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<document id="2B9624D21BCBE89EEE775E4E29E81C11" ID-CLB-Dataset="23993" ID-DOI="10.11646/zootaxa.4739.1.1" ID-GBIF-Dataset="d3e60fb6-636b-4e03-b55a-0e910f9a468e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3672547" ID-ZooBank="544B9C82-BF33-4EA1-9411-E1A307137466" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1582013879038" checkinUser="plazi" docAuthor="Salazar-Vallejo, Sergio I." docDate="2020" docId="2D1987E4FFA07369FF23FC44F69C60FD" docLanguage="en" docName="zootaxa.4739.1.1.pdf.imf" docOrigin="Zootaxa 4739 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leocrates rizzoae Salazar-Vallejo 2020, n. sp." docType="treatment" docVersion="13" lastPageNumber="70" masterDocId="D120FF9CFFE3732CFFB4FFF5F462610A" masterDocTitle="Revision of Leocrates Kinberg, 1866 and Leocratides Ehlers, 1908 (Annelida, Errantia, Hesionidae)" masterLastPageNumber="114" masterPageNumber="1" pageNumber="68" updateTime="1701997224814" updateUser="plazi">
<mods:mods id="55CA4AB29DA74512EF67E25D8637F0B1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="057922E3DE0B6DD908D3922B98298ECC">Revision of Leocrates Kinberg, 1866 and Leocratides Ehlers, 1908 (Annelida, Errantia, Hesionidae)</mods:title>
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<mods:namePart id="CC1631F9B1E0EE9B67CE9D70FBC44744">Salazar-Vallejo, Sergio I.</mods:namePart>
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<mods:title id="5B62E41A5CDDD608B89F2D59D270572C">Zootaxa</mods:title>
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<mods:date id="376177BBDC647689EC8EF7F09F07EF26">2020</mods:date>
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<mods:number id="01A8A52D22B2D7766AEB2FBF8CDCF9D0">2020-02-18</mods:number>
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<mods:number id="0B97FFE13B8BF0D98F3ED6324763CB05">4739</mods:number>
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<mods:identifier id="AA944288AAFF3439EEA1880CC6CD1085" type="DOI">10.11646/zootaxa.4739.1.1</mods:identifier>
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<treatment id="2D1987E4FFA07369FF23FC44F69C60FD" ID-DOI="http://doi.org/10.5281/zenodo.3680195" ID-GBIF-Taxon="162161390" ID-Zenodo-Dep="3680195" LSID="urn:lsid:plazi:treatment:2D1987E4FFA07369FF23FC44F69C60FD" httpUri="http://treatment.plazi.org/id/2D1987E4FFA07369FF23FC44F69C60FD" lastPageId="69" lastPageNumber="70" pageId="67" pageNumber="68">
<subSubSection id="EDAA6579FFA0736FFF23FC44F5D062C1" box="[151,434,945,971]" pageId="67" pageNumber="68" type="nomenclature">
<paragraph id="A50F36F2FFA0736FFF23FC44F5D062C1" blockId="67.[151,964,945,1043]" box="[151,434,945,971]" pageId="67" pageNumber="68">
<heading id="FE47819EFFA0736FFF23FC44F5D062C1" bold="true" box="[151,434,945,971]" fontSize="11" level="1" pageId="67" pageNumber="68" reason="1">
<emphasis id="97C4EAE0FFA0736FFF23FC44F5D062C1" bold="true" box="[151,434,945,971]" pageId="67" pageNumber="68">
<taxonomicName id="62B04D71FFA0736FFF23FC44F50B62C1" ID-CoL="8e2cb8eb-b75c-40da-b24a-9d95a24677cd" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[151,361,945,971]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="67" pageNumber="68" phylum="Annelida" rank="species" species="rizzoae" status="sp. nov.">
<emphasis id="97C4EAE0FFA0736FFF23FC44F50B62C1" bold="true" box="[151,361,945,971]" italics="true" pageId="67" pageNumber="68">Leocrates rizzoae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CF7579BFFA0736FFEC4FC44F5D062C1" box="[368,434,945,971]" pageId="67" pageNumber="68" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="EDAA6579FFA0736FFF23FC21F7A66519" pageId="67" pageNumber="68" type="description">
<paragraph id="A50F36F2FFA0736FFF23FC21F57662E5" blockId="67.[151,964,945,1043]" box="[151,276,980,1007]" pageId="67" pageNumber="68">
<figureCitation id="3D8B2A77FFA0736FFF23FC21F48E62E5" box="[151,236,980,1007]" captionStart="FIGURE 38" captionStartId="68.[151,250,988,1013]" captionTargetBox="[173,1417,184,959]" captionTargetId="figure@68.[170,1418,181,962]" captionTargetPageId="68" captionText="FIGURE 38. Leocrates rizzoae n. sp., holotype (USNM 37642). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, ventral view. D. Chaetiger 10, right parapodium, anterior view, MGS. Scale bars: A: 0.7 mm, B: 0.5 mm, C: 0.6 mm, D: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3672626" httpUri="https://zenodo.org/record/3672626/files/figure.png" pageId="67" pageNumber="68">Figs 38</figureCitation>
,
<figureCitation id="3D8B2A77FFA0736FFF4CFC20F57662E5" box="[248,276,981,1007]" captionStart="FIGURE 39" captionStartId="69.[151,250,1048,1073]" captionTargetBox="[172,1416,527,1021]" captionTargetId="figure@69.[170,1418,525,1023]" captionTargetPageId="69" captionText="FIGURE 39. Leocrates rizzoae n. sp., paratype (USNM 37641). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetiger 10, right parapodium, anterior view, MGS (insets: upper and lower neurochaetal blades, notacicular lobe). Scale bars: A: 0.8 mm, B, C: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3672628" httpUri="https://zenodo.org/record/3672628/files/figure.png" pageId="67" pageNumber="68">39</figureCitation>
