transverse rows of small denticles located along boundaries of tergites (
|
||
<figureCitation id="7B9E2A2FFFC6C64A07F77A315B045475" box="[1200,1290,799,825]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Fig. 2A</figureCitation>
|
||
). Eye mound large (length 0.6, width 0.6, height 0.45) and round, with 6–7 large black-tipped denticles and with several denticles on either sides (
|
||
<figureCitation id="7B9E2A2FFFC6C64A020C7A495FAB54CD" box="[331,421,871,897]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Fig. 2C</figureCitation>
|
||
); 3–4 teeth located in front of eye mound (
|
||
<figureCitation id="7B9E2A2FFFC6C64A00CC7A495A0254CD" box="[907,1036,871,897]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2A, C</figureCitation>
|
||
). Legs long, 1st pair not thickened, femora with longitudinal rows of spiky denticles (
|
||
<figureCitation id="7B9E2A2FFFC6C64A01877AA55D6E54E9" box="[704,864,907,933]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2A–B, D</figureCitation>
|
||
). Lengths of leg segments: I - 6.4+1.3+4.6+7.0+8.7 = 28.0; II - 10.0+1.4+7.8+8.7+18.2 = 46.1; III - 6.0+1.4+4.5+7.6+10.8 = 30.3; IV - 7.7+1.5+5.6+10.2+13.6 = 38.6. Pedipalp slender, long, all segments with only setae (
|
||
<figureCitation id="7B9E2A2FFFC6C64A01B47AFD5D6054A1" box="[755,878,979,1005]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2E–F</figureCitation>
|
||
); segment lengths: 4.3+1.0+2.2+4.6 = 12.1. Chelicera with large horn-shaped (2.8 long), rigid dorsal outgrowth (= apophysis) on second segment; basal segment dorsally with several teeth (= acute tubercles); second segment with teeth and setae dorsally and with setae laterally (
|
||
<figureCitation id="7B9E2A2FFFC6C64A03D87D115F2C5315" box="[159,290,1087,1113]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Figs 2G–H</figureCitation>
|
||
). Basal cheliceral segment 1.9 long, second segment 4.5 long. Abdomen, coxae and legs light yellow (sandy) in color. Coxae with bristles (
|
||
<figureCitation id="7B9E2A2FFFC6C64A01067D4D5C975331" box="[577,665,1123,1149]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Fig. 2B</figureCitation>
|
||
). Penis small (2.6 long), thin truncus with a wide base. Glans somewhat triangular in lateral view, with 2 pairs of distal setae (
|
||
<figureCitation id="7B9E2A2FFFC6C64A007E7DA95D9A53ED" box="[825,916,1159,1185]" captionStart="FIGURES 2" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[237,1350,181,1905]" captionTargetId="figure-20@3.[237,1350,181,1919]" captionTargetPageId="3" captionText="FIGURES 2A–K. Phalangium martensi spec. nov., holotype male. A, B Habitus (A dorsal, B lateral). C Eye mound, dorsal. D Femur I, dorsal. E, F Right pedipalp (E mesal, F lateral). G, H Right chelicera (G mesal, H lateral). I, J Penis (I dorsal, J lateral). K Close-up of penis glans, lateral. Scale lines: A, B, D–H: 1 mm; C, I–J: 0.5 mm; K: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.4928600" httpUri="https://zenodo.org/record/4928600/files/figure.png" pageId="4" pageNumber="77">Fig. 2K</figureCitation>
|
||
). Glans 0.4 long, stylus 0.15 long. Penis light yellow (sandy) in color, with darker edging in apical part. The penis of the
|
||
<typeStatus id="3C1E8808FFC6C64A008B7D855A225389" box="[972,1068,1195,1221]" pageId="4" pageNumber="77" type="paratype">paratype</typeStatus>
|
||
is the same shape, color, and size as that of the
|
||
<typeStatus id="3C1E8808FFC6C64A02697DE15F9A53A5" box="[302,404,1231,1257]" pageId="4" pageNumber="77" type="holotype">holotype</typeStatus>
|
||
. The
|
||
<typeStatus id="3C1E8808FFC6C64A02977DE15C3E53A5" box="[464,560,1231,1257]" pageId="4" pageNumber="77" type="paratype">paratype</typeStatus>
|
||
also does not differ from the
|
||
<typeStatus id="3C1E8808FFC6C64A003B7DE15DD053A5" box="[892,990,1231,1257]" pageId="4" pageNumber="77" type="holotype">holotype</typeStatus>
|
||
in other features of the body.
|
||
</paragraph>
|
||
<paragraph id="E31A36AAFFC6C64A03807DDD5FAC5241" blockId="4.[151,1437,151,1365]" box="[199,418,1267,1293]" pageId="4" pageNumber="77">
|
||
<emphasis id="D1D1EAB8FFC6C64A03807DDD5F105241" bold="true" box="[199,286,1267,1293]" pageId="4" pageNumber="77">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="E31A36AAFFC6C64A03807C385DE6527D" blockId="4.[151,1437,151,1365]" box="[199,1000,1302,1329]" pageId="4" pageNumber="77">
|
||
<emphasis id="D1D1EAB8FFC6C64A03807C385F2D527C" bold="true" box="[199,291,1302,1328]" pageId="4" pageNumber="77">Habitat</emphasis>
|
||
. Mountainous and sparsely vegetated, mostly with low bushes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ABBF6521FFC6C64A03807C145AC35219" box="[199,1229,1338,1365]" pageId="4" pageNumber="77" type="distribution">
|
||
<paragraph id="E31A36AAFFC6C64A03807C145AC35219" blockId="4.[151,1437,151,1365]" box="[199,1229,1338,1365]" pageId="4" pageNumber="77">
|
||
<emphasis id="D1D1EAB8FFC6C64A03807C145F575218" bold="true" box="[199,345,1338,1364]" pageId="4" pageNumber="77">Distribution</emphasis>
|
||
. Known only from the
|
||
<typeStatus id="3C1E8808FFC6C64A01277C155C9E5219" box="[608,656,1339,1365]" pageId="4" pageNumber="77">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="2161F848FFC6C64A00487C155DFC5219" box="[783,1010,1339,1365]" country="Iran" name="Khuzestan" pageId="4" pageNumber="77">Khuzestan Province</collectingRegion>
|
||
, southwestern
|
||
<collectingCountry id="9BB2763AFFC6C64A07DE7C155AC45219" box="[1177,1226,1339,1365]" name="Iran" pageId="4" pageNumber="77">Iran</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |