238 lines
31 KiB
XML
238 lines
31 KiB
XML
<document id="8D3E79B9145709540F1FFD880F375906" ID-CLB-Dataset="45374" ID-DOI="10.11646/zootaxa.4103.2.12" ID-GBIF-Dataset="fad60b2b-5a7b-4191-9609-8e5c778ca3ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="262498" ID-ZooBank="AC27CA02-845A-44D1-9937-BD4FF6D3095A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460444087176" checkinUser="plazi" docAuthor="Lozouet, Pierre & Krygelmans, Anouchka" docDate="2016" docId="03A087DBFFCFFFBAFF15FB0648DE3F6F" docLanguage="en" docName="zootaxa.4103.2.12.pdf.imf" docOrigin="Zootaxa 4103 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Modulidae P. Fischer 1884" docType="treatment" docVersion="11" lastPageNumber="196" masterDocId="FF99FFA3FFCFFFBBFF82FF964D5A3E10" masterDocTitle="A new species of Indo-Pacific Modulidae (Mollusca: Caenogastropoda)" masterLastPageNumber="200" masterPageNumber="195" pageNumber="195" updateTime="1698333199998" updateUser="ExternalLinkService">
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<mods:mods id="C992032490D65D62CD7A17FBFC1BD317" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FE6770CDD131FBFA0CAF39CB211C4F91">
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<mods:title id="EE559EB633F806CB6E972461295226AD">A new species of Indo-Pacific Modulidae (Mollusca: Caenogastropoda)</mods:title>
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<mods:name id="A41D184AB28FE14F4504654DD0DDDE76" type="personal">
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<mods:namePart id="266BE4472835E64D2CEF52312AA9CBAE">Lozouet, Pierre</mods:namePart>
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<mods:name id="7984B208EB877E6A82E381A85E72A367" type="personal">
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<mods:roleTerm id="DB127CFF738BEF14F7EDA28DA40A344F">Author</mods:roleTerm>
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<mods:namePart id="5A1F2E5F24E482205EB0582680567DD2">Krygelmans, Anouchka</mods:namePart>
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<mods:typeOfResource id="1A5AD0EB6CC31C11CDA776DCA1EADE63">text</mods:typeOfResource>
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<mods:title id="D9F3FB7643FA8D45825864C848658291">Zootaxa</mods:title>
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<mods:part id="8E2D7728A44357B8819C6581B3D41ECA">
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<mods:date id="CD09D78D8B95199BB4612DF029CFCFC3">2016</mods:date>
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<mods:number id="D52B16FA022F7239C8171D821EBF4343">4103</mods:number>
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<mods:number id="828AC422C9677C42861801C576A78D64">2</mods:number>
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<mods:classification id="E228A61A4EF42045015F78F6FF76E8B2">journal article</mods:classification>
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<mods:identifier id="E8B11FC0142E30A32D0CC60D2E78A9BD" type="DOI">10.11646/zootaxa.4103.2.12</mods:identifier>
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<treatment id="03A087DBFFCFFFBAFF15FB0648DE3F6F" ID-DOI="http://doi.org/10.5281/zenodo.6080861" ID-GBIF-Taxon="119508997" ID-Zenodo-Dep="6080861" LSID="urn:lsid:plazi:treatment:03A087DBFFCFFFBAFF15FB0648DE3F6F" httpUri="http://treatment.plazi.org/id/03A087DBFFCFFFBAFF15FB0648DE3F6F" lastPageId="1" lastPageNumber="196" pageId="0" pageNumber="195">
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<subSubSection id="C3136546FFCFFFBBFF15FB064FEB3B1F" pageId="0" pageNumber="195" type="nomenclature">
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<paragraph id="8BB636CDFFCFFFBBFF15FB064F1C3ABA" blockId="0.[151,1436,1168,2023]" box="[151,582,1168,1194]" pageId="0" pageNumber="195">
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<heading id="D0FE81A1FFCFFFBBFF15FB064F1C3ABA" bold="true" box="[151,582,1168,1194]" fontSize="11" level="1" pageId="0" pageNumber="195" reason="1">
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<emphasis id="B97DEADFFFCFFFBBFF15FB064F1C3ABA" bold="true" box="[151,582,1168,1194]" pageId="0" pageNumber="195">
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Family
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<taxonomicName id="4C094D4EFFCFFFBBFF74FB064F1C3ABA" ID-CoL="7NJTM" authority="P. Fischer, 1884" authorityName="P. Fischer" authorityYear="1884" box="[246,582,1168,1194]" class="Gastropoda" family="Modulidae" kingdom="Animalia" pageId="0" pageNumber="195" phylum="Mollusca" rank="family">Modulidae P. Fischer, 1884</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB636CDFFCFFFBBFF15FB244FEB3B1F" blockId="0.[151,1436,1168,2023]" pageId="0" pageNumber="195">
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<bibRefCitation id="EF984B3CFFCFFFBBFF15FB244CDB3AD9" box="[151,385,1202,1225]" pageId="0" pageNumber="195" refString="Landau, B. Vermeij, G. J. & Reich, S. (2014) Classification of the Modulidae (Caenogastropoda, Cerithioidea), with new genera and new fossil species from the Neogene of tropical America and Indonesia. Basteria, 78 (1 - 3), 1 - 29." type="journal article">
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Landau
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<emphasis id="B97DEADFFFCFFFBBFF70FB244C7F3AD9" box="[242,293,1202,1225]" italics="true" pageId="0" pageNumber="195">et al</emphasis>
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. (2014)
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</bibRefCitation>
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, working on both Recent and fossil taxa, divided the family
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<taxonomicName id="4C094D4EFFCFFFBBFBB4FB2449F33AD9" box="[1078,1193,1202,1225]" class="Gastropoda" family="Modulidae" kingdom="Animalia" pageId="0" pageNumber="195" phylum="Mollusca" rank="family">Modulidae</taxonomicName>
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into six genera. This classification is based on the general morphology of the shell. The aperture, and in particular the siphonal area (
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<figureCitation id="13322A48FFCFFFBBFAD6FB4348CC3AFC" box="[1364,1430,1237,1260]" captionStart="FIGURE 2" captionStartId="2.[151,244,1806,1828]" captionTargetBox="[161,1418,868,1770]" captionTargetId="figure@2.[151,1436,850,1785]" captionTargetPageId="2" captionText="FIGURE 2. Outlines of recent and fossil Modulidae. A. Indomodulus tectum, New Caledonia, Touho, Montrouzier expedition, sta. 1246, 20 ° 48, 8 ’ S, 165 ° 08.7 ' E [MNHN-IM- 2012 - 36610]. B. Modulus turbinoides (Locard, 1897) Cape Verde, Calypso 1959, sta. 65, 15 ° 37 ’ N, 23 ° 09 ’ W [MNHN-IM- 2012 - 36609]. C. Modulus modulus (Linnaeus, 1758), Sanibel I., Florida, coll. Staadt [MNHN-IM- 2012 - 36612]. D. Conomodulus neocaledonensis sp. nov., holotype, New Caledonia, Ouen I., Lagon campaign, Sta. 243, 22 ° 24 S, 167 ° 01 E [MNHN-IM- 2000 - 31117]. E. Conomodulus preangerensis (K. Martin, 1905), late Miocene, early Tortonian of Bontang, East Kalimantan, Indonesia (locality TF 505) [RGM. 794.179]. F. Incisilabium parisiensis (Deshayes, 1832), Middle Eocene, Bartonian, Paris Basin, St-Vaast-lès-Mello (Barisseuse), coll. Pacaud [MNHN. F. A 51573]. The arrows indicate the siphonal depression, reinforced in B and C." httpUri="https://zenodo.org/record/262500/files/figure.png" pageId="0" pageNumber="195">
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Fig.
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<docIssue id="CF6A4922FFCFFFBBFA05FB4348CC3AFC" box="[1415,1430,1237,1260]" pageId="0" pageNumber="195">2</docIssue>
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</figureCitation>
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, arrows) is regarded as the main diagnostic feature.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3136546FFCFFFBAFF45FA8C48DE3F6F" lastPageId="1" lastPageNumber="196" pageId="0" pageNumber="195" type="reference_group">
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<paragraph id="8BB636CDFFCFFFBBFF45FA8C4CF43879" blockId="0.[151,1436,1168,2023]" pageId="0" pageNumber="195">
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<taxonomicName id="4C094D4EFFCFFFBBFF45FA8C4F693B21" ID-CoL="7P6S7" authority="Landau et al., 2014" authorityName="Landau et al." authorityYear="2014" box="[199,563,1306,1329]" class="Gastropoda" family="Modulidae" genus="Laevimodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
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<emphasis id="B97DEADFFFCFFFBBFF45FA8C4C073B21" box="[199,349,1306,1329]" italics="true" pageId="0" pageNumber="195">Laevimodulus</emphasis>
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<bibRefCitation id="EF984B3CFFCFFFBBFEE6FA8C4F693B21" author="Landau" box="[356,563,1306,1329]" pageId="0" pageNumber="195" refString="Landau, B. Vermeij, G. J. & Reich, S. (2014) Classification of the Modulidae (Caenogastropoda, Cerithioidea), with new genera and new fossil species from the Neogene of tropical America and Indonesia. Basteria, 78 (1 - 3), 1 - 29." type="journal article" year="2014">
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Landau
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<emphasis id="B97DEADFFFCFFFBBFE39FA8C4CB03B21" box="[443,490,1306,1329]" italics="true" pageId="0" pageNumber="195">et al</emphasis>
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., 2014
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</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C094D4EFFCFFFBBFDEAFA8C4ECF3B21" ID-CoL="7PDNY" authority="Collins, 1934" authorityName="Collins" authorityYear="1934" box="[616,917,1306,1329]" class="Gastropoda" family="Modulidae" genus="Psammodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
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<emphasis id="B97DEADFFFCFFFBBFDEAFA8C4FA73B21" box="[616,765,1306,1329]" italics="true" pageId="0" pageNumber="195">Psammodulus</emphasis>
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Collins, 1934
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</taxonomicName>
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are exclusively fossil tropical American genera, with several records in the Miocene-Pliocene.
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<taxonomicName id="4C094D4EFFCFFFBBFD11FAAB494F3B44" ID-CoL="7PK2S" authority="Landau et al., 2014" authorityName="Landau et al." authorityYear="2014" box="[659,1045,1341,1364]" class="Gastropoda" family="Modulidae" genus="Trochomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
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<emphasis id="B97DEADFFFCFFFBBFD11FAAB4E623B44" box="[659,824,1341,1364]" italics="true" pageId="0" pageNumber="195">Trochomodulus</emphasis>
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<bibRefCitation id="EF984B3CFFCFFFBBFCC2FAAB494F3B44" author="Landau" box="[832,1045,1341,1364]" pageId="0" pageNumber="195" refString="Landau, B. Vermeij, G. J. & Reich, S. (2014) Classification of the Modulidae (Caenogastropoda, Cerithioidea), with new genera and new fossil species from the Neogene of tropical America and Indonesia. Basteria, 78 (1 - 3), 1 - 29." type="journal article" year="2014">
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Landau
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||
<emphasis id="B97DEADFFFCFFFBBFC1BFAAB4E903B44" box="[921,970,1341,1364]" italics="true" pageId="0" pageNumber="195">et al</emphasis>
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||
., 2014
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</bibRefCitation>
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</taxonomicName>
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is another tropical American genus represented from the late Early Miocene to the present-day Caribbean and Panamic regions. The genus
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<taxonomicName id="4C094D4EFFCFFFBBFB7EFAF64D963B89" ID-CoL="7P95L" authority="Gray, 1842" authorityName="Gray" authorityYear="1842" class="Gastropoda" family="Modulidae" genus="Modulus" kingdom="Animalia" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
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<emphasis id="B97DEADFFFCFFFBBFB7EFAF648033B67" box="[1276,1369,1376,1399]" italics="true" pageId="0" pageNumber="195">Modulus</emphasis>
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Gray, 1842
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</taxonomicName>
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contains most of the recent and fossil Caribbean-Panamic species and also includes the West-African species.
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<taxonomicName id="4C094D4EFFCFFFBBFF15FA334C803BAC" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" box="[151,474,1445,1468]" class="Gastropoda" family="Trochidae" genus="Trochus" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="tectum">
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<emphasis id="B97DEADFFFCFFFBBFF15FA334C613BAC" box="[151,315,1445,1468]" italics="true" pageId="0" pageNumber="195">Trochus tectum</emphasis>
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Gmelin, 1791
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</taxonomicName>
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, the only species of
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<taxonomicName id="4C094D4EFFCFFFBBFD3FFA334E6A3BAC" ID-CoL="7NJTM" box="[701,816,1445,1468]" class="Gastropoda" family="Modulidae" kingdom="Animalia" pageId="0" pageNumber="195" phylum="Mollusca" rank="family">Modulidae</taxonomicName>
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in the Indo-Pacific, is also the
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<typeStatus id="54B2886FFFCFFFBBFB0BFA3349ED3BAC" box="[1161,1207,1445,1468]" pageId="0" pageNumber="195">type</typeStatus>
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species of the genus
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<taxonomicName id="4C094D4EFFCFFFBBFF15FA5E4F5D3BCF" ID-CoL="7P5RB" authority="Landau et al., 2014" authorityName="Landau et al." authorityYear="2014" box="[151,519,1480,1503]" class="Gastropoda" family="Modulidae" genus="Indomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
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<emphasis id="B97DEADFFFCFFFBBFF15FA5E4C783BCF" box="[151,290,1480,1503]" italics="true" pageId="0" pageNumber="195">Indomodulus</emphasis>
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<bibRefCitation id="EF984B3CFFCFFFBBFEAEFA5E4F5D3BCF" author="Landau" box="[300,519,1480,1503]" pageId="0" pageNumber="195" refString="Landau, B. Vermeij, G. J. & Reich, S. (2014) Classification of the Modulidae (Caenogastropoda, Cerithioidea), with new genera and new fossil species from the Neogene of tropical America and Indonesia. Basteria, 78 (1 - 3), 1 - 29." type="journal article" year="2014">
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Landau
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<emphasis id="B97DEADFFFCFFFBBFE04FA5E4CED3BCF" box="[390,439,1480,1503]" italics="true" pageId="0" pageNumber="195">et al</emphasis>
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., 2014
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</bibRefCitation>
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||
</taxonomicName>
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.
