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<document id="15AD79385A2CBAB6CDE66E4D29954B22" ID-CLB-Dataset="22198" ID-DOI="10.5852/ejt.2017.277" ID-GBIF-Dataset="d168a1e0-88f9-48ab-a871-cc2d24b026d7" ID-ISSN="2118-9773" ID-Zenodo-Dep="3824583" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589383917281" checkinUser="carolina" docAuthor="Hsiao, Yun, Yu, Yali, Deng, Congshuang &amp; 4, Hong Pang" docDate="2017" docId="E55BF87FFFA33F7DFE45E5097483FD11" docLanguage="en" docName="ejt-2017-277.pdf.imf" docOrigin="European Journal of Taxonomy 277" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng 2017, sp. nov." docType="treatment" docUuid="F5AFB48F-0494-4CEC-B7F6-2B1742AC4F34" docUuidSource="ZooBank" docVersion="7" lastPageNumber="8" masterDocId="19628007FFA03F7AFFA4E4027002FF8E" masterDocTitle="The first fossil wedge-shaped beetle (Coleoptera, Ripiphoridae) from the middle Jurassic of China" masterLastPageNumber="13" masterPageNumber="1" pageNumber="4" updateTime="1698839382826" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="C16D8EF94E922B2FD81289A0CA973485" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8991CFD2249F7B9751D8F7735C62122E">
<mods:title id="435C94B21026E69C18B70E36642E8A0B">The first fossil wedge-shaped beetle (Coleoptera, Ripiphoridae) from the middle Jurassic of China</mods:title>
</mods:titleInfo>
<mods:name id="A3D7CD1FE87B0435E4CDEAE0C62E3452" type="personal">
<mods:role id="D41448A4E51BA374FFFDB5AF0E4F76FF">
<mods:roleTerm id="C47B133E0A3D90C67050DBA8F38DD256">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27C24DCC3D5B4944DD1F92B8C40543F7">Hsiao, Yun</mods:namePart>
</mods:name>
<mods:name id="06F24952F5C7041011CAE7BE3076139F" type="personal">
<mods:role id="0648F7216F10FA05C585BC89F442B436">
<mods:roleTerm id="49AD4E47B9320E33277031FF468C777C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FEBA984878631411F15EC649799DEA72">Yu, Yali</mods:namePart>
</mods:name>
<mods:name id="47484067A3B939AF4BEE6576FC76C841" type="personal">
<mods:role id="349A330EC22B321CE75C9063479D8E17">
<mods:roleTerm id="3066982A8F2AEEE592BBE32BC44E0F30">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E61BB1058C46D59F64773C756D3D7CEC">Deng, Congshuang</mods:namePart>
</mods:name>
<mods:name id="6061DA218AF14DFD7103CE5B16FECE0F" type="personal">
<mods:role id="F53FB1A4BE4E78E36665209445D5DFDA">
<mods:roleTerm id="9C55E75F6A1BDDFD38E06266B4B24EF5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="216EA54F62D1F486F55BAC90A3F8ED6C">4, Hong Pang</mods:namePart>
</mods:name>
<mods:typeOfResource id="EBED441588FE13D439E0DEA8A1A27468">text</mods:typeOfResource>
<mods:relatedItem id="5C2D0DF665893B6F5A0AD375CACBEAFE" type="host">
<mods:titleInfo id="35603525EE864A02DB9EBB4DBCE61DE7">
<mods:title id="FB0087F40B6EE1D4C5754F2DD2E0E8C6">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="A07148AB78248D47E83238191D396F4D">
<mods:date id="591190AF0A7A5BBBE1853CA451753227">2017</mods:date>
<mods:detail id="AA89DE1CBEE8BED023E8B2FAC52DB2F9" type="pubDate">
<mods:number id="833582AE78BFB63FA464AF55B62EA0E8">2017-02-14</mods:number>
</mods:detail>
<mods:detail id="A0DDB98F62E3691CACC6C2722D9427A4" type="issue">
<mods:number id="3B89FF6DC7E96B20F61F81529CCBCE7F">277</mods:number>
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<mods:start id="1D55606129C9DBE181E81D551DBF5CB8">1</mods:start>
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<mods:classification id="8DADC88C0657BF863B6FFA468F7450DC">journal article</mods:classification>
<mods:identifier id="223EE493FE714822B047CA2958A27A69" type="CLB-Dataset">22198</mods:identifier>
<mods:identifier id="C47BFB6A0985C24B9F00B43C396AAFAB" type="DOI">10.5852/ejt.2017.277</mods:identifier>
<mods:identifier id="8FE83CC4C84D525085D50F6401A99034" type="GBIF-Dataset">d168a1e0-88f9-48ab-a871-cc2d24b026d7</mods:identifier>
<mods:identifier id="3B9934C8CB64BF333827A03EE9DB9383" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="FF1CD593654BEDAD49E81CD00CA90A6B" type="Zenodo-Dep">3824583</mods:identifier>
<mods:identifier id="8A9478EE526172A249E5A79261AEA0C8" type="ZooBank">D4E1AD48-FE68-4C9B-91B1-BC6E918B5714</mods:identifier>
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<treatment id="E55BF87FFFA33F7DFE45E5097483FD11" ID-DOI="http://doi.org/10.5281/zenodo.3845817" ID-GBIF-Taxon="164129384" ID-Zenodo-Dep="3845817" LSID="urn:lsid:zoobank.org:act:F5AFB48F-0494-4CEC-B7F6-2B1742AC4F34" httpUri="http://treatment.plazi.org/id/E55BF87FFFA33F7DFE45E5097483FD11" lastPageId="7" lastPageNumber="8" pageId="3" pageNumber="4">
<subSubSection id="25E81AE2FFA33F79FE45E5097450FEA8" box="[481,1106,267,294]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="6D4D4969FFA33F79FE45E5097450FEA8" blockId="3.[366,1222,267,364]" box="[481,1106,267,294]" pageId="3" pageNumber="4">
<heading id="3605FE05FFA33F79FE45E5097450FEA8" box="[481,1106,267,294]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
<taxonomicName id="AAF232EAFFA33F79FE45E50973F3FEA8" authority="Hsiao, Yu &amp; Deng" authorityName="Hsiao, Yu &amp; Deng" authorityYear="2017" box="[481,1009,267,294]" class="Insecta" family="Ripiphoridae" genus="Archaeoripiphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nuwa" status="sp. nov.">
<emphasis id="5F86957BFFA33F79FE45E5097310FEA8" bold="true" box="[481,786,267,294]" italics="true" pageId="3" pageNumber="4">Archaeoripiphorus nuwa</emphasis>
Hsiao, Yu &amp; Deng
</taxonomicName>
<taxonomicNameLabel id="44B52800FFA33F79FC5CE50E7450FEA8" box="[1016,1106,268,294]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25E81AE2FFA33F79FECAE52C734EFEE2" pageId="3" pageNumber="4" type="description">
<paragraph id="6D4D4969FFA33F79FECAE52C74C4FEC7" blockId="3.[366,1222,267,364]" box="[366,1222,302,329]" pageId="3" pageNumber="4">
<uri id="1963456BFFA33F79FECAE52C74C4FEC7" box="[366,1222,302,329]" pageId="3" pageNumber="4">
urn:lsid:zoobank.org:act:
<uuid id="195473BCFFA33F79FD28E52C74C4FEC7" box="[652,1222,302,329]" pageId="3" pageNumber="4">F5AFB48F-0494-4CEC-B7F6-2B1742AC4F34</uuid>
</uri>
</paragraph>
<paragraph id="6D4D4969FFA33F79FD43E550734EFEE2" blockId="3.[366,1222,267,364]" box="[743,844,338,364]" pageId="3" pageNumber="4">
<figureCitation id="F5C955ECFFA33F79FD43E550734EFEE2" box="[743,844,338,364]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="3.[189,232,1964,1990]" captionStartId-1="4.[189,232,1965,1991]" captionStartId-2="5.[189,232,1930,1956]" captionStartId-3="6.[189,232,1928,1954]" captionTargetBox-0="[342,1246,569,1928]" captionTargetBox-1="[189,1399,291,1903]" captionTargetBox-2="[189,1398,773,1882]" captionTargetBox-3="[194,1390,1008,1882]" captionTargetId-0="figure@3.[342,1246,569,1928]" captionTargetId-1="figure@4.[189,1399,289,1918]" captionTargetId-2="figure@5.[189,1398,769,1888]" captionTargetId-3="figure@6.[188,1398,998,1886]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Fig. 1. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype, habitus, dorsal view. Scale bar = 5.0 mm. Photograph of dry specimen." captionText-1="Fig. 2. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A. Photograph of complete specimen. B. Habitus drawing. Scale bars = 5.0 mm." captionText-2="Fig. 3. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A. Head. B. Prothorax. C. Elytron. D. Antenna. E. Photograph of the surface of the body. Scale bars: A–C = 1.0 mm; D = 0.1 mm; E = 0.5 mm. The black arrows indicate the magnified portions." captionText-3="Fig. 4. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A–F. Tibial spurs (indicated by the red arrows). A–B. Fore leg. C–D. Mid leg. E–F. Hind leg. G–H. Tarsal claw of mid leg. A, C, E, G. Photographs. B, D, F, H. Hand drawings. Scale bars: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3824585" figureDoi-1="http://doi.org/10.5281/zenodo.3824587" figureDoi-2="http://doi.org/10.5281/zenodo.3825347" figureDoi-3="http://doi.org/10.5281/zenodo.3824593" httpUri-0="https://zenodo.org/record/3824585/files/figure.png" httpUri-1="https://zenodo.org/record/3824587/files/figure.png" httpUri-2="https://zenodo.org/record/3825347/files/figure.png" httpUri-3="https://zenodo.org/record/3824593/files/figure.png" pageId="3" pageNumber="4">Figs 1–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="25E81AE2FFA33F7FFF19E59172BFFEC6" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="6D4D4969FFA33F79FF19E5917134FE23" blockId="3.[189,310,403,429]" box="[189,310,403,429]" pageId="3" pageNumber="4">
<heading id="3605FE05FFA33F79FF19E5917134FE23" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="5F86957BFFA33F79FF19E5917134FE23" bold="true" box="[189,310,403,429]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6D4D4969FFA33F79FF19E5C07574FE71" blockId="3.[189,1398,450,512]" pageId="3" pageNumber="4">
As for the genus (
<emphasis id="5F86957BFFA33F79FE07E5C07221FE52" box="[419,547,450,476]" italics="true" pageId="3" pageNumber="4">vide supra</emphasis>
). In particular this species resembles the extant ptilophorine species,
<taxonomicName id="AAF232EAFFA33F79FF19E5E772E7FD8E" authority="(Nomura &amp; Nakane, 1959)" baseAuthorityName="Nomura &amp; Nakane" baseAuthorityYear="1959" box="[189,741,485,512]" class="Insecta" family="Ripiphoridae" genus="Trigonodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tokejii">
<emphasis id="5F86957BFFA33F79FF19E5E771A2FE71" box="[189,416,485,511]" italics="true" pageId="3" pageNumber="4">Trigonodera tokejii</emphasis>
(Nomura &amp; Nakane, 1959)
</taxonomicName>
in general appearance (
<figureCitation id="F5C955ECFFA33F79FC5AE5E77444FD8E" box="[1022,1094,485,512]" captionStart="Fig" captionStartId="7.[189,232,1929,1955]" captionTargetBox="[216,1354,1162,1854]" captionTargetId="figure@7.[189,1398,1103,1888]" captionTargetPageId="7" captionText="Fig. 5. An extant species of Ptilophorinae, Trigonodera tokejii (Nomura &amp; Nakane, 1959). A. Habitus, dorsal view. B. Habitus, lateral view. C. Head, frontal view. D. Antenna. Scale bars: A–B = 5.0 mm; C–D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3824595" httpUri="https://zenodo.org/record/3824595/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
), from which it differs by
</paragraph>
<caption id="398D19E1FFA33F79FF19E3AE72BFF867" ID-DOI="http://doi.org/10.5281/zenodo.3824585" ID-Zenodo-Dep="3824585" httpUri="https://zenodo.org/record/3824585/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1964,1990]" targetBox="[342,1246,569,1928]" targetPageId="3">
<paragraph id="6D4D4969FFA33F79FF19E3AE72BFF867" blockId="3.[189,1399,1964,2026]" pageId="3" pageNumber="4">
<emphasis id="5F86957BFFA33F79FF19E3AE710BF848" bold="true" box="[189,265,1964,1990]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
<taxonomicName id="AAF232EAFFA33F79FEABE3AE730CF848" authority="Hsiao, Yu &amp; Deng" authorityName="Hsiao, Yu &amp; Deng" authorityYear="2017" box="[271,782,1964,1990]" class="Insecta" family="Ripiphoridae" genus="Archaeoripiphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nuwa" status="gen. et sp. nov.">
<emphasis id="5F86957BFFA33F79FEABE3AE7236F848" box="[271,564,1964,1990]" italics="true" pageId="3" pageNumber="4">Archaeoripiphorus nuwa</emphasis>
Hsiao, Yu &amp; Deng
</taxonomicName>
<taxonomicNameLabel id="44B52800FFA33F79FCB0E3AE73C1F848" box="[788,963,1964,1990]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype, habitus, dorsal view. Scale bar = 5.0 mm. Photograph of dry specimen.
</paragraph>
</caption>
<caption id="398D19E1FFA43F7EFF19E3AF7324F864" ID-DOI="http://doi.org/10.5281/zenodo.3824587" ID-Zenodo-Dep="3824587" httpUri="https://zenodo.org/record/3824587/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1965,1991]" targetBox="[189,1399,291,1903]" targetPageId="4">
<paragraph id="6D4D4969FFA43F7EFF19E3AF7324F864" blockId="4.[189,1399,1965,2027]" pageId="4" pageNumber="5">
<emphasis id="5F86957BFFA43F7EFF19E3AF710BF849" bold="true" box="[189,265,1965,1991]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
<taxonomicName id="AAF232EAFFA43F7EFEABE3AF730EF849" authority="Hsiao, Yu &amp; Deng" authorityName="Hsiao, Yu &amp; Deng" authorityYear="2017" box="[271,780,1965,1991]" class="Insecta" family="Ripiphoridae" genus="Archaeoripiphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nuwa" status="gen. et sp. nov.">
<emphasis id="5F86957BFFA43F7EFEABE3AF7231F849" box="[271,563,1965,1991]" italics="true" pageId="4" pageNumber="5">Archaeoripiphorus nuwa</emphasis>
Hsiao, Yu &amp; Deng
</taxonomicName>
<taxonomicNameLabel id="44B52800FFA43F7EFCB5E3AF73BDF849" box="[785,959,1965,1991]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="5F86957BFFA43F7EFB99E3AF7450F849" bold="true" box="[1085,1106,1965,1991]" pageId="4" pageNumber="5">A</emphasis>
. Photograph of complete specimen.
<emphasis id="5F86957BFFA43F7EFE9EE3D2714CF864" bold="true" box="[314,334,2000,2026]" pageId="4" pageNumber="5">B</emphasis>
. Habitus drawing. Scale bars = 5.0 mm.
</paragraph>
</caption>
<paragraph id="6D4D4969FFA53F7FFF19E50972BFFEC6" blockId="5.[189,1398,267,329]" pageId="5" pageNumber="6">its shorter antennae and terminal maxillary palpomere, wider pronotum anteriorly, shorter elytra, and tibial edges without spiniform seta apically.</paragraph>
</subSubSection>
<subSubSection id="25E81AE2FFA53F7FFF19E57273D4FD9C" pageId="5" pageNumber="6" type="etymology">
<paragraph id="6D4D4969FFA53F7FFF19E5727146FE04" blockId="5.[189,324,368,394]" box="[189,324,368,394]" pageId="5" pageNumber="6">
<heading id="3605FE05FFA53F7FFF19E5727146FE04" bold="true" box="[189,324,368,394]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="5F86957BFFA53F7FFF19E5727146FE04" bold="true" box="[189,324,368,394]" pageId="5" pageNumber="6">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6D4D4969FFA53F7FFF19E5B373D4FD9C" blockId="5.[189,1399,433,530]" pageId="5" pageNumber="6">The specific name honors the great goddess in the ancient Chinese mythology, Nüwa, who created mankind and saved humanity from a major catastrophe by using the five-colored stones to repair the damage to the pillar of heaven caused by the water god, Gonggong.</paragraph>
</subSubSection>
<subSubSection id="25E81AE2FFA53F7FFF19E63B74FEFD28" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="6D4D4969FFA53F7FFF19E63B716EFDDD" blockId="5.[189,364,569,595]" box="[189,364,569,595]" pageId="5" pageNumber="6">
<heading id="3605FE05FFA53F7FFF19E63B716EFDDD" bold="true" box="[189,364,569,595]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="5F86957BFFA53F7FFF19E63B716EFDDD" bold="true" box="[189,364,569,595]" pageId="5" pageNumber="6">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="DD9A4334FFA53F7FFF4EE66A74FEFD28" ID-GBIF-Occurrence="2612932301" country="China" location="CHINA" pageId="5" pageNumber="6" specimenCode="CNU-C-NN-2006841" specimenCount="1" typeStatus="holotype">
<paragraph id="6D4D4969FFA53F7FFF4EE66A715EFD0C" blockId="5.[189,1276,616,678]" box="[234,348,616,642]" pageId="5" pageNumber="6">
<emphasis id="5F86957BFFA53F7FFF4EE66A715EFD0C" bold="true" box="[234,348,616,642]" pageId="5" pageNumber="6">
<typeStatus id="B249F7CBFFA53F7FFF4EE66A715EFD0C" box="[234,348,616,642]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D4D4969FFA53F7FFF19E68974FEFD28" blockId="5.[189,1276,616,678]" box="[189,1276,651,678]" pageId="5" pageNumber="6">
<collectingCountry id="15E509F9FFA53F7FFF19E689711DFD2B" box="[189,287,651,677]" name="China" pageId="5" pageNumber="6">CHINA</collectingCountry>
: nearly complete specimen in dorsal position; sex
<taxonomicName id="AAF232EAFFA53F7FFCC9E68973D8FD2B" box="[877,986,651,677]" class="Insecta" family="Ripiphoridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Unknown">unknown</taxonomicName>
(
<specimenCode id="3D54E112FFA53F7FFC4DE68974F2FD28" box="[1001,1264,651,678]" pageId="5" pageNumber="6">CNU-C-NN-2006841</specimenCode>
).
</paragraph>
</materialsCitation>
</subSubSection>
<caption id="398D19E1FFA53F7FFF19E3887404F865" ID-DOI="http://doi.org/10.5281/zenodo.3825347" ID-Zenodo-Dep="3825347" httpUri="https://zenodo.org/record/3825347/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1930,1956]" targetBox="[189,1398,773,1882]" targetPageId="5">
<paragraph id="6D4D4969FFA53F7FFF19E3887404F865" blockId="5.[189,1399,1930,2028]" pageId="5" pageNumber="6">
<emphasis id="5F86957BFFA53F7FFF19E388710EF82A" bold="true" box="[189,268,1930,1956]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="AAF232EAFFA53F7FFEB1E3897323F82B" authority="Hsiao, Yu &amp; Deng" authorityName="Hsiao, Yu &amp; Deng" authorityYear="2017" box="[277,801,1930,1957]" class="Insecta" family="Ripiphoridae" genus="Archaeoripiphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nuwa" status="gen. et sp. nov.">
<emphasis id="5F86957BFFA53F7FFEB1E389723EF82A" box="[277,572,1931,1957]" italics="true" pageId="5" pageNumber="6">Archaeoripiphorus nuwa</emphasis>
Hsiao, Yu &amp; Deng
</taxonomicName>
<taxonomicNameLabel id="44B52800FFA53F7FFC8DE38973E2F82B" box="[809,992,1931,1957]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="5F86957BFFA53F7FFBC0E388747BF82A" bold="true" box="[1124,1145,1930,1956]" pageId="5" pageNumber="6">A</emphasis>
. Head.
<emphasis id="5F86957BFFA53F7FFB7CE38974EEF82B" bold="true" box="[1240,1260,1931,1957]" pageId="5" pageNumber="6">B</emphasis>
. Prothorax.
<emphasis id="5F86957BFFA53F7FFF19E3AC70D0F846" bold="true" box="[189,210,1966,1992]" pageId="5" pageNumber="6">C</emphasis>
. Elytron.
<emphasis id="5F86957BFFA53F7FFEEAE3AC7161F846" bold="true" box="[334,355,1966,1992]" pageId="5" pageNumber="6">D</emphasis>
. Antenna.
<emphasis id="5F86957BFFA53F7FFE4DE3AC71FFF846" bold="true" box="[489,509,1966,1992]" pageId="5" pageNumber="6">E</emphasis>
. Photograph of the surface of the body. Scale bars: A–C = 1.0 mm; D = 0.1 mm; E = 0.5 mm. The black arrows indicate the magnified portions.
</paragraph>
</caption>
<subSubSection id="25E81AE2FFA63F7CFF19E509742CFE4A" pageId="6" pageNumber="7" type="distribution">
<paragraph id="6D4D4969FFA63F7CFF19E509715FFEAB" blockId="6.[189,349,267,293]" box="[189,349,267,293]" pageId="6" pageNumber="7">
<heading id="3605FE05FFA63F7CFF19E509715FFEAB" bold="true" box="[189,349,267,293]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="5F86957BFFA63F7CFF19E509715FFEAB" bold="true" box="[189,349,267,293]" pageId="6" pageNumber="7">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="6D4D4969FFA63F7CFF19E53874CBFEDD" blockId="6.[189,1225,313,340]" box="[189,1225,313,340]" pageId="6" pageNumber="7">
<collectingCountry id="15E509F9FFA63F7CFF19E538711DFEDA" box="[189,287,314,340]" name="China" pageId="6" pageNumber="7">CHINA</collectingCountry>
: NE China, Daohugou, Shantou Township, Ningcheng County,
<collectingRegion id="AF36878BFFA63F7CFBADE53874C7FEDD" box="[1033,1221,313,340]" country="China" name="Nei Mongol" pageId="6" pageNumber="7">Inner Mongolia</collectingRegion>
.
</paragraph>
<paragraph id="6D4D4969FFA63F7CFF19E5797160FE1B" blockId="6.[189,354,379,405]" box="[189,354,379,405]" pageId="6" pageNumber="7">
<heading id="3605FE05FFA63F7CFF19E5797160FE1B" bold="true" box="[189,354,379,405]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="5F86957BFFA63F7CFF19E5797160FE1B" bold="true" box="[189,354,379,405]" pageId="6" pageNumber="7">Type horizon</emphasis>
</heading>
</paragraph>
<paragraph id="6D4D4969FFA63F7CFF19E5AB742CFE4A" blockId="6.[189,1070,425,452]" box="[189,1070,425,452]" pageId="6" pageNumber="7">
Jiulongshan Formation, Middle Jurassic, about 165 Ma (
<bibRefCitation id="09633498FFA63F7CFCF7E5AB7420FE4A" author="Gao K. - Q. &amp; Ren D." box="[851,1058,425,452]" pageId="6" pageNumber="7" pagination="42 - 45" refId="ref4926" refString="Gao K. - Q. &amp; Ren D. 2006. Radiometric dating of ignimbrite from Inner Mongolia provides no indication of a post-Middle Jurassic Age for the Daohugou Beds. Acta Geologica Sinica 80: 42 - 45. http: // dx. doi. org / 10.1111 / j. 1755 - 6724.2006. tb 00793. x" type="journal article" year="2006">Gao &amp; Ren 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="25E81AE2FFA63F7DFF19E5E97483FD11" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="6D4D4969FFA63F7CFF19E5E9714CFD8B" blockId="6.[189,334,491,517]" box="[189,334,491,517]" pageId="6" pageNumber="7">
<heading id="3605FE05FFA63F7CFF19E5E9714CFD8B" bold="true" box="[189,334,491,517]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="5F86957BFFA63F7CFF19E5E9714CFD8B" bold="true" box="[189,334,491,517]" pageId="6" pageNumber="7">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6D4D4969FFA63F7CFF19E6187229FDF4" blockId="6.[189,1399,537,634]" pageId="6" pageNumber="7">
BODY (
<figureCitation id="F5C955ECFFA63F7CFEB7E61B7183FDBD" box="[275,385,537,563]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[189,232,1964,1990]" captionStartId-1="4.[189,232,1965,1991]" captionTargetBox-0="[342,1246,569,1928]" captionTargetBox-1="[189,1399,291,1903]" captionTargetId-0="figure@3.[342,1246,569,1928]" captionTargetId-1="figure@4.[189,1399,289,1918]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype, habitus, dorsal view. Scale bar = 5.0 mm. Photograph of dry specimen." captionText-1="Fig. 2. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A. Photograph of complete specimen. B. Habitus drawing. Scale bars = 5.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3824585" figureDoi-1="http://doi.org/10.5281/zenodo.3824587" httpUri-0="https://zenodo.org/record/3824585/files/figure.png" httpUri-1="https://zenodo.org/record/3824587/files/figure.png" pageId="6" pageNumber="7">Figs 1–2</figureCitation>
). Length
<quantity id="AA0AE48CFFA63F7CFE5FE61B726EFDBA" box="[507,620,537,564]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="6" pageNumber="7" unit="mm" value="15.5">15.5 mm</quantity>
, width 5.0 mm, head length 2.0 mm, antennal length
<quantity id="AA0AE48CFFA63F7CFAB4E61B7570FDBA" box="[1296,1394,537,564]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="6" pageNumber="7" unit="mm" value="3.4">3.4 mm</quantity>
, pronotum length
<quantity id="AA0AE48CFFA63F7CFE29E63E71EFFDD9" box="[397,493,572,599]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="7" unit="mm" value="3.5">3.5 mm</quantity>
, pronotum width 4.0 mm, elytra length 10.0 mm. Surface densely covered with fine pubescence (
<figureCitation id="F5C955ECFFA63F7CFE67E662721CFDF4" box="[451,542,608,634]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1398,773,1882]" captionTargetId="figure@5.[189,1398,769,1888]" captionTargetPageId="5" captionText="Fig. 3. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A. Head. B. Prothorax. C. Elytron. D. Antenna. E. Photograph of the surface of the body. Scale bars: A–C = 1.0 mm; D = 0.1 mm; E = 0.5 mm. The black arrows indicate the magnified portions." figureDoi="http://doi.org/10.5281/zenodo.3825347" httpUri="https://zenodo.org/record/3825347/files/figure.png" pageId="6" pageNumber="7">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="6D4D4969FFA63F7CFF19E6A0749EFC3C" blockId="6.[189,1399,673,946]" pageId="6" pageNumber="7">
HEAD (
<figureCitation id="F5C955ECFFA63F7CFEB7E6A37174FD35" box="[275,374,673,700]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1398,773,1882]" captionTargetId="figure@5.[189,1398,769,1888]" captionTargetPageId="5" captionText="Fig. 3. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A. Head. B. Prothorax. C. Elytron. D. Antenna. E. Photograph of the surface of the body. Scale bars: A–C = 1.0 mm; D = 0.1 mm; E = 0.5 mm. The black arrows indicate the magnified portions." figureDoi="http://doi.org/10.5281/zenodo.3825347" httpUri="https://zenodo.org/record/3825347/files/figure.png" pageId="6" pageNumber="7">Fig. 3A</figureCitation>
). Small and elongate, abruptly constricted posteriorly to form broad neck; surface sparsely punctate. Eyes oval, shallowly emarginate, distinctly separated from each other, ratio of eye diameter to interocular space 1.00:1.65; surface finely faceted. Frontoclypeal suture indistinct. Apical margin of clypeus slightly sinuate. Labrum rounded, subquadrate. Terminal maxillary palpomere elongate securiform, about 4.0 times as long as minimum width. Antennal insertions widely separated, exposed from above. Antennae 11-segmented. Scape and pedicel very short, antennomere III strongly elongate, IV to X rectangular or trapezoidal, antennomere XI with pointed apex (
<figureCitation id="F5C955ECFFA63F7CFBD7E77674D3FC00" box="[1139,1233,884,910]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1398,773,1882]" captionTargetId="figure@5.[189,1398,769,1888]" captionTargetPageId="5" captionText="Fig. 3. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A. Head. B. Prothorax. C. Elytron. D. Antenna. E. Photograph of the surface of the body. Scale bars: A–C = 1.0 mm; D = 0.1 mm; E = 0.5 mm. The black arrows indicate the magnified portions." figureDoi="http://doi.org/10.5281/zenodo.3825347" httpUri="https://zenodo.org/record/3825347/files/figure.png" pageId="6" pageNumber="7">Fig. 3D</figureCitation>
). Length ratio of antennomeres as follows: 1.20:1.00:2.30:1.35:1.20:1.35:1.10:0.90:0.90:0.90:1.45.
</paragraph>
<caption id="398D19E1FFA63F7CFF19E38A73B0F864" ID-DOI="http://doi.org/10.5281/zenodo.3824593" ID-Zenodo-Dep="3824593" httpUri="https://zenodo.org/record/3824593/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1928,1954]" targetBox="[194,1390,1008,1882]" targetPageId="6">
<paragraph id="6D4D4969FFA63F7CFF19E38A73B0F864" blockId="6.[189,1399,1928,2026]" pageId="6" pageNumber="7">
<emphasis id="5F86957BFFA63F7CFF19E38A710AF82D" bold="true" box="[189,264,1928,1955]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
<taxonomicName id="AAF232EAFFA63F7CFEA9E38B7304F82D" authority="Hsiao, Yu &amp; Deng" authorityName="Hsiao, Yu &amp; Deng" authorityYear="2017" box="[269,774,1929,1955]" class="Insecta" family="Ripiphoridae" genus="Archaeoripiphorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nuwa" status="gen. et sp. nov.">
<emphasis id="5F86957BFFA63F7CFEA9E38B7232F82D" box="[269,560,1929,1955]" italics="true" pageId="6" pageNumber="7">Archaeoripiphorus nuwa</emphasis>
Hsiao, Yu &amp; Deng
</taxonomicName>
<taxonomicNameLabel id="44B52800FFA63F7CFCAFE38B73B4F82D" box="[779,950,1929,1955]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype.
<emphasis id="5F86957BFFA63F7CFB97E38A746BF82C" bold="true" box="[1075,1129,1928,1954]" pageId="6" pageNumber="7">A–F</emphasis>
. Tibial spurs (indicated by the red arrows).
<emphasis id="5F86957BFFA63F7CFE04E3AE71D5F848" bold="true" box="[416,471,1964,1990]" pageId="6" pageNumber="7">A–B</emphasis>
. Fore leg.
<emphasis id="5F86957BFFA63F7CFDF1E3AE728CF848" bold="true" box="[597,654,1964,1990]" pageId="6" pageNumber="7">C–D</emphasis>
. Mid leg.
<emphasis id="5F86957BFFA63F7CFCA2E3AE7338F848" bold="true" box="[774,826,1964,1990]" pageId="6" pageNumber="7">E–F</emphasis>
. Hind leg.
<emphasis id="5F86957BFFA63F7CFC19E3AE73FBF848" bold="true" box="[957,1017,1964,1990]" pageId="6" pageNumber="7">G–H</emphasis>
. Tarsal claw of mid leg. A, C, E, G. Photographs. B, D, F, H. Hand drawings. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="6D4D4969FFA73F7DFF19E50E7318FE01" blockId="7.[189,1399,267,399]" pageId="7" pageNumber="8">
PROTHORAX (
<figureCitation id="F5C955ECFFA73F7DFEF4E50971A9FEAB" box="[336,427,267,294]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1398,773,1882]" captionTargetId="figure@5.[189,1398,769,1888]" captionTargetPageId="5" captionText="Fig. 3. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A. Head. B. Prothorax. C. Elytron. D. Antenna. E. Photograph of the surface of the body. Scale bars: A–C = 1.0 mm; D = 0.1 mm; E = 0.5 mm. The black arrows indicate the magnified portions." figureDoi="http://doi.org/10.5281/zenodo.3825347" httpUri="https://zenodo.org/record/3825347/files/figure.png" pageId="7" pageNumber="8">Fig. 3B</figureCitation>
). Almost triangular, about 0.88 times as long as wide, slightly narrower than elytra at base, tapering anteriorly; anterior margin truncate and straight; anterior angles rounded and indistinct, posterior margin trilobed, with posterior angles moderately protruding and obtuse; dorsal surface densely punctate. Scutellar shield minute, angulate apically.
</paragraph>
<paragraph id="6D4D4969FFA73F7DFF19E5B57415FE7D" blockId="7.[189,1399,438,500]" pageId="7" pageNumber="8">
ELYTRA (
<figureCitation id="F5C955ECFFA73F7DFE81E5B47180FE5E" box="[293,386,438,465]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1398,773,1882]" captionTargetId="figure@5.[189,1398,769,1888]" captionTargetPageId="5" captionText="Fig. 3. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A. Head. B. Prothorax. C. Elytron. D. Antenna. E. Photograph of the surface of the body. Scale bars: A–C = 1.0 mm; D = 0.1 mm; E = 0.5 mm. The black arrows indicate the magnified portions." figureDoi="http://doi.org/10.5281/zenodo.3825347" httpUri="https://zenodo.org/record/3825347/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
). About 2 times as long as wide, lateral sides slightly narrowing posteriorly; surface densely and regularly punctate. Abdomen with five tergites and ventrites.
</paragraph>
<paragraph id="6D4D4969FFA73F7DFF19E61E7483FD11" blockId="7.[189,1398,539,671]" pageId="7" pageNumber="8">
LEGS. Long and slender; tibiae moderately widening apically, without spiniform seta at apices, tibial spur formula 1-1-2 (
<figureCitation id="F5C955ECFFA73F7DFE0DE63C71ECFDD6" box="[425,494,574,600]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[194,1390,1008,1882]" captionTargetId="figure@6.[188,1398,998,1886]" captionTargetPageId="6" captionText="Fig. 4. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A–F. Tibial spurs (indicated by the red arrows). A–B. Fore leg. C–D. Mid leg. E–F. Hind leg. G–H. Tarsal claw of mid leg. A, C, E, G. Photographs. B, D, F, H. Hand drawings. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3824593" httpUri="https://zenodo.org/record/3824593/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
A–F); tarsi slender, tarsal formula 5-5-4; tarsus nearly as long as tibiae in fore and mid legs, slightly shorter than tibiae in hind legs; tarsomeres cylindrical, hind tarsomere I slightly longer than tarsomeres II and III combined. Claws slightly pectinate (
<figureCitation id="F5C955ECFFA73F7DFC49E6877435FD11" box="[1005,1079,645,671]" captionStart="Fig" captionStartId="6.[189,232,1928,1954]" captionTargetBox="[194,1390,1008,1882]" captionTargetId="figure@6.[188,1398,998,1886]" captionTargetPageId="6" captionText="Fig. 4. Archaeoripiphorus nuwa Hsiao, Yu &amp; Deng gen. et sp. nov., holotype. A–F. Tibial spurs (indicated by the red arrows). A–B. Fore leg. C–D. Mid leg. E–F. Hind leg. G–H. Tarsal claw of mid leg. A, C, E, G. Photographs. B, D, F, H. Hand drawings. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3824593" httpUri="https://zenodo.org/record/3824593/files/figure.png" pageId="7" pageNumber="8">Figs 4</figureCitation>
G–H).
</paragraph>
</subSubSection>
</treatment>
</document>
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