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<document id="C7E99FD2CED5648FE2764BF23A4C6569" ID-DOI="10.1649/072.068.0301" ID-ISSN="1938-4394" ID-Zenodo-Dep="10101811" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630152396499" checkinUser="valdenar" docAuthor="Bartolozzi, Luca, Keyzer, Roger De &amp; Zilioli, Michele" docDate="2014" docId="F11E44693902FF97FF2793657357FCF2" docLanguage="en" docName="072.068.0301.pdf.imf" docOrigin="The Coleopterists Bulletin 68 (3)" docSource="http://dx.doi.org/10.1649/072.068.0301" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Lissotes krakei Bartolozzi, de Keyzer, and Zilioli, new species" docType="treatment" docUuid="C5131217-9732-4B0B-8362-BFE766A345A5" docUuidSource="ZooBank" docVersion="4" lastPageNumber="348" masterDocId="0D273C113903FF94FF9491207034FFC8" masterDocTitle="Contribution to the Knowledge of the Australian Stag Beetles, with Description of a New Species ofLissotesWestwood, 1855 (Coleoptera: Lucanidae)" masterLastPageNumber="350" masterPageNumber="345" pageNumber="346" updateTime="1699618221065" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B5C72A195CC08A7043E5EA421CC53505">Contribution to the Knowledge of the Australian Stag Beetles, with Description of a New Species ofLissotesWestwood, 1855 (Coleoptera: Lucanidae)</mods:title>
</mods:titleInfo>
<mods:name id="BA24B66AC2F09E97D90D74018180B47F" type="personal">
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<mods:namePart id="EA2291AF4C3053B452F0D358E847E9B9">Bartolozzi, Luca</mods:namePart>
</mods:name>
<mods:name id="8D21BF9BDD920187A2BA4F904C53D764" type="personal">
<mods:role id="7D173AD937008D0D3F1788D888C1BF58">
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<mods:namePart id="FB360BBB17A44F702887F1F05A2F5672">Keyzer, Roger De</mods:namePart>
</mods:name>
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<mods:namePart id="0D3F069DFE34A46A5A3E6D51C830EFA1">Zilioli, Michele</mods:namePart>
</mods:name>
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<mods:title id="659AAEFBA8C5C0F2748380D7E1B0A6EC">The Coleopterists Bulletin</mods:title>
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<mods:part id="F4A9BC60E8E5A130F030B0265A8DE8E3">
<mods:date id="0D09FDCD0FFAF1C1658E8B5EF1B98168">2014</mods:date>
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<mods:number id="B588105F679EBE05EF83CC2BF47A16FB">2014-09-30</mods:number>
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<mods:number id="7679CF60F4A45C31D625C83A30FACF12">68</mods:number>
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<mods:url id="C16878302711AFC676E3771053B7EDBC">http://dx.doi.org/10.1649/072.068.0301</mods:url>
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<mods:classification id="A13D75453E83B107A26BB8654173AD59">journal article</mods:classification>
<mods:identifier id="5E2724B6D60895E974E194BB3AC99649" type="DOI">10.1649/072.068.0301</mods:identifier>
<mods:identifier id="CF9305AC783A4139F953BAD814FA30BF" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="729D3772CAA70C10051A9868DD4B8C45" type="Zenodo-Dep">10101811</mods:identifier>
<mods:identifier id="BB34E84475C4A34F37BB4A9B7120C8A8" type="ZooBank">700DD1A0-5B96-4FCE-806E-C6A3A717B7AD</mods:identifier>
</mods:mods>
<treatment id="F11E44693902FF97FF2793657357FCF2" LSID="urn:lsid:zoobank.org:act:C5131217-9732-4B0B-8362-BFE766A345A5" httpUri="http://treatment.plazi.org/id/F11E44693902FF97FF2793657357FCF2" lastPageId="3" lastPageNumber="348" pageId="1" pageNumber="346">
<subSubSection id="31ADA6F43902FF95FF27936571DFFDBD" pageId="1" pageNumber="346" type="nomenclature">
<paragraph id="7908F57F3902FF95FF27936571DFFDBD" blockId="1.[179,610,581,656]" pageId="1" pageNumber="346">
<heading id="224042133902FF95FF27936571DFFDBD" centered="true" fontSize="10" level="2" pageId="1" pageNumber="346" reason="2">
<emphasis id="4BC3296D3902FF95FF27936571DFFDBD" bold="true" pageId="1" pageNumber="346">
<taxonomicName id="BEB78EFC3902FF95FF279365715CFDBD" ID-CoL="3VGD3" authority="Bartolozzi, de Keyzer, and Zilioli" authorityName="Bartolozzi, de Keyzer, and Zilioli" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="346" phylum="Arthropoda" rank="species" species="krakei" status="sp. nov.">
<emphasis id="4BC3296D3902FF95FF2793657173FD92" bold="true" box="[179,327,581,602]" italics="true" pageId="1" pageNumber="346">Lissotes krakei</emphasis>
Bartolozzi, de Keyzer, and Zilioli
</taxonomicName>
,
<taxonomicNameLabel id="50F094163902FF95FEE1934071DFFDBD" box="[373,491,608,629]" pageId="1" pageNumber="346" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="31ADA6F43902FF95FEC1935B71F4FD58" box="[341,448,634,656]" pageId="1" pageNumber="346" type="description">
<paragraph id="7908F57F3902FF95FEC1935B71F4FD58" blockId="1.[179,610,581,656]" box="[341,448,634,656]" pageId="1" pageNumber="346">
(
<figureCitation id="E18CE9FA3902FF95FEC8935B718DFD58" box="[348,441,634,656]" captionStart-0="Figs" captionStart-1="Fig" captionStart-2="Figs" captionStart-3="Figs" captionStartId-0="1.[175,215,1710,1729]" captionStartId-1="2.[197,229,575,594]" captionStartId-2="2.[197,237,1710,1729]" captionStartId-3="4.[197,237,1066,1085]" captionTargetBox-0="[171,1153,794,1688]" captionTargetBox-1="[180,649,197,552]" captionTargetBox-2="[173,1193,992,1686]" captionTargetBox-3="[216,1193,196,1043]" captionTargetId-0="figure-253@1.[151,1172,772,1705]" captionTargetId-1="figure-287@2.[173,659,183,552]" captionTargetId-2="figure-298@2.[172,1193,991,1686]" captionTargetId-3="figure-246@4.[172,1193,183,1043]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="2" captionTargetPageId-3="4" captionText-0="Figs. 12. Lissotes krakei, habitus in dorsal view. 1) Male (total length 20.2 mm); 2) Female (total length 18.4 mm)." captionText-1="Fig. 3. Lissotes krakei, female, head and prothorax." captionText-2="Figs. 46. Lissotes krakei, aedeagus. 4) Dorsal view; 5) Ventral view; 6) Lateral view." captionText-3="Figs. 710. Lissotes species, male heads and mandibles. 7) L. krakei; 8) L. darlingtoni; 9) L. furcicornis;" figureDoi-0="http://doi.org/10.5281/zenodo.10101813" figureDoi-1="http://doi.org/10.5281/zenodo.10101815" figureDoi-2="http://doi.org/10.5281/zenodo.10101817" figureDoi-3="http://doi.org/10.5281/zenodo.10101819" httpUri-0="https://zenodo.org/record/10101813/files/figure.png" httpUri-1="https://zenodo.org/record/10101815/files/figure.png" httpUri-2="https://zenodo.org/record/10101817/files/figure.png" httpUri-3="https://zenodo.org/record/10101819/files/figure.png" pageId="1" pageNumber="346">Figs. 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="31ADA6F43902FF96FF3B93857172FC9D" lastPageId="2" lastPageNumber="347" pageId="1" pageNumber="346" type="materials_examined">
<paragraph id="7908F57F3902FF95FF3B9385724AFD72" blockId="1.[151,638,677,725]" box="[175,638,677,699]" pageId="1" pageNumber="346">
<emphasis id="4BC3296D3902FF95FF3B93857237FD72" bold="true" box="[175,515,677,698]" pageId="1" pageNumber="346">
Type Material.
<typeStatus id="A60C4BDD3902FF95FEC09385718EFD72" box="[340,442,677,698]" pageId="1" pageNumber="346" type="holotype">Holotype</typeStatus>
male:
</emphasis>
“Victorian
</paragraph>
<paragraph id="7908F57F3902FF95FF0393E07249FD1D" blockId="1.[151,638,677,725]" box="[151,637,704,725]" pageId="1" pageNumber="346">
Alps, Mt. Wills,
<date id="0D09D3BF3902FF95FEA993E07199FD1D" box="[317,429,704,725]" pageId="1" pageNumber="346" value="1999-12-07">7.Dec.1999</date>
, leg. S. Smith/ COL-
</paragraph>
<paragraph id="7908F57F3902FF96FD3991E27172FC9D" blockId="1.[685,1172,194,725]" lastBlockId="2.[173,660,645,932]" lastPageId="2" lastPageNumber="347" pageId="1" pageNumber="346">
115022/ Mus. Vic. Ento 2012-15L/
<taxonomicName id="BEB78EFC3902FF95FBBE91E274A0FF1F" box="[1066,1172,194,215]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="346" phylum="Arthropoda" rank="species" species="alpinus">
<emphasis id="4BC3296D3902FF95FBBE91E274A0FF1F" box="[1066,1172,194,215]" italics="true" pageId="1" pageNumber="346">L. alpinus</emphasis>
</taxonomicName>
G. K.”.
<typeStatus id="A60C4BDD3902FF95FC9691FD736AFF3A" box="[770,862,221,242]" pageId="1" pageNumber="346" type="holotype">Holotype</typeStatus>
deposited in MV.
<emphasis id="4BC3296D3902FF95FBB791FD74B9FF3A" bold="true" box="[1059,1165,221,242]" pageId="1" pageNumber="346">
<typeStatus id="A60C4BDD3902FF95FBB791FD74B9FF3A" box="[1059,1165,221,242]" pageId="1" pageNumber="346" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<materialsCitation id="C9DFFF223902FF95FD3991D8731FFEE0" collectionCode="LB, RDK" pageId="1" pageNumber="346" specimenCount="3" specimenCount-male="3">
<specimenCount id="6FB13EF63902FF95FD3991D872EAFEC5" box="[685,734,247,269]" pageId="1" pageNumber="346" type="male">3♂♂</specimenCount>
with same collecting data as holotype (
<collectionCode id="1FA66DBA3902FF95FBE091D874A0FEC5" box="[1140,1172,248,269]" pageId="1" pageNumber="346">LB</collectionCode>
and 2
<collectionCode id="1FA66DBA3902FF95FD7A90337310FEE0" box="[750,804,275,296]" pageId="1" pageNumber="346">RDK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FCA3903373F4FE8B" collectingDate="1940-01" collectionCode="MV" collectorName="C. Oke" location="Australian Alps" municipality="Vic" pageId="1" pageNumber="346" specimenCount="1" specimenCount-male="1">
<specimenCount id="6FB13EF63902FF95FCA390337361FEE0" box="[823,853,274,296]" pageId="1" pageNumber="346" type="male">1♂</specimenCount>
<location id="7C68A3A43902FF95FCCA903373CCFEE0" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43902FF95FCCA903373CCFEE0" box="[862,1016,275,296]" municipality="Vic" name="Australian Alps" pageId="1" pageNumber="346">Australian Alps</location>
,
<collectingMunicipality id="996C6F053902FF95FB919033741CFEE0" box="[1029,1064,275,296]" pageId="1" pageNumber="346">Vic</collectingMunicipality>
,
<date id="0D09D3BF3902FF95FBA1903374A4FEE0" box="[1077,1168,275,296]" pageId="1" pageNumber="346" value="1940-01">
<collectingDate id="1D4D2A573902FF95FBA1903374A4FEE0" box="[1077,1168,275,296]" pageId="1" pageNumber="346" value="1940-01">Jan 1940</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FD39900E72C7FE8B" box="[685,755,302,323]" pageId="1" pageNumber="346">C. Oke</collectorName>
/ COL-86147/ (
<collectionCode id="1FA66DBA3902FF95FC1A900E738DFE8B" box="[910,953,302,323]" country="0" name="University of Montana Museum" pageId="1" pageNumber="346">MV</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FC59900E745EFE95" collectingDate="1952-02-22" collectionCode="MV" collectorName="F. E. Wilson" location="Glen Wills" municipality="Vic" pageId="1" pageNumber="346" specimenCount="1" specimenCount-male="1">
<specimenCount id="6FB13EF63902FF95FC59900E73DFFE8B" box="[973,1003,301,323]" pageId="1" pageNumber="346" type="male">1♂</specimenCount>
<location id="7C68A3A43902FF95FC60900E7454FE8B" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43902FF95FC60900E7454FE8B" box="[1012,1120,302,323]" municipality="Vic" name="Glen Wills" pageId="1" pageNumber="346">Glen Wills</location>
,
<collectingMunicipality id="996C6F053902FF95FBF8900E74BBFE8B" box="[1132,1167,302,323]" pageId="1" pageNumber="346">Vic</collectingMunicipality>
,
<date id="0D09D3BF3902FF95FD399068731FFE95" box="[685,811,328,349]" pageId="1" pageNumber="346" value="1952-02-22">
<collectingDate id="1D4D2A573902FF95FD399068731FFE95" box="[685,811,328,349]" pageId="1" pageNumber="346" value="1952-02-22">22 Feb 1952</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FCA39068739DFE95" box="[823,937,328,349]" pageId="1" pageNumber="346">F.E. Wilson</collectorName>
/COL-86143/ (
<collectionCode id="1FA66DBA3902FF95FBAC90687456FE95" box="[1080,1122,328,349]" country="0" name="University of Montana Museum" pageId="1" pageNumber="346">MV</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FBE290687422FE5B" collectingDate="2002-03-08" collectionCode="AP" collectorName="J. Balderson" county="Vic" location="Strawberry Saddle" municipality="Bogong High Plains" pageId="1" pageNumber="346" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="6FB13EF63902FF95FBE2906874A0FE96" box="[1142,1172,328,350]" pageId="1" pageNumber="346" type="female">1♀</specimenCount>
<specimenCount id="6FB13EF63902FF95FD39904372E9FEB1" box="[685,733,355,377]" pageId="1" pageNumber="346" type="male">2♂♂</specimenCount>
<location id="7C68A3A43902FF95FD77904373A2FEB0" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43902FF95FD77904373A2FEB0" box="[739,918,355,376]" county="Vic" municipality="Bogong High Plains" name="Strawberry Saddle" pageId="1" pageNumber="346">Strawberry Saddle</location>
,
<collectingMunicipality id="996C6F053902FF95FC0B90437457FEB0" box="[927,1123,355,376]" pageId="1" pageNumber="346">Bogong High Plains</collectingMunicipality>
,
<collectingCounty id="90698DF33902FF95FBF9904374A4FEB0" box="[1133,1168,355,376]" pageId="1" pageNumber="346">Vic</collectingCounty>
,
<date id="0D09D3BF3902FF95FD39905E7372FE5B" box="[685,838,382,403]" pageId="1" pageNumber="346" value="2002-03-08">
<collectingDate id="1D4D2A573902FF95FD39905E7372FE5B" box="[685,838,382,403]" pageId="1" pageNumber="346" value="2002-03-08">8 March 2002</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FCC0905E73E3FE5B" box="[852,983,382,403]" pageId="1" pageNumber="346">J. Balderson</collectorName>
(
<collectionCode id="1FA66DBA3902FF95FC7D905E743AFE5B" box="[1001,1038,382,403]" pageId="1" pageNumber="346">AP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FBB0905E7369FE01" collectingDate="26 Dec 1992 and 30 Jan 1996" collectionCode="AP" collectorName="J Balderson" county="Vic" location="Strawberry Saddle" municipality="Bogong High Plains" pageId="1" pageNumber="346" specimenCount="2" specimenCount-male="2">
<specimenCount id="6FB13EF63902FF95FBB0905E7462FE5B" box="[1060,1110,381,403]" pageId="1" pageNumber="346" type="male">2♂♂</specimenCount>
same location data,
<collectingDate id="1D4D2A573902FF95FCD390B974A4FE66" box="[839,1168,409,430]" isEnumeration="true" pageId="1" pageNumber="346">
<date id="0D09D3BF3902FF95FCD390B973E4FE66" box="[839,976,409,430]" pageId="1" pageNumber="346" value="1992-12-26">26 Dec 1992</date>
and
<date id="0D09D3BF3902FF95FB9E90B974A4FE66" box="[1034,1168,409,430]" pageId="1" pageNumber="346" value="1996-01-30">30 Jan 1996</date>
</collectingDate>
,
<collectorName id="D44290A93902FF95FD3990947317FE01" box="[685,803,436,457]" pageId="1" pageNumber="346">J Balderson</collectorName>
(
<collectionCode id="1FA66DBA3902FF95FCA790947362FE01" box="[819,854,436,457]" pageId="1" pageNumber="346">AP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FCFF909472EBFE36" collectingDate="1992-12-25" collectionCode="AP" collectorName="J. Balderson" county="Vic" location="West Viaduct" municipality="Bogong High Plains" pageId="1" pageNumber="346" specimenCount="2" specimenCount-male="2">
<specimenCount id="6FB13EF63902FF95FCFF909473AFFE01" box="[875,923,435,457]" pageId="1" pageNumber="346" type="male">2♂♂</specimenCount>
Langford
<location id="7C68A3A43902FF95FB9D909474A4FE01" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43902FF95FB9D909474A4FE01" box="[1033,1168,436,457]" county="Vic" municipality="Bogong High Plains" name="West Viaduct" pageId="1" pageNumber="346">West Viaduct</location>
,
<collectingMunicipality id="996C6F053902FF95FD3990EE735FFE2B" box="[685,875,462,483]" pageId="1" pageNumber="346">Bogong High Plains</collectingMunicipality>
,
<collectingCounty id="90698DF33902FF95FCE090EE73A2FE2B" box="[884,918,462,483]" pageId="1" pageNumber="346">Vic</collectingCounty>
,
<date id="0D09D3BF3902FF95FC3490EE742CFE2B" box="[928,1048,462,483]" pageId="1" pageNumber="346" value="1992-12-25">
<collectingDate id="1D4D2A573902FF95FC3490EE742CFE2B" box="[928,1048,462,483]" pageId="1" pageNumber="346" value="1992-12-25">25 Dec 1992</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FBB690EE74A0FE2B" box="[1058,1172,462,483]" pageId="1" pageNumber="346">J. Balderson</collectorName>
(
<collectionCode id="1FA66DBA3902FF95FD2190C972ECFE36" box="[693,728,489,510]" pageId="1" pageNumber="346">AP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FD6490C9734FFDD1" collectingDate="1994-02-05" collectingDateMax="1994-02-13" collectingDateMin="1994-02-05" collectionCode="AP" collectorName="J. Balderson" county="Vic" location="West Viaduct" municipality="Bogong High Plains" pageId="1" pageNumber="346" specimenCount="1" specimenCount-male="1">
<specimenCount id="6FB13EF63902FF95FD6490C9733AFE37" box="[752,782,489,511]" pageId="1" pageNumber="346" type="male">1♂</specimenCount>
same location data,
<date id="0D09D3BF3902FF95FC7A90C974A4FE36" box="[1006,1168,489,511]" pageId="1" pageNumber="346" value="1994-02-05" valueMax="1994-02-13" valueMin="1994-02-05">
<collectingDate id="1D4D2A573902FF95FC7A90C974A4FE36" box="[1006,1168,489,511]" pageId="1" pageNumber="346" value="1994-02-05" valueMax="1994-02-13" valueMin="1994-02-05">513 Feb 1994</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FD3993247303FDD1" box="[685,823,516,537]" pageId="1" pageNumber="346">J. Balderson</collectorName>
(
<collectionCode id="1FA66DBA3902FF95FCD993247347FDD1" box="[845,883,516,537]" pageId="1" pageNumber="346">AP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FC19932473CCFDFC" collectingDate="1996-01-28" collectingDateMax="1994-02-13" collectingDateMin="1994-02-05" collectionCode="AP" collectorName="J. Balderson" county="Vic" location="West Viaduct" municipality="Bogong High Plains" pageId="1" pageNumber="346" specimenCount="1" specimenCount-female="1">
<specimenCount id="6FB13EF63902FF95FC1993247398FDD2" box="[909,940,516,538]" pageId="1" pageNumber="346" type="female">1♀</specimenCount>
same location data,
<date id="0D09D3BF3902FF95FD39933F731BFDFC" box="[685,815,543,564]" pageId="1" pageNumber="346" value="1996-01-28">
<collectingDate id="1D4D2A573902FF95FD39933F731BFDFC" box="[685,815,543,564]" pageId="1" pageNumber="346" value="1996-01-28">28 Jan 1996</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FCAA933F738FFDFC" box="[830,955,543,564]" pageId="1" pageNumber="346">J. Balderson</collectorName>
(
<collectionCode id="1FA66DBA3902FF95FC5A933F73C5FDFC" box="[974,1009,543,564]" pageId="1" pageNumber="346">AP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FB93933F7465FDA1" collectingDate="1994-02-10" collectionCode="AP" collectorName="J. Balderson &amp; P. K. Christensen" county="Vic" location="Buckety Plain" municipality="Bogong High Plains" pageId="1" pageNumber="346" specimenCount="3" specimenCount-male="3">
<specimenCount id="6FB13EF63902FF95FB93933F7403FDFC" box="[1031,1079,542,564]" pageId="1" pageNumber="346" type="male">3♂♂</specimenCount>
<location id="7C68A3A43902FF95FBD7933F72D5FD87" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43902FF95FBD7933F72D5FD87" county="Vic" municipality="Bogong High Plains" name="Buckety Plain" pageId="1" pageNumber="346">Buckety Plain</location>
,
<collectingMunicipality id="996C6F053902FF95FD65931A73F1FD87" box="[753,965,570,591]" pageId="1" pageNumber="346">Bogong High Plains</collectingMunicipality>
,
<collectingCounty id="90698DF33902FF95FC41931A73CDFD87" box="[981,1017,570,591]" pageId="1" pageNumber="346">Vic</collectingCounty>
,
<date id="0D09D3BF3902FF95FB9C931A74A4FD87" box="[1032,1168,570,591]" pageId="1" pageNumber="346" value="1994-02-10">
<collectingDate id="1D4D2A573902FF95FB9C931A74A4FD87" box="[1032,1168,570,591]" pageId="1" pageNumber="346" value="1994-02-10">10 Feb 1994</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FD3993747307FDA1" box="[685,819,596,617]" pageId="1" pageNumber="346">J. Balderson</collectorName>
&amp;
<collectorName id="D44290A93902FF95FCCB93747424FDA1" box="[863,1040,596,617]" pageId="1" pageNumber="346">P.K. Christensen</collectorName>
(
<collectionCode id="1FA66DBA3902FF95FBB09374747DFDA1" box="[1060,1097,596,617]" pageId="1" pageNumber="346">AP</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF95FBF6937472EDFD72" collectingDate="1997-01-09" collectionCode="RDK" collectorName="R. de Keyzer &amp; A. Sundholm" country="American Samoa" county="Vic" location="Mt Wills" municipality="Sunnyside" pageId="1" pageNumber="346" specimenCount="5" specimenCount-male="5">
<specimenCount id="6FB13EF63902FF95FBF6937474A0FDA2" box="[1122,1172,596,618]" pageId="1" pageNumber="346" type="male">5♂♂</specimenCount>
<location id="7C68A3A43902FF95FD39934F733AFD4C" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43902FF95FD39934F733AFD4C" box="[685,782,623,644]" country="American Samoa" county="Vic" municipality="Sunnyside" name="Mt Wills" pageId="1" pageNumber="346">Mt Wills</location>
,
<quantity id="BE4F589A3902FF95FC8B934F7362FD4C" box="[799,854,623,644]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="346" unit="km" value="2.0">2 km</quantity>
N. of
<collectingMunicipality id="996C6F053902FF95FC3D934F7421FD4C" box="[937,1045,623,644]" pageId="1" pageNumber="346">Sunnyside</collectingMunicipality>
,
<collectingCounty id="90698DF33902FF95FBB1934F747EFD4C" box="[1061,1098,623,644]" pageId="1" pageNumber="346">Vic</collectingCounty>
,
<date id="0D09D3BF3902FF95FBCE934F72D5FD57" pageId="1" pageNumber="346" value="1997-01-09">
<collectingDate id="1D4D2A573902FF95FBCE934F72D5FD57" pageId="1" pageNumber="346" value="1997-01-09">9 Jan 1997</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FD7B93AA7342FD57" box="[751,886,650,671]" pageId="1" pageNumber="346">R. de Keyzer</collectorName>
&amp;
<collectorName id="D44290A93902FF95FC0F93AA7415FD57" box="[923,1057,650,671]" pageId="1" pageNumber="346">A. Sundholm</collectorName>
(
<collectionCode id="1FA66DBA3902FF95FBA693AA7453FD57" box="[1074,1127,650,671]" pageId="1" pageNumber="346">RDK</collectionCode>
and
<collectingCountry id="01A0B5EF3902FF95FD39938572E5FD72" box="[685,721,677,698]" name="American Samoa" pageId="1" pageNumber="346">AS</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223902FF96FD7D938570C6FD53" collectingDate="1997-01-09" collectionCode="RDK" collectorName="R. de Keyzer &amp; A. Sundholm" country="American Samoa" county="Vic" lastPageId="2" lastPageNumber="347" location="Mt Wills" municipality="Sunnyside" pageId="1" pageNumber="346" specimenCount="1" specimenCount-male="1">
<specimenCount id="6FB13EF63902FF95FD7D9385733CFD72" box="[745,776,676,698]" pageId="1" pageNumber="346" type="male">1♂</specimenCount>
<location id="7C68A3A43902FF95FC8093857342FD72" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43902FF95FC8093857342FD72" box="[788,886,677,698]" country="American Samoa" county="Vic" municipality="Sunnyside" name="Mt Wills" pageId="1" pageNumber="346">Mt Wills</location>
,
<quantity id="BE4F589A3902FF95FC11938573E5FD72" box="[901,977,677,698]" metricMagnitude="3" metricUnit="m" metricValue="8.8" pageId="1" pageNumber="346" unit="km" value="8.8">8.8 km</quantity>
N. of
<collectingMunicipality id="996C6F053902FF95FBB5938574A4FD72" box="[1057,1168,677,698]" pageId="1" pageNumber="346">Sunnyside</collectingMunicipality>
,
<collectingCounty id="90698DF33902FF95FD3993E072E5FD1D" box="[685,721,704,725]" pageId="1" pageNumber="346">Vic</collectingCounty>
,
<date id="0D09D3BF3902FF95FD4B93E07367FD1D" box="[735,851,704,725]" pageId="1" pageNumber="346" value="1997-01-09">
<collectingDate id="1D4D2A573902FF95FD4B93E07367FD1D" box="[735,851,704,725]" pageId="1" pageNumber="346" value="1997-01-09">9 Jan 1997</collectingDate>
</date>
,
<collectorName id="D44290A93902FF95FCF593E073DCFD1D" box="[865,1000,704,725]" pageId="1" pageNumber="346">R. de Keyzer</collectorName>
&amp;
<collectorName id="D44290A93902FF95FB9A93E074A0FD1D" box="[1038,1172,704,725]" pageId="1" pageNumber="346">A. Sundholm</collectorName>
(
<collectionCode id="1FA66DBA3901FF96FF2093A670D8FD53" box="[180,236,646,667]" pageId="2" pageNumber="347">RDK</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223901FF96FF6B93A6717EFD18" collectingDate="1997-01-09" collectionCode="RDK" collectorName="R. de Keyzer &amp; A. Sundholm" country="American Samoa" county="Vic" location="Mt Wills" municipality="Sunnyside" pageId="2" pageNumber="347" specimenCount="4" specimenCount-male="4">
<specimenCount id="6FB13EF63901FF96FF6B93A6711BFD53" box="[255,303,645,667]" pageId="2" pageNumber="347" type="male">4♂♂</specimenCount>
<location id="7C68A3A43901FF96FEA393A671A6FD53" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43901FF96FEA393A671A6FD53" box="[311,402,646,667]" country="American Samoa" county="Vic" municipality="Sunnyside" name="Mt Wills" pageId="2" pageNumber="347">Mt Wills</location>
,
<quantity id="BE4F589A3901FF96FE0A93A671D7FD53" box="[414,483,646,667]" metricMagnitude="3" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="347" unit="km" value="6.2">6.2 km</quantity>
N. of
<collectingMunicipality id="996C6F053901FF96FDBC93A672A4FD53" box="[552,656,646,667]" pageId="2" pageNumber="347">Sunnyside</collectingMunicipality>
,
<collectingCounty id="90698DF33901FF96FF39938070E5FD7D" box="[173,209,672,693]" pageId="2" pageNumber="347">Vic</collectingCounty>
,
<date id="0D09D3BF3901FF96FF4B93807166FD7D" box="[223,338,672,693]" pageId="2" pageNumber="347" value="1997-01-09">
<collectingDate id="1D4D2A573901FF96FF4B93807166FD7D" box="[223,338,672,693]" pageId="2" pageNumber="347" value="1997-01-09">9 Jan 1997</collectingDate>
</date>
,
<collectorName id="D44290A93901FF96FEF4938071D3FD7D" box="[352,487,672,693]" pageId="2" pageNumber="347">R. de Keyzer</collectorName>
&amp;
<collectorName id="D44290A93901FF96FD99938072A7FD7D" box="[525,659,672,693]" pageId="2" pageNumber="347">A. Sundholm</collectorName>
(
<collectionCode id="1FA66DBA3901FF96FF27939B70DCFD18" box="[179,232,699,720]" pageId="2" pageNumber="347">RDK</collectionCode>
and
<collectingCountry id="01A0B5EF3901FF96FEB4939B7176FD18" box="[288,322,699,720]" name="American Samoa" pageId="2" pageNumber="347">AS</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223901FF96FECD939B71D0FCCD" collectingDate="1997-10-25" collectionCode="RDK" collectorName="R. de Keyzer &amp; A. Sundholm" country="American Samoa" county="Vic" location="Mt Wills" municipality="Sunnyside" pageId="2" pageNumber="347" specimenCount="6" specimenCount-female="1" specimenCount-male="5">
<specimenCount id="6FB13EF63901FF96FECD939B71BDFD18" box="[345,393,698,720]" pageId="2" pageNumber="347" type="male">5♂♂</specimenCount>
<specimenCount id="6FB13EF63901FF96FE00939B7187FD18" box="[404,435,698,720]" pageId="2" pageNumber="347" type="female">1♀</specimenCount>
<location id="7C68A3A43901FF96FE29939B722EFD18" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43901FF96FE29939B722EFD18" box="[445,538,699,720]" country="American Samoa" county="Vic" municipality="Sunnyside" name="Mt Wills" pageId="2" pageNumber="347">Mt Wills</location>
,
<quantity id="BE4F589A3901FF96FDBD939B7245FD18" box="[553,625,699,720]" metricMagnitude="3" metricUnit="m" metricValue="6.2" pageId="2" pageNumber="347" unit="km" value="6.2">6.2 km</quantity>
N. of
<collectingMunicipality id="996C6F053901FF96FF5E93F5711AFD22" box="[202,302,725,746]" pageId="2" pageNumber="347">Sunnyside</collectingMunicipality>
,
<collectingCounty id="90698DF33901FF96FEAE93F57169FD22" box="[314,349,725,746]" pageId="2" pageNumber="347">Vic</collectingCounty>
,
<date id="0D09D3BF3901FF96FEFE93F571D3FD22" box="[362,487,725,746]" pageId="2" pageNumber="347" value="1997-10-25">
<collectingDate id="1D4D2A573901FF96FEFE93F571D3FD22" box="[362,487,725,746]" pageId="2" pageNumber="347" value="1997-10-25">25 Oct 1997</collectingDate>
</date>
,
<collectorName id="D44290A93901FF96FE6193F57243FD22" box="[501,631,725,746]" pageId="2" pageNumber="347">R. de Keyzer</collectorName>
&amp;
<collectorName id="D44290A93901FF96FF3993D07103FCCD" box="[173,311,752,773]" pageId="2" pageNumber="347">A. Sundholm</collectorName>
(
<collectionCode id="1FA66DBA3901FF96FEDD93D0714AFCCD" box="[329,382,752,773]" pageId="2" pageNumber="347">RDK</collectionCode>
and
<collectingCountry id="01A0B5EF3901FF96FE2D93D071E8FCCD" box="[441,476,752,773]" name="American Samoa" pageId="2" pageNumber="347">AS</collectingCountry>
)
</materialsCitation>
;
<materialsCitation id="C9DFFF223901FF96FE6793D07176FC9D" collectingDate="1997-10-25" collectionCode="RDK" collectorName="R. de Keyzer &amp; A. Sundholm" country="American Samoa" location="Mt Wills" municipality="Mt Wills" pageId="2" pageNumber="347" specimenCount="6" specimenCount-male="6">
<specimenCount id="6FB13EF63901FF96FE6793D07210FCCE" box="[499,548,752,774]" pageId="2" pageNumber="347" type="male">6♂♂</specimenCount>
<location id="7C68A3A43901FF96FDBB93D072BBFCCD" LSID="urn:lsid:plazi:treatment:F11E44693902FF97FF2793657357FCF2:7C68A3A43901FF96FDBB93D072BBFCCD" box="[559,655,752,773]" country="American Samoa" municipality="Mt Wills" name="Mt Wills" pageId="2" pageNumber="347">Mt Wills</location>
,
<collectingMunicipality id="996C6F053901FF96FF39922B7124FCE8" box="[173,272,779,800]" pageId="2" pageNumber="347">Mt Wills</collectingMunicipality>
track, 200 metres below summit,
<date id="0D09D3BF3901FF96FF399205710CFCF2" box="[173,312,805,826]" pageId="2" pageNumber="347" value="1997-10-25">
<collectingDate id="1D4D2A573901FF96FF399205710CFCF2" box="[173,312,805,826]" pageId="2" pageNumber="347" value="1997-10-25">25 Oct 1997</collectingDate>
</date>
,
<collectorName id="D44290A93901FF96FEDD920571EEFCF2" box="[329,474,805,826]" pageId="2" pageNumber="347">R. de Keyzer</collectorName>
&amp;
<collectorName id="D44290A93901FF96FD91920572A6FCF2" box="[517,658,805,826]" pageId="2" pageNumber="347">A. Sundholm</collectorName>
(
<collectionCode id="1FA66DBA3901FF96FF27926070D2FC9D" box="[179,230,832,853]" pageId="2" pageNumber="347">RDK</collectionCode>
and
<collectingCountry id="01A0B5EF3901FF96FE8E9260710FFC9D" box="[282,315,832,853]" name="American Samoa" pageId="2" pageNumber="347">AS</collectingCountry>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="2DC8A5F73902FF95FF3B978E74A1F909" ID-DOI="http://doi.org/10.5281/zenodo.10101813" ID-Zenodo-Dep="10101813" box="[175,1173,1710,1729]" httpUri="https://zenodo.org/record/10101813/files/figure.png" pageId="1" pageNumber="346" startId="1.[175,215,1710,1729]" targetBox="[171,1153,794,1688]" targetPageId="1">
<paragraph id="7908F57F3902FF95FF3B978E74A1F909" blockId="1.[175,1173,1710,1729]" box="[175,1173,1710,1729]" pageId="1" pageNumber="346">
<emphasis id="4BC3296D3902FF95FF3B978E713DF909" bold="true" box="[175,265,1710,1729]" pageId="1" pageNumber="346">Figs. 12.</emphasis>
<taxonomicName id="BEB78EFC3902FF95FE8A978E71AFF909" box="[286,411,1710,1729]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="346" phylum="Arthropoda" rank="species" species="krakei">
<emphasis id="4BC3296D3902FF95FE8A978E71AFF909" box="[286,411,1710,1729]" italics="true" pageId="1" pageNumber="346">Lissotes krakei</emphasis>
</taxonomicName>
, habitus in dorsal view.
<emphasis id="4BC3296D3902FF95FDF3978E724FF909" bold="true" box="[615,635,1710,1729]" pageId="1" pageNumber="346">1)</emphasis>
Male (total length 20.2 mm);
<emphasis id="4BC3296D3902FF95FCE3978E73BEF909" bold="true" box="[887,906,1710,1729]" pageId="1" pageNumber="346">2)</emphasis>
Female (total length 18.4 mm).
</paragraph>
</caption>
<caption id="2DC8A5F73901FF96FF51931F72A0FD9A" ID-DOI="http://doi.org/10.5281/zenodo.10101815" ID-Zenodo-Dep="10101815" box="[197,660,575,594]" httpUri="https://zenodo.org/record/10101815/files/figure.png" pageId="2" pageNumber="347" startId="2.[197,229,575,594]" targetBox="[180,649,197,552]" targetPageId="2">
<paragraph id="7908F57F3901FF96FF51931F72A0FD9A" blockId="2.[197,660,575,594]" box="[197,660,575,594]" pageId="2" pageNumber="347">
<emphasis id="4BC3296D3901FF96FF51931F7135FD9A" bold="true" box="[197,257,575,594]" pageId="2" pageNumber="347">Fig. 3.</emphasis>
<taxonomicName id="BEB78EFC3901FF96FE82931F71A1FD9A" box="[278,405,575,594]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="347" phylum="Arthropoda" rank="species" species="krakei">
<emphasis id="4BC3296D3901FF96FE82931F71A1FD9A" box="[278,405,575,594]" italics="true" pageId="2" pageNumber="347">Lissotes krakei</emphasis>
</taxonomicName>
, female, head and prothorax.
</paragraph>
</caption>
<subSubSection id="31ADA6F43901FF97FF51927A71E7F9CF" lastPageId="3" lastPageNumber="348" pageId="2" pageNumber="347" type="description">
<paragraph id="7908F57F3901FF97FF51927A723CFD48" blockId="2.[173,660,645,932]" lastBlockId="3.[151,639,194,1729]" lastPageId="3" lastPageNumber="348" pageId="2" pageNumber="347">
<emphasis id="4BC3296D3901FF96FF51927A7186FCA7" bold="true" box="[197,434,858,879]" pageId="2" pageNumber="347">
Description.