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<taxonomicName id="4C094D4EFFCFFFBBFD96FA5E4FAA3BCF" ID-CoL="3PND8" box="[532,752,1480,1503]" class="Gastropoda" family="Modulidae" genus="Indomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="tectum">
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<emphasis id="B97DEADFFFCFFFBBFD96FA5E4FAA3BCF" box="[532,752,1480,1503]" italics="true" pageId="0" pageNumber="195">Indomodulus tectum</emphasis>
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</taxonomicName>
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is widespread throughout the Indo-Pacific. However,
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<bibRefCitation id="EF984B3CFFCFFFBBFACAFA5E4D843811" author="Trondle" pageId="0" pageNumber="195" refString="Trondle, J. (1989) The Modulidae of French Polynesia. Xenophora, 46, 16 - 18." type="journal article" year="1989">Trondlé (1989)</bibRefCitation>
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argued that based principally on the shell colour of the aperture at least two species can be distinguished in
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<collectingCountry id="F31E765DFFCFFFBBFAD3FA7C4C593834" name="French Polynesia" pageId="0" pageNumber="195">French Polynesia</collectingCountry>
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:
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<taxonomicName id="4C094D4EFFCFFFBBFE8EF99B4CBF3834" ID-CoL="3PND8" box="[268,485,1549,1572]" class="Gastropoda" family="Modulidae" genus="Indomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="tectum">
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<emphasis id="B97DEADFFFCFFFBBFE8EF99B4CBF3834" box="[268,485,1549,1572]" italics="true" pageId="0" pageNumber="195">Indomodulus tectum</emphasis>
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||
</taxonomicName>
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and
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<taxonomicName id="4C094D4EFFCFFFBBFD9AF99B4E503834" ID-CoL="43TWW" box="[536,778,1549,1572]" class="Gastropoda" family="Modulidae" genus="Indomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="candidus">
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||
<emphasis id="B97DEADFFFCFFFBBFD9AF99B4E503834" box="[536,778,1549,1572]" italics="true" pageId="0" pageNumber="195">Indomodulus candidus</emphasis>
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||
</taxonomicName>
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||
(Petit de la Saussaye, 1853). A molecular analysis is urgently needed to clarify this systematic debate. The genus
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<taxonomicName id="4C094D4EFFCFFFBBFD39F9A649793857" ID-CoL="7P27V" authority="Landau et al., 2014" authorityName="Landau et al." authorityYear="2014" box="[699,1059,1584,1607]" class="Gastropoda" family="Modulidae" genus="Conomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
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||
<emphasis id="B97DEADFFFCFFFBBFD39F9A64E153857" box="[699,847,1584,1607]" italics="true" pageId="0" pageNumber="195">Conomodulus</emphasis>
|
||
<bibRefCitation id="EF984B3CFFCFFFBBFCD7F9A649793857" author="Landau" box="[853,1059,1584,1607]" pageId="0" pageNumber="195" refString="Landau, B. Vermeij, G. J. & Reich, S. (2014) Classification of the Modulidae (Caenogastropoda, Cerithioidea), with new genera and new fossil species from the Neogene of tropical America and Indonesia. Basteria, 78 (1 - 3), 1 - 29." type="journal article" year="2014">
|
||
Landau
|
||
<emphasis id="B97DEADFFFCFFFBBFC2EF9A64EBB3857" box="[940,993,1584,1607]" italics="true" pageId="0" pageNumber="195">et al.</emphasis>
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||
, 2014
|
||
</bibRefCitation>
|
||
</taxonomicName>
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||
is based on two fossil species from the Miocene of
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<collectingCountry id="F31E765DFFCFFFBBFEBDF9C44CF03879" box="[319,426,1618,1641]" name="Indonesia" pageId="0" pageNumber="195">Indonesia</collectingCountry>
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.
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||
</paragraph>
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<paragraph id="8BB636CDFFCFFFBBFF45F9E34E6338E4" blockId="0.[151,1436,1168,2023]" pageId="0" pageNumber="195">
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||
Most of the American species classified in the genera
|
||
<taxonomicName id="4C094D4EFFCFFFBBFC90F9E34E35389C" ID-CoL="7P95L" box="[786,879,1653,1676]" class="Gastropoda" family="Modulidae" genus="Modulus" kingdom="Animalia" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
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||
<emphasis id="B97DEADFFFCFFFBBFC90F9E34E35389C" box="[786,879,1653,1676]" italics="true" pageId="0" pageNumber="195">Modulus</emphasis>
|
||
</taxonomicName>
|
||
,
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<taxonomicName id="4C094D4EFFCFFFBBFCFFF9E3497B389C" ID-CoL="7PK2S" box="[893,1057,1653,1676]" class="Gastropoda" family="Modulidae" genus="Trochomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEADFFFCFFFBBFCFFF9E3497B389C" box="[893,1057,1653,1676]" italics="true" pageId="0" pageNumber="195">Trochomodulus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C094D4EFFCFFFBBFBDDF9E349AE389C" ID-CoL="7P6S7" box="[1119,1268,1653,1676]" class="Gastropoda" family="Modulidae" genus="Laevimodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEADFFFCFFFBBFBDDF9E349AE389C" box="[1119,1268,1653,1676]" italics="true" pageId="0" pageNumber="195">Laevimodulus</emphasis>
|
||
</taxonomicName>
|
||
share the same reinforced siphonal depression, which is well-delimited by a siphonal ridge (
|
||
<figureCitation id="13322A48FFCFFFBBFC52F90E495538BF" box="[976,1039,1688,1711]" captionStart="FIGURE 2" captionStartId="2.[151,244,1806,1828]" captionTargetBox="[161,1418,868,1770]" captionTargetId="figure@2.[151,1436,850,1785]" captionTargetPageId="2" captionText="FIGURE 2. Outlines of recent and fossil Modulidae. A. Indomodulus tectum, New Caledonia, Touho, Montrouzier expedition, sta. 1246, 20 ° 48, 8 ’ S, 165 ° 08.7 ' E [MNHN-IM- 2012 - 36610]. B. Modulus turbinoides (Locard, 1897) Cape Verde, Calypso 1959, sta. 65, 15 ° 37 ’ N, 23 ° 09 ’ W [MNHN-IM- 2012 - 36609]. C. Modulus modulus (Linnaeus, 1758), Sanibel I., Florida, coll. Staadt [MNHN-IM- 2012 - 36612]. D. Conomodulus neocaledonensis sp. nov., holotype, New Caledonia, Ouen I., Lagon campaign, Sta. 243, 22 ° 24 S, 167 ° 01 E [MNHN-IM- 2000 - 31117]. E. Conomodulus preangerensis (K. Martin, 1905), late Miocene, early Tortonian of Bontang, East Kalimantan, Indonesia (locality TF 505) [RGM. 794.179]. F. Incisilabium parisiensis (Deshayes, 1832), Middle Eocene, Bartonian, Paris Basin, St-Vaast-lès-Mello (Barisseuse), coll. Pacaud [MNHN. F. A 51573]. The arrows indicate the siphonal depression, reinforced in B and C." httpUri="https://zenodo.org/record/262500/files/figure.png" pageId="0" pageNumber="195">Fig. 2</figureCitation>
|
||
B, C). The monotypic Mio-Pliocene American genus
|
||
<taxonomicName id="4C094D4EFFCFFFBBFECEF92C4CBB38C1" ID-CoL="7PDNY" box="[332,481,1722,1745]" class="Gastropoda" family="Modulidae" genus="Psammodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEADFFFCFFFBBFECEF92C4CBB38C1" box="[332,481,1722,1745]" italics="true" pageId="0" pageNumber="195">Psammodulus</emphasis>
|
||
</taxonomicName>
|
||
has a shallow siphonal depression, but this genus is unusual amongst
|
||
<taxonomicName id="4C094D4EFFCFFFBBFB4FF92C481A38C1" ID-CoL="7NJTM" box="[1229,1344,1722,1745]" class="Gastropoda" family="Modulidae" kingdom="Animalia" pageId="0" pageNumber="195" phylum="Mollusca" rank="family">Modulidae</taxonomicName>
|
||
because of the coarse grains of sand adhering to the surface of the shell.
|
||
</paragraph>
|
||
<paragraph id="8BB636CDFFCFFFBBFF45F8964F6339D4" blockId="0.[151,1436,1168,2023]" pageId="0" pageNumber="195">
|
||
The European fossil genus
|
||
<taxonomicName id="4C094D4EFFCFFFBBFE7EF89649DF3907" authority="Cossmann, 1918 (Middle Eocene, Bartonian)" authorityName="Cossmann, 1918 (Middle Eocene, Bartonian" box="[508,1157,1792,1815]" class="Gastropoda" family="Modulidae" genus="Incisilabium" higherTaxonomySource="CoL" kingdom="Animalia" order="Archaeogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEADFFFCFFFBBFE7EF8964FDB3907" box="[508,641,1792,1815]" italics="true" pageId="0" pageNumber="195">Incisilabium</emphasis>
|
||
Cossmann, 1918 (Middle Eocene, Bartonian)
|
||
</taxonomicName>
|
||
, the oldest unambiguous member of
|
||
<taxonomicName id="4C094D4EFFCFFFBBFE99F8B44CD43929" ID-CoL="7NJTM" box="[283,398,1826,1849]" class="Gastropoda" family="Modulidae" kingdom="Animalia" pageId="0" pageNumber="195" phylum="Mollusca" rank="family">Modulidae</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF984B3CFFCFFFBBFE23F8B44ED33929" author="Cossmann" box="[417,905,1826,1849]" pageId="0" pageNumber="195" refString="Cossmann, M. & Pissarro, G. (1907) Iconographie complete des coquilles fossiles de l'Eocene des environs de Paris. Tome II, pars - In- 4 º, Paris (Hermann), pl. I - IX." type="journal volume" year="1907">Cossmann & Pissarro 1907, pl. 3 fig. 22-1</bibRefCitation>
|
||
), bears a broad siphonal depression that is not reinforced and not delimited by a groove or ridge (
|
||
<figureCitation id="13322A48FFCFFFBBFD35F8D34FA2394C" box="[695,760,1861,1884]" captionStart="FIGURE 2" captionStartId="2.[151,244,1806,1828]" captionTargetBox="[161,1418,868,1770]" captionTargetId="figure@2.[151,1436,850,1785]" captionTargetPageId="2" captionText="FIGURE 2. Outlines of recent and fossil Modulidae. A. Indomodulus tectum, New Caledonia, Touho, Montrouzier expedition, sta. 1246, 20 ° 48, 8 ’ S, 165 ° 08.7 ' E [MNHN-IM- 2012 - 36610]. B. Modulus turbinoides (Locard, 1897) Cape Verde, Calypso 1959, sta. 65, 15 ° 37 ’ N, 23 ° 09 ’ W [MNHN-IM- 2012 - 36609]. C. Modulus modulus (Linnaeus, 1758), Sanibel I., Florida, coll. Staadt [MNHN-IM- 2012 - 36612]. D. Conomodulus neocaledonensis sp. nov., holotype, New Caledonia, Ouen I., Lagon campaign, Sta. 243, 22 ° 24 S, 167 ° 01 E [MNHN-IM- 2000 - 31117]. E. Conomodulus preangerensis (K. Martin, 1905), late Miocene, early Tortonian of Bontang, East Kalimantan, Indonesia (locality TF 505) [RGM. 794.179]. F. Incisilabium parisiensis (Deshayes, 1832), Middle Eocene, Bartonian, Paris Basin, St-Vaast-lès-Mello (Barisseuse), coll. Pacaud [MNHN. F. A 51573]. The arrows indicate the siphonal depression, reinforced in B and C." httpUri="https://zenodo.org/record/262500/files/figure.png" pageId="0" pageNumber="195">Fig. 2</figureCitation>
|
||
F). The
|
||
<typeStatus id="54B2886FFFCFFFBBFCCEF8D34E20394C" box="[844,890,1861,1884]" pageId="0" pageNumber="195">type</typeStatus>
|
||
species is also characterized by its regular rows of nodes and the total lack of an umbilicus.