</paragraph>
<paragraph id="A50F36F2FFA0736FFF23FC0DF7A66519" blockId="67.[151,964,945,1043]" box="[151,964,1016,1043]" pageId="67" pageNumber="68">
<uri id="D1213AF0FFA0736FFF23FC0DF7A66519" box="[151,964,1016,1043]" pageId="67" pageNumber="68">
urn:lsid:zoobank.org:act:
<uuid id="D1160C27FFA0736FFE1AFC0DF7A66519" box="[430,964,1016,1043]" pageId="67" pageNumber="68">2D64FB95-619F-47F1-A19C-54B6D11CA155</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFA0736FFF23FBCBF046655F" box="[151,1060,1084,1109]" pageId="67" pageNumber="68" type="reference_group">
<paragraph id="A50F36F2FFA0736FFF23FBCBF046655F" blockId="67.[151,1060,1084,1109]" box="[151,1060,1084,1109]" pageId="67" pageNumber="68">
<treatmentCitationGroup id="85A011DCFFA0736FFF23FBCBF046655F" box="[151,1060,1084,1109]" pageId="67" pageNumber="68">
<treatmentCitation id="241110E3FFA0736FFF23FBCBF648655F" author="Fauvel, P." box="[151,554,1084,1109]" page="371" pageId="67" pageNumber="68" year="1919">
<taxonomicName id="62B04D71FFA0736FFF23FBCBF648655F" ID-CoL="6P997" authority=": Fauvel 1919: 371" authorityName=": Fauvel" authorityPageNumber="371" authorityYear="1919" box="[151,554,1084,1109]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="67" pageNumber="68" phylum="Annelida" rank="species" species="claparedii">
<emphasis id="97C4EAE0FFA0736FFF23FBCBF50A655F" box="[151,360,1085,1109]" italics="true" pageId="67" pageNumber="68">Leocrates claparedii</emphasis>
:
<bibRefCitation id="C1214B03FFA0736FFEC2FBC9F648655F" author="Fauvel" box="[374,554,1084,1109]" firstAuthor="Fauvel" pageId="67" pageNumber="68" pagination="315 - 473" refId="ref83663" refString="Fauvel, P. (1919) Annelides polychetes de Madagascar, de Djibouti et du Golfe Persique. Archives de Zoologie Experimentale et Generale, 58, 315 - 473. [http: // www. biodiversitylibrary. org / item / 29986 # page / 497 / mode / 1 up] https: // doi. org / 10.5962 / bhl. part. 8154" type="journal article" year="1919">Fauvel 1919: 371</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="241110E3FFA0736FFD81FBC9F6BB655F" author="Fauvel, P." box="[565,729,1084,1109]" page="61" pageId="67" pageNumber="68" year="1932">
<bibRefCitation id="C1214B03FFA0736FFD81FBC9F6BB655F" author="Fauvel" box="[565,729,1084,1109]" firstAuthor="Fauvel" pageId="67" pageNumber="68" pagination="1 - 262" refId="ref83869" refString="Fauvel, P. (1932) Annelida Polychaeta of the Indian Museum, Calcutta. Memoirs of the Indian Museum, 12, 1 - 262. [http: // faunaofindia. nic. in / PDFVolumes / memoirs / 012 / 01 / 0001 - 0262. pdf]" type="journal article" year="1932">Fauvel 1932: 61</bibRefCitation>
</treatmentCitation>
(
<emphasis id="97C4EAE0FFA0736FFD5CFBCBF76C655F" box="[744,782,1086,1109]" italics="true" pageId="67" pageNumber="68">non</emphasis>
<collectingRegion id="6774F810FFA0736FFCA0FBC8F72C655F" box="[788,846,1085,1109]" country="Tanzania" name="Pwani" pageId="67" pageNumber="68">Costa</collectingRegion>
<emphasis id="97C4EAE0FFA0736FFCE0FBC8F70A655E" box="[852,872,1085,1108]" italics="true" pageId="67" pageNumber="68">in</emphasis>
<bibRefCitation id="C1214B03FFA0736FFCDBFBC9F07B655F" author="Claparede" box="[879,1049,1084,1109]" firstAuthor="Claparede" pageId="67" pageNumber="68" pagination="313 - 584" refId="ref82602" refString="Claparede, E. (1868) Les annelides chetopodes du Golfe de Naples. Memoires de la Societe de Physique et d'Histoire Naturelle de Geneve, 19 (2), 313 - 584. [http: // www. biodiversitylibrary. org / item / 50163 # page / 375 / mode / 1 up]" type="journal article" year="1868">Claparède, 1868</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFA0736FFF23FB76F72D6747" pageId="67" pageNumber="68" type="materials_examined">
<paragraph id="A50F36F2FFA0736FFF23FB76F1666427" blockId="67.[151,1437,1154,2046]" pageId="67" pageNumber="68">
<emphasis id="97C4EAE0FFA0736FFF23FB76F5946597" bold="true" box="[151,502,1154,1181]" pageId="67" pageNumber="68">Type material. Indian Ocean.</emphasis>
<materialsCitation id="15D83CAFFFA0736FFE49FB77F76665EF" ID-GBIF-Occurrence="2571525054" collectingDate="1964-02-24" collectionCode="USNM" collectorName="J. Bohlke" country="Seychelles" latitude="-4.28" location="International Indian Ocean Expedition" longLatPrecision="21" longitude="55.746666" municipality="Islands" pageId="67" pageNumber="68" specimenCode="USNM 37642" specimenCount="8" specimenCount-male="8" typeStatus="holotype">
<emphasis id="97C4EAE0FFA0736FFE49FB77F6B66597" bold="true" box="[509,724,1154,1181]" pageId="67" pageNumber="68">
<collectingCountry id="DDA77662FFA0736FFE49FB77F6146597" box="[509,630,1154,1181]" name="Seychelles" pageId="67" pageNumber="68">Seychelles</collectingCountry>
<collectingMunicipality id="456BAC88FFA0736FFDCAFB76F6B66597" box="[638,724,1155,1181]" pageId="67" pageNumber="68">Islands</collectingMunicipality>
</emphasis>
.
<typeStatus id="7A0B8850FFA0736FFD56FB76F7286597" box="[738,842,1155,1181]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
(
<specimenCode id="F5169E89FFA0736FFCECFB76F79E6594" box="[856,1020,1155,1182]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="67" pageNumber="68" type="Museum">USNM 37642</specimenCode>
),
<location id="A06F6029FFA0736FFBA4FB76F4A765CB" LSID="urn:lsid:plazi:treatment:2D1987E4FFA07369FF23FC44F69C60FD:A06F6029FFA0736FFBA4FB76F4A765CB" country="Seychelles" county="D'Arros Island" latitude="-4.28" longLatPrecision="21" longitude="55.746666" municipality="Islands" name="International Indian Ocean Expedition" pageId="67" pageNumber="68">International Indian Ocean Expedition</location>
,
<location id="A06F6029FFA0736FFF65FB52F59065C8" LSID="urn:lsid:plazi:treatment:2D1987E4FFA07369FF23FC44F69C60FD:A06F6029FFA0736FFF65FB52F59065C8" box="[209,498,1191,1218]" country="Seychelles" county="D'Arros Island" latitude="-4.28" longLatPrecision="21" longitude="55.746666" municipality="Islands" name="Seychell Islands Program" pageId="67" pageNumber="68">Seychell Islands Program</location>
, Sta. F-67,
<location id="A06F6029FFA0736FFDC6FB52F74365CB" LSID="urn:lsid:plazi:treatment:2D1987E4FFA07369FF23FC44F69C60FD:A06F6029FFA0736FFDC6FB52F74365CB" box="[626,801,1191,1217]" country="Seychelles" county="D'Arros Island" latitude="-4.28" longLatPrecision="21" longitude="55.746666" municipality="Islands" name="Curieuse Island" pageId="67" pageNumber="68">Curieuse Island</location>
(=
<location id="A06F6029FFA0736FFCF4FB53F7D165C8" LSID="urn:lsid:plazi:treatment:2D1987E4FFA07369FF23FC44F69C60FD:A06F6029FFA0736FFCF4FB53F7D165C8" box="[832,947,1190,1218]" country="Seychelles" latitude="-4.28" longLatPrecision="21" longitude="55.746666" municipality="Islands" name="Ile Rouge" pageId="67" pageNumber="68">Île Rouge</location>
;
<geoCoordinate id="C0845035FFA0736FFC0AFB52F02965CB" box="[958,1099,1191,1217]" degrees="04" direction="south" minutes="16" orientation="latitude" pageId="67" pageNumber="68" precision="15" seconds="48" value="-4.28">04°16’48” S</geoCoordinate>
,
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), SE end,
<location id="A06F6029FFA0736FFAEFFB52F1FE65CB" LSID="urn:lsid:plazi:treatment:2D1987E4FFA07369FF23FC44F69C60FD:A06F6029FFA0736FFAEFFB52F1FE65CB" box="[1371,1436,1191,1217]" country="Seychelles" county="D'Arros Island" latitude="-4.28" longLatPrecision="21" longitude="55.746666" municipality="Islands" name="South" pageId="67" pageNumber="68">South</location>
point of
<location id="A06F6029FFA0736FFF4CFB3EF51765EC" LSID="urn:lsid:plazi:treatment:2D1987E4FFA07369FF23FC44F69C60FD:A06F6029FFA0736FFF4CFB3EF51765EC" box="[248,373,1227,1254]" country="Seychelles" county="D'Arros Island" latitude="-4.28" longLatPrecision="21" longitude="55.746666" municipality="Islands" name="Laraie Bay" pageId="67" pageNumber="68">Laraie Bay</location>
,
<quantity id="62489B17FFA0736FFE35FB3EF5CC65EC" box="[385,430,1227,1254]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="67" pageNumber="68" unit="m" value="8.0">
<specimenCount id="B3B6FD7BFFA0736FFE35FB3EF5CC65EC" box="[385,430,1227,1254]" pageId="67" pageNumber="68" type="male">8 m</specimenCount>
</quantity>
,
<date id="D10E1032FFA0736FFE0FFB3EF63065EF" box="[443,594,1227,1253]" pageId="67" pageNumber="68" value="1964-02-24">
<collectingDate id="C14AE9DAFFA0736FFE0FFB3EF63065EF" box="[443,594,1227,1253]" pageId="67" pageNumber="68" value="1964-02-24">24 Feb. 1964</collectingDate>
</date>
,
<collectorName id="08455324FFA0736FFDEAFB39F6AE65EF" box="[606,716,1227,1254]" pageId="67" pageNumber="68">J. Bohlke</collectorName>
, coll
</materialsCitation>
.
<materialsCitation id="15D83CAFFFA0736FFCA4FB3EF1666427" ID-GBIF-Occurrence="2571525048" collectingDate="1964-03-08" collectionCode="USNM" collectorName="One &amp; International Indian Ocean Expedition &amp; Seychelles Islands Program &amp; Sta. &amp; J. Bohlke &amp; D. Dockins &amp; R. Rosenblatt &amp; W. Starck &amp; J. Tyler &amp; K. Hill" country="Seychelles" latitude="-5.4" longitude="53.883335" pageId="67" pageNumber="68" specimenCode="USNM 37641" typeStatus="paratype">
<collectorName id="08455324FFA0736FFCA4FB3EF75D65EF" box="[784,831,1227,1253]" pageId="67" pageNumber="68">One</collectorName>
<typeStatus id="7A0B8850FFA0736FFCF3FB39F7C565EC" box="[839,935,1228,1254]" pageId="67" pageNumber="68" type="paratype">paratype</typeStatus>
(
<specimenCode id="F5169E89FFA0736FFC02FB3EF03965EC" box="[950,1115,1227,1254]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="67" pageNumber="68" type="Museum">USNM 37641</specimenCode>
),
<collectorName id="08455324FFA0736FFBDBFB3EF5776403" pageId="67" pageNumber="68">International Indian Ocean Expedition</collectorName>
,
<collectorName id="08455324FFA0736FFE95FB1AF6396400" box="[289,603,1263,1290]" pageId="67" pageNumber="68">
<location id="A06F6029FFA0736FFE95FB1AF5926403" LSID="urn:lsid:plazi:treatment:2D1987E4FFA07369FF23FC44F69C60FD:A06F6029FFA0736FFE95FB1AF5926403" box="[289,496,1263,1289]" country="Seychelles" county="D'Arros Island" latitude="-5.4" longLatPrecision="1305" longitude="53.883335" municipality="Amirante Islands" name="Seychelles Islands" pageId="67" pageNumber="68">Seychelles Islands</location>
Program
</collectorName>
,
<collectorName id="08455324FFA0736FFDDCFB1AF6F16403" box="[616,659,1263,1289]" pageId="67" pageNumber="68">Sta.</collectorName>
F-103,
<collectingMunicipality id="456BAC88FFA0736FFD5FFB1AF7CD6403" box="[747,943,1263,1289]" pageId="67" pageNumber="68">Amirante Islands</collectingMunicipality>
,
<collectingCounty id="4C6E4E7EFFA0736FFC0FFB1AF0066403" box="[955,1124,1263,1289]" pageId="67" pageNumber="68">D’Arros Island</collectingCounty>
(
<geoCoordinate id="C0845035FFA0736FFBC1FB1BF0BB6403" box="[1141,1241,1262,1289]" degrees="05" direction="south" minutes="24" orientation="latitude" pageId="67" pageNumber="68" precision="925" value="-5.4">05°24’ S</geoCoordinate>
,
<geoCoordinate id="C0845035FFA0736FFB53FB1BF12F6400" box="[1255,1357,1262,1290]" degrees="53" direction="east" minutes="53" orientation="longitude" pageId="67" pageNumber="68" precision="925" value="53.883335">53°53’ E</geoCoordinate>
), off E side,
<quantity id="62489B17FFA0736FFF66FAE7F5556424" box="[210,311,1298,1326]" metricMagnitude="1" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="67" pageNumber="68" unit="m" value="27.0" valueMax="29.0" valueMin="25.0">
<elevation id="2E9DD1C1FFA0736FFF66FAE7F5556424" box="[210,311,1298,1326]" metricMagnitude="1" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="67" pageNumber="68" unit="m" value="27.0" valueMax="29.0" valueMin="25.0">25–29 m</elevation>
</quantity>
,
<date id="D10E1032FFA0736FFEF7FAE6F5AC6427" box="[323,462,1299,1326]" pageId="67" pageNumber="68" value="1964-03-08">
<collectingDate id="C14AE9DAFFA0736FFEF7FAE6F5AC6427" box="[323,462,1299,1326]" pageId="67" pageNumber="68" value="1964-03-08">8 Mar. 1964</collectingDate>
</date>
,
<collectorName id="08455324FFA0736FFE6DFAE1F6246427" box="[473,582,1299,1326]" pageId="67" pageNumber="68">J. Bohlke</collectorName>
,
<collectorName id="08455324FFA0736FFDE5FAE1F6B16427" box="[593,723,1299,1326]" pageId="67" pageNumber="68">D. Dockins</collectorName>
,
<collectorName id="08455324FFA0736FFD6AFAE1F7186427" box="[734,890,1299,1326]" pageId="67" pageNumber="68">R. Rosenblatt</collectorName>
,
<collectorName id="08455324FFA0736FFC30FAE1F7966427" box="[900,1012,1299,1326]" pageId="67" pageNumber="68">W. Starck</collectorName>
,
<collectorName id="08455324FFA0736FFBB4FAE1F0316427" box="[1024,1107,1299,1326]" pageId="67" pageNumber="68">J. Tyler</collectorName>
&amp;
<collectorName id="08455324FFA0736FFBC3FAE1F0A56427" box="[1143,1223,1299,1326]" pageId="67" pageNumber="68">K. Hill</collectorName>
, coll.
</materialsCitation>
</paragraph>
<paragraph id="A50F36F2FFA0736FFF73FAC3F72D6747" blockId="67.[151,1437,1154,2046]" pageId="67" pageNumber="68">
<emphasis id="97C4EAE0FFA0736FFF73FAC3F5D1645B" bold="true" box="[199,435,1334,1361]" pageId="67" pageNumber="68">Additional material</emphasis>
.
<emphasis id="97C4EAE0FFA0736FFE75FAC3F634645B" bold="true" box="[449,598,1334,1361]" pageId="67" pageNumber="68">Arabian Sea</emphasis>
. One specimen (
<collectionCode id="C3A1AE37FFA0736FFCA6FAC2F70B645B" box="[786,873,1335,1361]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="67" pageNumber="68">MNHN</collectionCode>
A71a), Mission Gravier Djibouti, Recif de la Menagerie, in corals,
<date id="D10E1032FFA0736FFED6FAA9F583647F" box="[354,481,1371,1398]" pageId="67" pageNumber="68" value="1904-01-07">7 Jan. 1904</date>
[
<quantity id="62489B17FFA0736FFE5BFAAEF65F647C" box="[495,573,1371,1398]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="67" pageNumber="68" unit="mm" value="11.0">11 mm</quantity>
long,
<quantity id="62489B17FFA0736FFD36FAAEF6A0647C" box="[642,706,1371,1398]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="mm" value="2.0">2 mm</quantity>
wide]. Two specimens (
<collectionCode id="C3A1AE37FFA0736FFC79FAAEF046647F" box="[973,1060,1371,1397]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="67" pageNumber="68">MNHN</collectionCode>
A71b), Mission Gravier 1904, Île Moucha (
<geoCoordinate id="C0845035FFA0736FFEB6FA8AF50A6490" box="[258,360,1407,1434]" degrees="11" direction="north" minutes="43" orientation="latitude" pageId="67" pageNumber="68" precision="925" value="11.716666">11°43’ N</geoCoordinate>
,
<geoCoordinate id="C0845035FFA0736FFEC6FA8AF5B46490" box="[370,470,1407,1434]" degrees="43" direction="east" minutes="11" orientation="longitude" pageId="67" pageNumber="68" precision="925" value="43.183334">43°11’ E</geoCoordinate>
), Gulf of Tadjoura, Djibouti, dredged,
<quantity id="62489B17FFA0736FFC34FA8AF7D86490" box="[896,954,1407,1434]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="m" value="20.0">20 m</quantity>
,
<date id="D10E1032FFA0736FFC70FA8AF0386493" box="[964,1114,1407,1434]" pageId="67" pageNumber="68" value="1904-05-13">13 May 1904</date>
, C. Gravier, coll. [
<quantity id="62489B17FFA0736FFA96FA8AF1FF6490" box="[1314,1437,1407,1434]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="67" pageNumber="68" unit="mm" value="14.0" valueMax="18.0" valueMin="10.0">10–18 mm</quantity>
long,
<quantity id="62489B17FFA0736FFF63FA56F55664B4" box="[215,308,1443,1470]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="67" pageNumber="68" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
wide].
<emphasis id="97C4EAE0FFA0736FFE3CFA57F67964B7" bold="true" box="[392,539,1442,1469]" pageId="67" pageNumber="68">Gulf of Suez</emphasis>
. One specimen (
<collectionCode id="C3A1AE37FFA0736FFD62FA56F74F64B7" box="[726,813,1443,1469]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="67" pageNumber="68">MNHN</collectionCode>
A360), Mission R.P. Dollfus, Sta. 11,
<date id="D10E1032FFA0736FFB6DFA56F10664B7" box="[1241,1380,1443,1470]" pageId="67" pageNumber="68" value="1928-12-08">8 Dec. 1928</date>
, R.P. Dollfus, coll. [
<quantity id="62489B17FFA0736FFE8EFA33F5F164E8" box="[314,403,1478,1506]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="67" pageNumber="68" unit="mm" value="8.5">8.5 mm</quantity>
long,
<quantity id="62489B17FFA0736FFE6FFA33F65064E8" box="[475,562,1478,1506]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="67" pageNumber="68" unit="mm" value="1.5">1.5 mm</quantity>
wide]. One specimen (
<collectionCode id="C3A1AE37FFA0736FFC8CFA32F7ED64EB" box="[824,911,1479,1505]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="67" pageNumber="68">MNHN</collectionCode>
A372), R/V
<taxonomicName id="62B04D71FFA0736FFB96FA32F0DA64EB" authority=", Sta." authorityName="Sta." box="[1058,1208,1479,1505]" class="Liliopsida" family="Orchidaceae" genus="Calypso" kingdom="Plantae" order="Asparagales" pageId="67" pageNumber="68" phylum="Tracheophyta" rank="genus">Calypso, Sta.</taxonomicName>
unknown (probably Southern Saudi Arabian coast), Drach, Cherbonnier &amp; Mercier, coll. [
<quantity id="62489B17FFA0736FFC2DFA1EF7B8670C" box="[921,986,1515,1542]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="67" pageNumber="68" unit="mm" value="6.0">6 mm</quantity>
long,
<quantity id="62489B17FFA0736FFBABFA1EF03D670C" box="[1055,1119,1515,1542]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="67" pageNumber="68" unit="mm" value="1.0">1 mm</quantity>
wide].
<emphasis id="97C4EAE0FFA0736FFB05FA1FF093670F" bold="true" box="[1201,1265,1514,1541]" pageId="67" pageNumber="68">
<collectingCountry id="DDA77662FFA0736FFB05FA1FF093670F" box="[1201,1265,1514,1541]" name="India" pageId="67" pageNumber="68">India</collectingCountry>
</emphasis>
. One specimen (
<collectionCode id="C3A1AE37FFA0736FFF2BF9FAF4946723" box="[159,246,1551,1577]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="67" pageNumber="68">MNHN</collectionCode>
A419), Nicobar Islands, R/VInvestigator, Indian Survey, Sta. 593, Paway Island (Pahua,
<geoCoordinate id="C0845035FFA0736FFB6AF9FAF1246720" box="[1246,1350,1551,1578]" degrees="07" direction="north" minutes="22" orientation="latitude" pageId="67" pageNumber="68" precision="925" value="7.366667">07°22’ N</geoCoordinate>
,
<geoCoordinate id="C0845035FFA0736FFAE6F9FAF4C96744" degrees="93" direction="east" minutes="37" orientation="longitude" pageId="67" pageNumber="68" precision="925" value="93.61667">93°37’ E</geoCoordinate>
), Bay of Bengal, shore collecting [
<quantity id="62489B17FFA0736FFD84F9C6F6E36744" box="[560,641,1587,1614]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="67" pageNumber="68" unit="mm" value="12.0">12 mm</quantity>
long,
<quantity id="62489B17FFA0736FFD73F9C6F7636747" box="[711,769,1587,1613]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="mm" value="2.0">2mm</quantity>
wide].
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFA07368FF73F9A3F7B764E7" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68" type="description">
<paragraph id="A50F36F2FFA0736FFF73F9A3F13867D7" blockId="67.[151,1437,1154,2046]" pageId="67" pageNumber="68">
<emphasis id="97C4EAE0FFA0736FFF73F9A3F532677B" bold="true" box="[199,336,1622,1649]" pageId="67" pageNumber="68">Description</emphasis>
.
<typeStatus id="7A0B8850FFA0736FFED4F9A2F5A5677B" box="[352,455,1623,1649]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
(USNM 37642), complete, slightly damaged, bent, sigmoid, middle and posterior cirri missing. Body obconic, blunt anteriorly, tapered posteriorly (
<figureCitation id="3D8B2A77FFA0736FFC87F98EF7FA679F" box="[819,920,1659,1685]" captionStart="FIGURE 38" captionStartId="68.[151,250,988,1013]" captionTargetBox="[173,1417,184,959]" captionTargetId="figure@68.[170,1418,181,962]" captionTargetPageId="68" captionText="FIGURE 38. Leocrates rizzoae n. sp., holotype (USNM 37642). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, ventral view. D. Chaetiger 10, right parapodium, anterior view, MGS. Scale bars: A: 0.7 mm, B: 0.5 mm, C: 0.6 mm, D: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3672626" httpUri="https://zenodo.org/record/3672626/files/figure.png" pageId="67" pageNumber="68">Fig. 38A</figureCitation>
),
<quantity id="62489B17FFA0736FFC1FF98EF79A679C" box="[939,1016,1659,1686]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="67" pageNumber="68" unit="mm" value="14.0">14 mm</quantity>
long,
<quantity id="62489B17FFA0736FFB88F98FF0ED679C" box="[1084,1167,1658,1686]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="67" pageNumber="68" unit="mm" value="2.5">2.5 mm</quantity>
wide, 16 chaetigers; left parapodium of chaetiger 9 previously removed (not in container), right parapodium of chaetiger 10 dissected (kept in container). Tentacular and anterior dorsal cirri almost complete, others partially broken or missing. Body pale.
</paragraph>
<paragraph id="A50F36F2FFA0736FFF73F912F10F6640" blockId="67.[151,1437,1154,2046]" pageId="67" pageNumber="68">
Prostomium longer than wide, slightly wider anteriorly (
<figureCitation id="3D8B2A77FFA0736FFC88F912F7C0660B" box="[828,930,1767,1793]" captionStart="FIGURE 38" captionStartId="68.[151,250,988,1013]" captionTargetBox="[173,1417,184,959]" captionTargetId="figure@68.[170,1418,181,962]" captionTargetPageId="68" captionText="FIGURE 38. Leocrates rizzoae n. sp., holotype (USNM 37642). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, ventral view. D. Chaetiger 10, right parapodium, anterior view, MGS. Scale bars: A: 0.7 mm, B: 0.5 mm, C: 0.6 mm, D: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3672626" httpUri="https://zenodo.org/record/3672626/files/figure.png" pageId="67" pageNumber="68">Fig. 38B</figureCitation>
). Lateral antennae with ceratophores distinct, antennae directed ventrally, almost as long as prostomium, slightly longer than palps; palpophores twice longer than palpostyles. Median antenna broken, probably not reaching anterior prostomial margin, inserted between eyes.
</paragraph>
<paragraph id="A50F36F2FFA0736FFF73F8A1F7F06698" blockId="67.[151,1437,1154,2046]" pageId="67" pageNumber="68">
Eyes brownish, anterior eyes each
<superScript id="52C59BBAFFA0736FFDE3F8A7F63D666A" attach="right" box="[599,607,1874,1888]" fontSize="6" pageId="67" pageNumber="68">1</superScript>
/
<subScript id="393434B7FFA0736FFDD3F891F60D6678" attach="left" box="[615,623,1892,1906]" fontSize="6" pageId="67" pageNumber="68">6</subScript>
prostomial width, twice larger than posterior ones, slightly emarginate anterolaterally, more distant to each other than posterior round eyes.
</paragraph>
<paragraph id="A50F36F2FFA0736FFF73F86EF03566F4" blockId="67.[151,1437,1154,2046]" pageId="67" pageNumber="68">Nuchal organs lobes horizontal C-shaped, completely concealed by anterior margin of tentacular belt; lateral ciliated bands narrow, not visible dorsally. Tentacular cirri without tips, longer ones reaching chaetiger 8. Lateral cushions low, entire; longitudinal striae not visible (probably by integument damage).</paragraph>
<caption id="F1CF667AFFA77368FF23FC29F5046536" ID-DOI="http://doi.org/10.5281/zenodo.3672626" ID-Zenodo-Dep="3672626" httpUri="https://zenodo.org/record/3672626/files/figure.png" pageId="68" pageNumber="69" startId="68.[151,250,988,1013]" targetBox="[173,1417,184,959]" targetPageId="68">
<paragraph id="A50F36F2FFA77368FF23FC29F5046536" blockId="68.[151,1437,988,1085]" pageId="68" pageNumber="69">
<emphasis id="97C4EAE0FFA77368FF23FC29F57B62FF" bold="true" box="[151,281,988,1013]" pageId="68" pageNumber="69">FIGURE 38</emphasis>
.
<taxonomicName id="62B04D71FFA77368FE92FC28F5B862FE" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[294,474,989,1012]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="68" pageNumber="69" phylum="Annelida" rank="species" species="rizzoae" status="sp. nov.">
<emphasis id="97C4EAE0FFA77368FE92FC28F5B862FE" box="[294,474,989,1012]" italics="true" pageId="68" pageNumber="69">Leocrates rizzoae</emphasis>
</taxonomicName>
<emphasis id="97C4EAE0FFA77368FE54FC28F67B62FF" bold="true" box="[480,537,989,1013]" pageId="68" pageNumber="69">
<taxonomicNameLabel id="8CF7579BFFA77368FE54FC28F67B62FF" box="[480,537,989,1013]" pageId="68" pageNumber="69" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (USNM 37642). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, ventral view. D. Chaetiger 10, right parapodium, anterior view, MGS. Scale bars: A: 0.7 mm, B: 0.5 mm, C: 0.6 mm, D: 0.2 mm.
</paragraph>
</caption>
<paragraph id="A50F36F2FFA77368FF73FB9EF52865C7" blockId="68.[151,1437,1131,2022]" pageId="68" pageNumber="69">
Pharynx barely exposed (
<figureCitation id="3D8B2A77FFA77368FE52FB9EF62F658F" box="[486,589,1131,1157]" captionStart="FIGURE 38" captionStartId="68.[151,250,988,1013]" captionTargetBox="[173,1417,184,959]" captionTargetId="figure@68.[170,1418,181,962]" captionTargetPageId="68" captionText="FIGURE 38. Leocrates rizzoae n. sp., holotype (USNM 37642). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, ventral view. D. Chaetiger 10, right parapodium, anterior view, MGS. Scale bars: A: 0.7 mm, B: 0.5 mm, C: 0.6 mm, D: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3672626" httpUri="https://zenodo.org/record/3672626/files/figure.png" pageId="68" pageNumber="69">Fig. 38C</figureCitation>
), jaws observed through previous dissection. Lateral vesicles not seen. Anterior margin with irregular constrictions. Dorsal and ventral jaws brownish, exposed, tapered, ventral jaw smaller than dorsal one.
</paragraph>
<paragraph id="A50F36F2FFA77368FF73FB22F108648B" blockId="68.[151,1437,1131,2022]" pageId="68" pageNumber="69">
Dorsal cirri as long as body width (including parapodia). Chaetigers 1–4 without notochaetae; notochaetae present along chaetigers 5–16, about 50 per bundle, delicate, some arranged in bundles, notochaetae subdistally denticulate, denticles fine. Notacicular lobes tapered; neuracicular lobes triangular, blunt, slightly longer than wide (
<figureCitation id="3D8B2A77FFA77368FADBFAEAF4A96457" captionStart="FIGURE 38" captionStartId="68.[151,250,988,1013]" captionTargetBox="[173,1417,184,959]" captionTargetId="figure@68.[170,1418,181,962]" captionTargetPageId="68" captionText="FIGURE 38. Leocrates rizzoae n. sp., holotype (USNM 37642). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Anterior end, ventral view. D. Chaetiger 10, right parapodium, anterior view, MGS. Scale bars: A: 0.7 mm, B: 0.5 mm, C: 0.6 mm, D: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3672626" httpUri="https://zenodo.org/record/3672626/files/figure.png" pageId="68" pageNumber="69">Fig. 38D</figureCitation>
). Ventral cirri as long as neurochaetal lobe. Neurochaetae about 20 per bundle, blades bidentate, most missing, decreasing in size ventrally, remaining blades 5–9 times longer than wide, guards approaching subdistal tooth.
</paragraph>
<paragraph id="A50F36F2FFA77368FF73FA7EF6D664C3" blockId="68.[151,1437,1131,2022]" pageId="68" pageNumber="69">Posterior region tapered, almost without cirri. Preanal segment with dorsal cirri twice longer than ventral ones. Pygidium with anus terminal, anal cirri missing.</paragraph>
<paragraph id="A50F36F2FFA77368FF73FA26F7B764E7" blockId="68.[151,1437,1131,2022]" box="[199,981,1491,1517]" pageId="68" pageNumber="69">Oocytes not seen. Gonad fragments are testis; no oocytes were found.</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFA77368FF73FA02F5436777" pageId="68" pageNumber="69" type="etymology">
<paragraph id="A50F36F2FFA77368FF73FA02F5436777" blockId="68.[151,1437,1131,2022]" pageId="68" pageNumber="69">
<emphasis id="97C4EAE0FFA77368FF73FA02F52A671B" bold="true" box="[199,328,1527,1553]" pageId="68" pageNumber="69">Etymology</emphasis>
. This species name is after Alexandra Elaine Rizzo from the
<collectingRegion id="6774F810FFA77368FBA3FA02F162671B" box="[1047,1280,1527,1553]" country="Brazil" name="Rio de Janeiro" pageId="68" pageNumber="69">Rio de Janeiro State</collectingRegion>
University, a much-appreciated Brazilian colleague, and in recognition of her research activities, especially as a specialist of several polychaete groups, including
<taxonomicName id="62B04D71FFA77368FDA2F9CAF6FA6753" authorityName="Grube" authorityYear="1850" box="[534,664,1599,1625]" class="Polychaeta" family="Hesionidae" kingdom="Animalia" order="Phyllodocida" pageId="68" pageNumber="69" phylum="Annelida" rank="family">Hesionidae</taxonomicName>
. The species-group name is a noun in the genitive case (
<bibRefCitation id="C1214B03FFA77368FAA6F9CAF1FA6753" author="ICZN" box="[1298,1432,1599,1625]" firstAuthor="ICZN" pageId="68" pageNumber="69" refId="ref85916" refString="ICZN (International Commission on Zoological Nomenclature). (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature in association with the British Museum (Natural History), London, 306 pp. [http: // www. iczn. org / iczn / index. jsp]" type="book" year="1999">ICZN 1999</bibRefCitation>
,
<collectingRegion id="6774F810FFA77368FF23F996F4A26777" box="[151,192,1635,1661]" country="Djibouti" name="Arta" pageId="68" pageNumber="69">Art</collectingRegion>
. 31.1.2).
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFA77368FF73F973F66E66EC" pageId="68" pageNumber="69" type="description">
<paragraph id="A50F36F2FFA77368FF73F973F66E66EC" blockId="68.[151,1437,1131,2022]" pageId="68" pageNumber="69">
<emphasis id="97C4EAE0FFA77368FF73F973F55A67AB" bold="true" box="[199,312,1670,1697]" pageId="68" pageNumber="69">Variation</emphasis>
. The
<typeStatus id="7A0B8850FFA77368FEC9F972F5BF67AB" box="[381,477,1671,1697]" pageId="68" pageNumber="69" type="paratype">paratype</typeStatus>
(USNM 37641) is complete, anterior half depressed, probably by labels, most cirri and chaetal blades missing; anteroventral dissection along mouth to chaetiger 4 previously made, left parapodia of chaetigers 4 and 7 previously removed (missing), right parapodium of chaetiger 10 dissected (kept in container). Body with parallel sides, blunt anteriorly, tapered posteriorly (
<figureCitation id="3D8B2A77FFA77368FCEDF906F7A06607" box="[857,962,1779,1805]" captionStart="FIGURE 39" captionStartId="69.[151,250,1048,1073]" captionTargetBox="[172,1416,527,1021]" captionTargetId="figure@69.[170,1418,525,1023]" captionTargetPageId="69" captionText="FIGURE 39. Leocrates rizzoae n. sp., paratype (USNM 37641). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetiger 10, right parapodium, anterior view, MGS (insets: upper and lower neurochaetal blades, notacicular lobe). Scale bars: A: 0.8 mm, B, C: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3672628" httpUri="https://zenodo.org/record/3672628/files/figure.png" pageId="68" pageNumber="69">Fig. 39A</figureCitation>
),
<quantity id="62489B17FFA77368FC63F906F04A6607" box="[983,1064,1779,1805]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="68" pageNumber="69" unit="mm" value="13.0">13 mm</quantity>
long,
<quantity id="62489B17FFA77368FBC4F906F0D06607" box="[1136,1202,1779,1805]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="68" pageNumber="69" unit="mm" value="2.0">2 mm</quantity>
wide, 16 chaetigers. Prostomium longer than wide, barely wider anteriorly (
<figureCitation id="3D8B2A77FFA77368FCBEF8E2F713663B" box="[778,881,1815,1841]" captionStart="FIGURE 39" captionStartId="69.[151,250,1048,1073]" captionTargetBox="[172,1416,527,1021]" captionTargetId="figure@69.[170,1418,525,1023]" captionTargetPageId="69" captionText="FIGURE 39. Leocrates rizzoae n. sp., paratype (USNM 37641). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetiger 10, right parapodium, anterior view, MGS (insets: upper and lower neurochaetal blades, notacicular lobe). Scale bars: A: 0.8 mm, B, C: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3672628" httpUri="https://zenodo.org/record/3672628/files/figure.png" pageId="68" pageNumber="69">Fig. 39B</figureCitation>
); left lateral antenna slightly shorter than prostomium, slightly longer than left palp; palpophores twice longer than palpostyles. Eyes dark brown, anterior ones twice larger than posterior ones, slightly farther apart and anterolaterally emarginate. Notochaetae from chaetiger 5, about 50 per bundle, finely denticulate subdistally; notacicular lobes tapered, neuracicular lobes conical, blunt (
<figureCitation id="3D8B2A77FFA77368FF14F852F56566CB" box="[160,263,1959,1985]" captionStart="FIGURE 39" captionStartId="69.[151,250,1048,1073]" captionTargetBox="[172,1416,527,1021]" captionTargetId="figure@69.[170,1418,525,1023]" captionTargetPageId="69" captionText="FIGURE 39. Leocrates rizzoae n. sp., paratype (USNM 37641). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetiger 10, right parapodium, anterior view, MGS (insets: upper and lower neurochaetal blades, notacicular lobe). Scale bars: A: 0.8 mm, B, C: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3672628" httpUri="https://zenodo.org/record/3672628/files/figure.png" pageId="68" pageNumber="69">Fig. 39C</figureCitation>
); neurochaetae about 30 per bundle; blades bidentate, 4–16 times longer than wide, guards approaching subdistal tooth (
<figureCitation id="3D8B2A77FFA77368FEFFF83EF5D366EF" box="[331,433,1995,2021]" captionStart="FIGURE 39" captionStartId="69.[151,250,1048,1073]" captionTargetBox="[172,1416,527,1021]" captionTargetId="figure@69.[170,1418,525,1023]" captionTargetPageId="69" captionText="FIGURE 39. Leocrates rizzoae n. sp., paratype (USNM 37641). A. Anterior region, dorsal view. B. Anterior end, dorsal view. C. Chaetiger 10, right parapodium, anterior view, MGS (insets: upper and lower neurochaetal blades, notacicular lobe). Scale bars: A: 0.8 mm, B, C: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3672628" httpUri="https://zenodo.org/record/3672628/files/figure.png" pageId="68" pageNumber="69">Fig. 39C</figureCitation>
, insets).
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFA67369FF73FF6CF51B60A5" pageId="69" pageNumber="70" type="discussion">
<paragraph id="A50F36F2FFA67369FF73FF6CF51B60A5" blockId="69.[151,1437,153,503]" pageId="69" pageNumber="70">
<emphasis id="97C4EAE0FFA67369FF73FF6CF55661B9" bold="true" box="[199,308,153,179]" pageId="69" pageNumber="70">Remarks</emphasis>
.
<taxonomicName id="62B04D71FFA67369FEF7FF6FF66861B9" authority="Salazar-Vallejo, 2020" authorityName="Salazar-Vallejo" authorityYear="2020" box="[323,522,154,179]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="69" pageNumber="70" phylum="Annelida" rank="species" species="rizzoae" status="sp. nov.">
<emphasis id="97C4EAE0FFA67369FEF7FF6FF66861B9" box="[323,522,154,179]" italics="true" pageId="69" pageNumber="70">Leocrates rizzoae</emphasis>
</taxonomicName>
<emphasis id="97C4EAE0FFA67369FDA5FF6CF63361B9" bold="true" box="[529,593,153,179]" pageId="69" pageNumber="70">
<taxonomicNameLabel id="8CF7579BFFA67369FDA5FF6CF63361B9" box="[529,593,153,179]" pageId="69" pageNumber="70" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="62B04D71FFA67369FD64FF6FF7CA61B9" authority="Grube, 1878" authorityName="Grube" authorityYear="1878" box="[720,936,153,179]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="69" pageNumber="70" phylum="Annelida" rank="species" species="iris">
<emphasis id="97C4EAE0FFA67369FD64FF6FF77661B9" box="[720,788,154,179]" italics="true" pageId="69" pageNumber="70">L. iris</emphasis>
<bibRefCitation id="C1214B03FFA67369FCA8FF6CF7CA61B9" author="Grube" box="[796,936,153,179]" firstAuthor="Grube" pageId="69" pageNumber="70" pagination="1 - 300" refId="ref84847" refString="Grube, A. E. (1878) Annulata Semperiana. Beitrage zur Kenntnis der Annelidenfauna der Philippinen nach den von Herrn Prof. Semper mitgebrachten Sammlungen. Memoires de l'Academie Imperiale des Sciences de St. - Petersbourg, Serie 7, 25, 8, 1 - 300, pls. 1 - 15. https: // doi. org / 10.5962 / bhl. title. 85345" type="journal article" year="1878">Grube, 1878</bibRefCitation>
</taxonomicName>
from the
<collectingCountry id="DDA77662FFA67369FBACFF6CF0F461B9" box="[1048,1174,153,179]" name="Philippines" pageId="69" pageNumber="70">Philippines</collectingCountry>
and
<collectingCountry id="DDA77662FFA67369FB79FF6CF17E61B9" box="[1229,1308,153,179]" name="Samoa" pageId="69" pageNumber="70">Samoa</collectingCountry>
, reinstated, because they have large eyes, anterior eyes larger than posterior ones, notacicular lobes entire, three times longer than wide, and middle chaetigers with about 30 neurochaetae per bundle. These two species differ in the shape of anterior eyes, and by the size proportions of palpophore to palpostyles; in
<taxonomicName id="62B04D71FFA67369FC6BFEF3F02D6015" authorityName="Salazar-Vallejo" authorityYear="2020" box="[991,1103,262,287]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="69" pageNumber="70" phylum="Annelida" rank="species" species="rizzoae">
<emphasis id="97C4EAE0FFA67369FC6BFEF3F02D6015" box="[991,1103,262,287]" italics="true" pageId="69" pageNumber="70">L. rizzoae</emphasis>
</taxonomicName>
anterior eyes are emarginate, and palpophores are twice larger than palpostyles, whereas in
<taxonomicName id="62B04D71FFA67369FCF3FEDFF7EB6049" authorityName="Grube" authorityYear="1878" box="[839,905,298,323]" class="Polychaeta" family="Hesionidae" genus="Leocrates" kingdom="Animalia" order="Phyllodocida" pageId="69" pageNumber="70" phylum="Annelida" rank="species" species="iris">
<emphasis id="97C4EAE0FFA67369FCF3FEDFF7EB6049" box="[839,905,298,323]" italics="true" pageId="69" pageNumber="70">L. iris</emphasis>
</taxonomicName>
anterior eyes are reniform, and palpophores are 2–3 times longer than palpostyles. The inclusion of the Red Sea specimens is with hesitation because of the condition of the specimens (one dried out, the other collapsed), and better specimens will help clarify its northwestern distributional range.
</paragraph>
</subSubSection>
<subSubSection id="EDAA6579FFA67369FF73FE4DF69C60FD" pageId="69" pageNumber="70" type="distribution">
<paragraph id="A50F36F2FFA67369FF73FE4DF69C60FD" blockId="69.[151,1437,153,503]" pageId="69" pageNumber="70">
<emphasis id="97C4EAE0FFA67369FF73FE4DF53B60D9" bold="true" box="[199,345,440,467]" pageId="69" pageNumber="70">Distribution</emphasis>
. Arabian Sea, the
<collectingCountry id="DDA77662FFA67369FD90FE4CF6FD60D9" box="[548,671,441,467]" name="Seychelles" pageId="69" pageNumber="70">Seychelles</collectingCountry>
, and
<collectingCountry id="DDA77662FFA67369FD6CFE4CF7E960D9" box="[728,907,441,467]" name="India" pageId="69" pageNumber="70">Nicobar Islands</collectingCountry>
, Indian Ocean, (Red Sea?), in intertidal to
<quantity id="62489B17FFA67369FAD0FE4CF1FE60D9" box="[1380,1436,441,467]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="69" pageNumber="70" unit="m" value="30.0">30 m</quantity>
depth in mixed bottoms, including coralline substrates.
</paragraph>
</subSubSection>
</treatment>
</document>
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