<typeStatus id="A60C4BDD3901FF96FEDA927A719AFCA7" box="[334,430,858,879]" pageId="2" pageNumber="347" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="E18CE9FA3901FF96FE69927A7203FCA7" box="[509,567,858,879]" captionStart="Figs" captionStartId="1.[175,215,1710,1729]" captionTargetBox="[171,1153,794,1688]" captionTargetId="figure-253@1.[151,1172,772,1705]" captionTargetPageId="1" captionText="Figs. 12. Lissotes krakei, habitus in dorsal view. 1) Male (total length 20.2 mm); 2) Female (total length 18.4 mm)." figureDoi="http://doi.org/10.5281/zenodo.10101813" httpUri="https://zenodo.org/record/10101813/files/figure.png" pageId="2" pageNumber="347">Fig. 1</figureCitation>
). Length
<quantity id="BE4F589A3901FF96FF3992557138FC42" box="[173,268,885,906]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" pageId="2" pageNumber="347" unit="mm" value="20.2">20.2 mm</quantity>
; width (at prothorax) 8.0 mm; width (at elytra)
<quantity id="BE4F589A3901FF96FE8092AF7157FC6C" box="[276,355,911,932]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="2" pageNumber="347" unit="mm" value="7.6">7.6 mm</quantity>
. Color uniformly black.
<emphasis id="4BC3296D3901FF96FDC292AF72B8FC6C" bold="true" box="[598,652,911,932]" pageId="2" pageNumber="347">Head</emphasis>
: Form short, transverse, about twice as long as wide; clypeus very small, subtrapezoidal, flat, shiny, with some punctures and golden hairs distally; frons concave, with medial, slightly bilobate, strong protuberance; eyes rather small, canthus small, somewhat triangular, running about 1/3 length of the eye, postocular margin slightly protruding; surface of head punctate, except an almost smooth area under medial protuberance; mandibles strong, slightly longer than head, with a strong, vertically directed medial tooth on upper margin; tip of tooth curved inwards; apex pointed, inner side concave, inferior margin with a very small basal tooth followed by a large lamina with irregular border; mentum small, subrectangular, finely punctate; antennae 10-jointed, scape about as long as funicle and club together, club consisting of 3 antennomeres.
<emphasis id="4BC3296D3901FF96FC2893A67412FD53" bold="true" box="[956,1062,646,667]" pageId="2" pageNumber="347">Prothorax</emphasis>
: Transverse, large, with longitudinal, shallow medial groove; anterior and posterior borders straight, lateral margins slightly convex, anterior angles present; surface shiny with rather dense punctures, except along the anterior margin; scutellum very small, semicircular; underside strongly and finely punctate. Hindwings absent.
<emphasis id="4BC3296D3901FF96FC2A92607435FC9D" bold="true" box="[958,1025,832,853]" pageId="2" pageNumber="347">Elytra</emphasis>
: Fused, slightly longer than mandibles, head, and prothorax together; sides slightly convex; apex regularly rounded; surface with dense punctures in longitudinal rows, very short golden pubescence inside the punctures along lateral margins and at declivity.
<emphasis id="4BC3296D3900FF97FF0391D770CEFEC4" bold="true" box="[151,250,247,268]" pageId="3" pageNumber="348">Abdomen</emphasis>
: Segments strongly and finely punctate.
<emphasis id="4BC3296D3900FF97FF03903270F2FEEF" bold="true" box="[151,198,274,295]" pageId="3" pageNumber="348">Legs</emphasis>
: Short, slender; protibiae very slightly curved inwards with apical fork and 4 small teeth on the outer margin, and a strong, sharp apical spine; mesotibiae straight with a sharp postmedial tooth on the outer margin, preceded by 2 very small ones, at apex 2 spines, 1 longer than the other; metatibiae straight with a postmedial tooth on the outer margin, at apex 2 strong spines, one longer than the other; all tarsi slender.
<emphasis id="4BC3296D3900FF97FD8290C67243FE33" bold="true" box="[534,631,486,507]" pageId="3" pageNumber="348">Aedeagus</emphasis>
: Shape typical of Lucaninae, with parameres shorter than basal piece, and with very short, golden pubescence on their distal margin; permanently everted internal sac about as long as parameres and basal piece together (
<figureCitation id="E18CE9FA3900FF97FE0F934B71C8FD48" box="[411,508,619,641]" captionStart="Figs" captionStartId="2.[197,237,1710,1729]" captionTargetBox="[173,1193,992,1686]" captionTargetId="figure-298@2.[172,1193,991,1686]" captionTargetPageId="2" captionText="Figs. 46. Lissotes krakei, aedeagus. 4) Dorsal view; 5) Ventral view; 6) Lateral view." figureDoi="http://doi.org/10.5281/zenodo.10101817" httpUri="https://zenodo.org/record/10101817/files/figure.png" pageId="3" pageNumber="348">Figs. 46</figureCitation>
).
</paragraph>
<caption id="2DC8A5F73901FF96FF51978E73FEF909" ID-DOI="http://doi.org/10.5281/zenodo.10101817" ID-Zenodo-Dep="10101817" box="[197,970,1710,1729]" httpUri="https://zenodo.org/record/10101817/files/figure.png" pageId="2" pageNumber="347" startId="2.[197,237,1710,1729]" targetBox="[173,1193,992,1686]" targetPageId="2">
<paragraph id="7908F57F3901FF96FF51978E73FEF909" blockId="2.[197,970,1710,1729]" box="[197,970,1710,1729]" pageId="2" pageNumber="347">
<emphasis id="4BC3296D3901FF96FF51978E7114F909" bold="true" box="[197,288,1710,1729]" pageId="2" pageNumber="347">Figs. 46.</emphasis>
<taxonomicName id="BEB78EFC3901FF96FEA2978E7180F909" box="[310,436,1710,1729]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="347" phylum="Arthropoda" rank="species" species="krakei">
<emphasis id="4BC3296D3901FF96FEA2978E7180F909" box="[310,436,1710,1729]" italics="true" pageId="2" pageNumber="347">Lissotes krakei</emphasis>
</taxonomicName>
, aedeagus.
<emphasis id="4BC3296D3901FF96FD8F978E721AF909" bold="true" box="[539,558,1710,1729]" pageId="2" pageNumber="347">4)</emphasis>
Dorsal view;
<emphasis id="4BC3296D3901FF96FD3E978E728AF909" bold="true" box="[682,702,1710,1729]" pageId="2" pageNumber="347">5)</emphasis>
Ventral view;
<emphasis id="4BC3296D3901FF96FCAA978E7366F909" bold="true" box="[830,850,1710,1729]" pageId="2" pageNumber="347">6)</emphasis>
Lateral view.
</paragraph>
</caption>
<paragraph id="7908F57F3900FF97FF3B93A671E7F9CF" blockId="3.[151,639,194,1729]" pageId="3" pageNumber="348">
<emphasis id="4BC3296D3900FF97FF3B93A67128FD53" bold="true" box="[175,284,646,667]" pageId="3" pageNumber="348">
<typeStatus id="A60C4BDD3900FF97FF3B93A6712CFD53" box="[175,280,646,667]" pageId="3" pageNumber="348" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Males as in
<typeStatus id="A60C4BDD3900FF97FE3293A671CEFD53" box="[422,506,646,667]" pageId="3" pageNumber="348" type="holotype">holotype</typeStatus>
except varying as follows: Length 17.624.0 mm; width (at prothorax)
<quantity id="BE4F589A3900FF97FF68939B7159FD18" box="[252,365,698,720]" metricMagnitude="-3" metricUnit="m" metricValue="7.95" metricValueMax="9.4" metricValueMin="6.5" pageId="3" pageNumber="348" unit="mm" value="7.95" valueMax="9.4" valueMin="6.5">6.59.4 mm</quantity>
; width (at elytra)
<quantity id="BE4F589A3900FF97FD9D939B724EFD18" box="[521,634,698,720]" metricMagnitude="-3" metricUnit="m" metricValue="7.55" metricValueMax="8.7" metricValueMin="6.4" pageId="3" pageNumber="348" unit="mm" value="7.55" valueMax="8.7" valueMin="6.4">6.48.7 mm</quantity>
; frontal protuberance less developed in small males to distinctly bilobed in large males; mandible with vertically directed medial tooth on upper margin becoming more pronounced and apex flattened in large males; protibial outer margin with 36 small teeth; mesotibiae with 23 small, sharp teeth preceding sharp postmedial tooth on outer margin. Females (
<figureCitation id="E18CE9FA3900FF97FF6292AF7166FC6C" box="[246,338,911,933]" captionStart-0="Figs" captionStart-1="Fig" captionStartId-0="1.[175,215,1710,1729]" captionStartId-1="2.[197,229,575,594]" captionTargetBox-0="[171,1153,794,1688]" captionTargetBox-1="[180,649,197,552]" captionTargetId-0="figure-253@1.[151,1172,772,1705]" captionTargetId-1="figure-287@2.[173,659,183,552]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Figs. 12. Lissotes krakei, habitus in dorsal view. 1) Male (total length 20.2 mm); 2) Female (total length 18.4 mm)." captionText-1="Fig. 3. Lissotes krakei, female, head and prothorax." figureDoi-0="http://doi.org/10.5281/zenodo.10101813" figureDoi-1="http://doi.org/10.5281/zenodo.10101815" httpUri-0="https://zenodo.org/record/10101813/files/figure.png" httpUri-1="https://zenodo.org/record/10101815/files/figure.png" pageId="3" pageNumber="348">Figs. 23</figureCitation>
) as in
<typeStatus id="A60C4BDD3900FF97FE0992AF71DBFC6C" box="[413,495,911,932]" pageId="3" pageNumber="348" type="holotype">holotype</typeStatus>
except as follows: Length
<quantity id="BE4F589A3900FF97FE8B928A719AFC77" box="[287,430,937,959]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" metricValueMax="1.8699999999999999" metricValueMin="1.81" pageId="3" pageNumber="348" unit="mm" value="18.4" valueMax="18.7" valueMin="18.1">18.118.7 mm</quantity>
; width (at prothorax)
<quantity id="BE4F589A3900FF97FF0392E4713BFC11" box="[151,271,964,986]" metricMagnitude="-3" metricUnit="m" metricValue="7.45" metricValueMax="7.7" metricValueMin="7.2" pageId="3" pageNumber="348" unit="mm" value="7.45" valueMax="7.7" valueMin="7.2">7.27.7 mm</quantity>
; width (at elytra)
<quantity id="BE4F589A3900FF97FE5092E47208FC11" box="[452,572,964,986]" metricMagnitude="-3" metricUnit="m" metricValue="7.75" metricValueMax="8.2" metricValueMin="7.3" pageId="3" pageNumber="348" unit="mm" value="7.75" valueMax="8.2" valueMin="7.3">7.38.2 mm</quantity>
. Color uniformly black to dark reddish brown.
<emphasis id="4BC3296D3900FF97FDD292FF7249FC3C" bold="true" box="[582,637,991,1012]" pageId="3" pageNumber="348">Head</emphasis>
(
<figureCitation id="E18CE9FA3900FF97FF0B92DA70EAFBC7" box="[159,222,1018,1039]" captionStart="Fig" captionStartId="2.[197,229,575,594]" captionTargetBox="[180,649,197,552]" captionTargetId="figure-287@2.[173,659,183,552]" captionTargetPageId="2" captionText="Fig. 3. Lissotes krakei, female, head and prothorax." figureDoi="http://doi.org/10.5281/zenodo.10101815" httpUri="https://zenodo.org/record/10101815/files/figure.png" pageId="3" pageNumber="348">Fig. 3</figureCitation>
): Form short, transverse, about twice as wide as long, less than 2/3 width of pronotum; frons with concave area on each side, no medial protuberance, sparsely punctate; eyes small; surface punctate except for smooth area at middle of vertex; mandibles simple, shorter than head, with outer edge gently curved towards apex and single median tooth on inner edge; mentum with sparse, golden hairs.
<emphasis id="4BC3296D3900FF97FEFA95EE71E0FB2B" bold="true" box="[366,468,1230,1251]" pageId="3" pageNumber="348">Prothorax</emphasis>
: Anterior border and angle narrower than posterior border; surface less densely punctate along medial area of anterior margin, with shiny, very finely punctate area on both anterior sides of the medial groove.
<emphasis id="4BC3296D3900FF97FDAC94187243FA85" bold="true" box="[568,631,1336,1357]" pageId="3" pageNumber="348">Elytra</emphasis>
: Longer than mandibles, head, and prothorax together; sides more convex and apex more tapered than in males.
<emphasis id="4BC3296D3900FF97FE0F94A871FAFA55" bold="true" box="[411,462,1416,1437]" pageId="3" pageNumber="348">Legs</emphasis>
: Protibial outer margin with 46 small teeth; mesotibiae with 34 small, sharp teeth preceding much longer sharp postmedial tooth on outer margin; metatibiae with strong, sharp medial tooth.
</paragraph>
</subSubSection>
<subSubSection id="31ADA6F43900FF97FF3B972D7126F944" pageId="3" pageNumber="348" type="etymology">
<paragraph id="7908F57F3900FF97FF3B972D7126F944" blockId="3.[151,639,194,1729]" pageId="3" pageNumber="348">
<emphasis id="4BC3296D3900FF97FF3B972D7116F9EA" bold="true" box="[175,290,1549,1570]" pageId="3" pageNumber="348">Etymology.</emphasis>
This species is named in honor of Mr. Graeme J. Krake, with whom one of us (LB) has been in friendly correspondence for many years and who kindly donated a specimen of the new species.
</paragraph>
</subSubSection>
<subSubSection id="31ADA6F43900FF97FF3B97B27309FEEF" pageId="3" pageNumber="348" type="distribution">
<paragraph id="7908F57F3900FF97FF3B97B27309FEEF" blockId="3.[151,639,194,1729]" lastBlockId="3.[685,1172,189,826]" pageId="3" pageNumber="348">
<emphasis id="4BC3296D3900FF97FF3B97B27106F96F" bold="true" box="[175,306,1682,1703]" pageId="3" pageNumber="348">Distribution.</emphasis>
<taxonomicName id="BEB78EFC3900FF97FEAB97B171DCF96E" box="[319,488,1681,1702]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="krakei">
<emphasis id="4BC3296D3900FF97FEAB97B171DCF96E" box="[319,488,1681,1702]" italics="true" pageId="3" pageNumber="348">Lissotes krakei</emphasis>
</taxonomicName>
seems to be endemic to the Victorian Alps, which extends over
<quantity id="BE4F589A3900FF97FD7791E27377FF1F" box="[739,835,194,215]" metricMagnitude="6" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="348" unit="km" value="3000.0">3,000 km</quantity>
<superScript id="8EC258373900FF97FCD0919D7379FF03" attach="left" box="[836,845,189,203]" fontSize="6" pageId="3" pageNumber="348">2</superScript>
at over
<quantity id="BE4F589A3900FF97FC3191E273CCFF1F" box="[933,1016,194,215]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="348" unit="m" value="1200.0">1,200 m</quantity>
elevation. The collecting sites, Mount Wills, Buckety Plain, and Bogong High Plains, are located in Alpine National Park.
</paragraph>
</subSubSection>
<subSubSection id="31ADA6F43900FF97FD51900C7357FCF2" pageId="3" pageNumber="348" type="diagnosis">
<paragraph id="7908F57F3900FF97FD51900C7357FCF2" blockId="3.[685,1172,189,826]" pageId="3" pageNumber="348">
<emphasis id="4BC3296D3900FF97FD51900C731AFE89" bold="true" box="[709,814,300,321]" pageId="3" pageNumber="348">Diagnosis.</emphasis>
<taxonomicName id="BEB78EFC3900FF97FCA8900C73F8FE89" box="[828,972,300,321]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="krakei">
<emphasis id="4BC3296D3900FF97FCA8900C73F8FE89" box="[828,972,300,321]" italics="true" pageId="3" pageNumber="348">Lissotes krakei</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="BEB78EFC3900FF97FD399066743FFE94" authority="Benesh, 1943" authorityName="Benesh" authorityYear="1943" box="[685,1035,326,348]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="4BC3296D3900FF97FD3990667342FE93" box="[685,886,326,347]" italics="true" pageId="3" pageNumber="348">Lissotes darlingtoni</emphasis>
Benesh, 1943
</taxonomicName>
and
<taxonomicName id="BEB78EFC3900FF97FBD1906673E7FEBF" authority="Westwood, 1871" authorityName="Westwood" authorityYear="1871" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="furcicornis">
<emphasis id="4BC3296D3900FF97FBD19066732CFEBE" italics="true" pageId="3" pageNumber="348">Lissotes furcicornis</emphasis>
Westwood, 1871
</taxonomicName>
. The males of the three species can be easily distinguished by the different shapes of heads and mandibles (
<figureCitation id="E18CE9FA3900FF97FD2190917315FE0E" box="[693,801,433,455]" captionStart="Figs" captionStartId="4.[197,237,1066,1085]" captionTargetBox="[216,1193,196,1043]" captionTargetId="figure-246@4.[172,1193,183,1043]" captionTargetPageId="4" captionText="Figs. 710. Lissotes species, male heads and mandibles. 7) L. krakei; 8) L. darlingtoni; 9) L. furcicornis;" figureDoi="http://doi.org/10.5281/zenodo.10101819" httpUri="https://zenodo.org/record/10101819/files/figure.png" pageId="3" pageNumber="348">Figs. 710</figureCitation>
). In
<taxonomicName id="BEB78EFC3900FF97FCC59091739DFE0E" box="[849,937,433,454]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="krakei">
<emphasis id="4BC3296D3900FF97FCC59091739DFE0E" box="[849,937,433,454]" italics="true" pageId="3" pageNumber="348">L. krakei</emphasis>
</taxonomicName>
, the frons has a medial protuberance which is missing in the two other species; the canthus shape is also peculiar and totally different from the other taxa. The mandibles of
<taxonomicName id="BEB78EFC3900FF97FC84933B735DFDF8" box="[784,873,539,560]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="krakei">
<emphasis id="4BC3296D3900FF97FC84933B735DFDF8" box="[784,873,539,560]" italics="true" pageId="3" pageNumber="348">L. krakei</emphasis>
</taxonomicName>
(
<figureCitation id="E18CE9FA3900FF97FCEF933B738DFDF8" box="[891,953,539,560]" captionStart="Figs" captionStartId="4.[197,237,1066,1085]" captionTargetBox="[216,1193,196,1043]" captionTargetId="figure-246@4.[172,1193,183,1043]" captionTargetPageId="4" captionText="Figs. 710. Lissotes species, male heads and mandibles. 7) L. krakei; 8) L. darlingtoni; 9) L. furcicornis;" figureDoi="http://doi.org/10.5281/zenodo.10101819" httpUri="https://zenodo.org/record/10101819/files/figure.png" pageId="3" pageNumber="348">Fig. 7</figureCitation>
) are rather similar to those of
<taxonomicName id="BEB78EFC3900FF97FC99931573A9FD82" authorityName="Benesh" authorityYear="1943" box="[781,925,565,586]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="4BC3296D3900FF97FC99931573A9FD82" box="[781,925,565,586]" italics="true" pageId="3" pageNumber="348">L. darlingtoni</emphasis>
</taxonomicName>
(
<figureCitation id="E18CE9FA3900FF97FC24931673C4FD83" box="[944,1008,566,587]" captionStart="Figs" captionStartId="4.[197,237,1066,1085]" captionTargetBox="[216,1193,196,1043]" captionTargetId="figure-246@4.[172,1193,183,1043]" captionTargetPageId="4" captionText="Figs. 710. Lissotes species, male heads and mandibles. 7) L. krakei; 8) L. darlingtoni; 9) L. furcicornis;" figureDoi="http://doi.org/10.5281/zenodo.10101819" httpUri="https://zenodo.org/record/10101819/files/figure.png" pageId="3" pageNumber="348">Fig. 8</figureCitation>
), but they lack the strong basal tooth on the inferior margin; in
<taxonomicName id="BEB78EFC3900FF97FD39934B7309FD48" authorityName="Westwood" authorityYear="1871" box="[685,829,619,640]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="furcicornis">
<emphasis id="4BC3296D3900FF97FD39934B7309FD48" box="[685,829,619,640]" italics="true" pageId="3" pageNumber="348">L. furcicornis</emphasis>
</taxonomicName>
(
<figureCitation id="E18CE9FA3900FF97FCC5934B73A0FD48" box="[849,916,619,640]" captionStart="Figs" captionStartId="4.[197,237,1066,1085]" captionTargetBox="[216,1193,196,1043]" captionTargetId="figure-246@4.[172,1193,183,1043]" captionTargetPageId="4" captionText="Figs. 710. Lissotes species, male heads and mandibles. 7) L. krakei; 8) L. darlingtoni; 9) L. furcicornis;" figureDoi="http://doi.org/10.5281/zenodo.10101819" httpUri="https://zenodo.org/record/10101819/files/figure.png" pageId="3" pageNumber="348">Fig. 9</figureCitation>
), the mandibles have a much stronger medial vertical tooth, and the apex is not sharply pointed but has a double point. The female of
<taxonomicName id="BEB78EFC3900FF97FC17939A73E8FD07" box="[899,988,698,719]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="krakei">
<emphasis id="4BC3296D3900FF97FC17939A73E8FD07" box="[899,988,698,719]" italics="true" pageId="3" pageNumber="348">L. krakei</emphasis>
</taxonomicName>
can be easily distinguished from the females of
<taxonomicName id="BEB78EFC3900FF97FB9793F574A0FD22" authorityName="Benesh" authorityYear="1943" box="[1027,1172,725,746]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="4BC3296D3900FF97FB9793F574A0FD22" box="[1027,1172,725,746]" italics="true" pageId="3" pageNumber="348">L. darlingtoni</emphasis>
</taxonomicName>
and
<taxonomicName id="BEB78EFC3900FF97FD4893D07350FCCD" authorityName="Westwood" authorityYear="1871" box="[732,868,752,773]" class="Insecta" family="Lucanidae" genus="Lissotes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="furcicornis">
<emphasis id="4BC3296D3900FF97FD4893D07350FCCD" box="[732,868,752,773]" italics="true" pageId="3" pageNumber="348">L. furcicornis</emphasis>
</taxonomicName>
by the shape of the canthus and by the punctures and shape of the head and pronotum (
<figureCitation id="E18CE9FA3900FF97FC8E92057363FCF2" box="[794,855,805,826]" captionStart="Fig" captionStartId="2.[197,229,575,594]" captionTargetBox="[180,649,197,552]" captionTargetId="figure-287@2.[173,659,183,552]" captionTargetPageId="2" captionText="Fig. 3. Lissotes krakei, female, head and prothorax." figureDoi="http://doi.org/10.5281/zenodo.10101815" httpUri="https://zenodo.org/record/10101815/files/figure.png" pageId="3" pageNumber="348">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>