|
||
<taxonomicName id="4C094D4EFFCFFFBBFDCEF8FE4E8A396F" ID-CoL="3PND8" authority="Gmelin, 1791" authorityName="Gmelin" authorityYear="1791" box="[588,976,1896,1919]" class="Gastropoda" family="Modulidae" genus="Indomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="tectum">
|
||
<emphasis id="B97DEADFFFCFFFBBFDCEF8FE4E7F396F" box="[588,805,1896,1919]" italics="true" pageId="0" pageNumber="195">Indomodulus tectum</emphasis>
|
||
(Gmelin, 1791)
|
||
</taxonomicName>
|
||
(and its varieties) bears a shallow siphonal depression, which is poorly delimited (
|
||
<figureCitation id="13322A48FFCFFFBBFDBFF81C4F2539B1" box="[573,639,1930,1953]" captionStart="FIGURE 2" captionStartId="2.[151,244,1806,1828]" captionTargetBox="[161,1418,868,1770]" captionTargetId="figure@2.[151,1436,850,1785]" captionTargetPageId="2" captionText="FIGURE 2. Outlines of recent and fossil Modulidae. A. Indomodulus tectum, New Caledonia, Touho, Montrouzier expedition, sta. 1246, 20 ° 48, 8 ’ S, 165 ° 08.7 ' E [MNHN-IM- 2012 - 36610]. B. Modulus turbinoides (Locard, 1897) Cape Verde, Calypso 1959, sta. 65, 15 ° 37 ’ N, 23 ° 09 ’ W [MNHN-IM- 2012 - 36609]. C. Modulus modulus (Linnaeus, 1758), Sanibel I., Florida, coll. Staadt [MNHN-IM- 2012 - 36612]. D. Conomodulus neocaledonensis sp. nov., holotype, New Caledonia, Ouen I., Lagon campaign, Sta. 243, 22 ° 24 S, 167 ° 01 E [MNHN-IM- 2000 - 31117]. E. Conomodulus preangerensis (K. Martin, 1905), late Miocene, early Tortonian of Bontang, East Kalimantan, Indonesia (locality TF 505) [RGM. 794.179]. F. Incisilabium parisiensis (Deshayes, 1832), Middle Eocene, Bartonian, Paris Basin, St-Vaast-lès-Mello (Barisseuse), coll. Pacaud [MNHN. F. A 51573]. The arrows indicate the siphonal depression, reinforced in B and C." httpUri="https://zenodo.org/record/262500/files/figure.png" pageId="0" pageNumber="195">Fig. 2</figureCitation>
|
||
A), and has the highest whorl expansion rate resulting in the largest body whorl relative to the spire of the group.
|
||
</paragraph>
|
||
<paragraph id="8BB636CDFFCFFFBAFF45F8464F133F29" blockId="0.[151,1436,1168,2023]" lastBlockId="1.[151,1436,152,383]" lastPageId="1" lastPageNumber="196" pageId="0" pageNumber="195">
|
||
Species of
|
||
<taxonomicName id="4C094D4EFFCFFFBBFEC2F8464C8E39F7" ID-CoL="7P27V" box="[320,468,2000,2023]" class="Gastropoda" family="Modulidae" genus="Conomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEADFFFCFFFBBFEC2F8464C8E39F7" box="[320,468,2000,2023]" italics="true" pageId="0" pageNumber="195">Conomodulus</emphasis>
|
||
</taxonomicName>
|
||
have a high spire and differ from other modulid genera by a wider and deeper siphonal groove (
|
||
<bibRefCitation id="EF984B3CFFCEFFBAFF6CFF0E4CE63EBF" author="Landau" box="[238,444,152,175]" pageId="1" pageNumber="196" refString="Landau, B. Vermeij, G. J. & Reich, S. (2014) Classification of the Modulidae (Caenogastropoda, Cerithioidea), with new genera and new fossil species from the Neogene of tropical America and Indonesia. Basteria, 78 (1 - 3), 1 - 29." type="journal article" year="2014">
|
||
Landau
|
||
<emphasis id="B97DEADFFFCEFFBAFEC4FF0E4C2E3EBF" box="[326,372,152,175]" italics="true" pageId="1" pageNumber="196">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
). We have examined the specimens of
|
||
<taxonomicName id="4C094D4EFFCEFFBAFCD7FF0E48723EBF" ID-CoL="XRCH" authority="Martin, 1905" authorityName="Martin" authorityYear="1905" box="[853,1320,152,175]" class="Gastropoda" family="Modulidae" genus="Conomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="1" pageNumber="196" phylum="Mollusca" rank="species" species="preangerensis">
|
||
<emphasis id="B97DEADFFFCEFFBAFCD7FF0E49DF3EBF" box="[853,1157,152,175]" italics="true" pageId="1" pageNumber="196">Conomodulus preangerensis</emphasis>
|
||
(Martin, 1905)
|
||
</taxonomicName>
|
||
figured by
|
||
<bibRefCitation id="EF984B3CFFCEFFBAFF15FF2C4C2A3EC1" box="[151,368,186,209]" pageId="1" pageNumber="196" refString="Landau, B. Vermeij, G. J. & Reich, S. (2014) Classification of the Modulidae (Caenogastropoda, Cerithioidea), with new genera and new fossil species from the Neogene of tropical America and Indonesia. Basteria, 78 (1 - 3), 1 - 29." type="journal article">
|
||
Landau
|
||
<emphasis id="B97DEADFFFCEFFBAFF6FFF2C4C793EC1" box="[237,291,186,209]" italics="true" pageId="1" pageNumber="196">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
[RGM.794.179, late Miocene, early Tortonian of Bontang, East Kalimantan,
|
||
<collectingCountry id="F31E765DFFCEFFBAFB2CFF2C484C3EC1" box="[1198,1302,186,209]" name="Indonesia" pageId="1" pageNumber="196">Indonesia</collectingCountry>
|
||
(locality TF 505), Height 12.0 mm; RGM.312.287, middle Miocene, lower Menkrawit beds, East Kalimantan,
|
||
<collectingCountry id="F31E765DFFCEFFBAFB53FF4B48633EE4" box="[1233,1337,221,244]" name="Indonesia" pageId="1" pageNumber="196">Indonesia</collectingCountry>
|
||
(locality 391), Height 5.0 mm (incomplete)] and have observed that these specimens have weak lirae on the inside of the outer lip and a shallow siphonal groove (
|
||
<figureCitation id="13322A48FFCEFFBAFE6BFEB44F733F29" box="[489,553,290,313]" captionStart="FIGURE 2" captionStartId="2.[151,244,1806,1828]" captionTargetBox="[161,1418,868,1770]" captionTargetId="figure@2.[151,1436,850,1785]" captionTargetPageId="2" captionText="FIGURE 2. Outlines of recent and fossil Modulidae. A. Indomodulus tectum, New Caledonia, Touho, Montrouzier expedition, sta. 1246, 20 ° 48, 8 ’ S, 165 ° 08.7 ' E [MNHN-IM- 2012 - 36610]. B. Modulus turbinoides (Locard, 1897) Cape Verde, Calypso 1959, sta. 65, 15 ° 37 ’ N, 23 ° 09 ’ W [MNHN-IM- 2012 - 36609]. C. Modulus modulus (Linnaeus, 1758), Sanibel I., Florida, coll. Staadt [MNHN-IM- 2012 - 36612]. D. Conomodulus neocaledonensis sp. nov., holotype, New Caledonia, Ouen I., Lagon campaign, Sta. 243, 22 ° 24 S, 167 ° 01 E [MNHN-IM- 2000 - 31117]. E. Conomodulus preangerensis (K. Martin, 1905), late Miocene, early Tortonian of Bontang, East Kalimantan, Indonesia (locality TF 505) [RGM. 794.179]. F. Incisilabium parisiensis (Deshayes, 1832), Middle Eocene, Bartonian, Paris Basin, St-Vaast-lès-Mello (Barisseuse), coll. Pacaud [MNHN. F. A 51573]. The arrows indicate the siphonal depression, reinforced in B and C." httpUri="https://zenodo.org/record/262500/files/figure.png" pageId="1" pageNumber="196">Fig. 2</figureCitation>
|
||
E).
|
||
</paragraph>
|
||
<paragraph id="8BB636CDFFCEFFBAFF45FED348DE3F6F" blockId="1.[151,1436,152,383]" pageId="1" pageNumber="196">
|
||
Judging from the general shape, particularly by the high spire, the new species from
|
||
<collectingCountry id="F31E765DFFCEFFBAFBFCFED3487D3F4C" box="[1150,1319,325,348]" name="New Caledonia" pageId="1" pageNumber="196">New Caledonia</collectingCountry>
|
||
cannot be included in the Indo-Pacific genus
|
||
<taxonomicName id="4C094D4EFFCEFFBAFD89FEFE4FCC3F6F" ID-CoL="7P5RB" box="[523,662,360,383]" class="Gastropoda" family="Modulidae" genus="Indomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="1" pageNumber="196" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEADFFFCEFFBAFD89FEFE4FCC3F6F" box="[523,662,360,383]" italics="true" pageId="1" pageNumber="196">Indomodulus</emphasis>
|
||
</taxonomicName>
|
||
and appears related to the Indo-Pacific Neogene genus
|
||
<taxonomicName id="4C094D4EFFCEFFBAFB6BFEFE48273F6F" ID-CoL="7P27V" box="[1257,1405,360,383]" class="Gastropoda" family="Modulidae" genus="Conomodulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caenogastropoda" pageId="1" pageNumber="196" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B97DEADFFFCEFFBAFB6BFEFE48273F6F" box="[1257,1405,360,383]" italics="true" pageId="1" pageNumber="196">Conomodulus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |