170 lines
18 KiB
XML
170 lines
18 KiB
XML
<document id="E3BFAABD60E3BC4BA1CA822943E854CA" ID-ISSN="0003-0090" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630274675836" checkinUser="felipe" docAuthor="Dimitrov, Dimitar & Hormiga, Gustavo" docDate="2009" docId="A23B87F1FFC6FFD5E8DAB6CB7CDCF95A" docLanguage="en" docName="B317.pdf" docOrigin="Bulletin of the American Museum of Natural History 2009 (317)" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.3:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="3" docTitle="Cyrtognatha orphana Dimitrov & Hormiga 2009, new species" docType="treatment" docVersion="2" lastPageNumber="53" masterDocId="5E02FF89FFF2FFE1E81AB40C7F03FF85" masterDocTitle="Revision And Cladistic Analysis Of The Orbweaving Spider Genus Cyrtognatha Keyserling, 1881 (Araneae, Tetragnathidae)" masterLastPageNumber="140" masterPageNumber="1" pageNumber="53" updateTime="1699208653156" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="E54EC0E8FFA114DFFA18F2BB00DF2800" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="846A857FCD56195921E6A9112EB88270">
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<mods:title id="2900F0C094EE9A0B700846B09CBA8727">Revision And Cladistic Analysis Of The Orbweaving Spider Genus Cyrtognatha Keyserling, 1881 (Araneae, Tetragnathidae)</mods:title>
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<mods:name id="BF6F1878510EA3123BCA514ED449C102" type="personal">
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<mods:roleTerm id="0AA9217624EE8428BF62F23DA1CC1CDB">Author</mods:roleTerm>
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<mods:namePart id="07DCEDFF51E5494FB180F54E94944F96">Dimitrov, Dimitar</mods:namePart>
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</mods:name>
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<mods:name id="8D049363AD955CB389B18CFBC57C831E" type="personal">
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<mods:roleTerm id="C0EBC19A8E2E5FEC4A8E0D78BE01C8DE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="58FEDA74E93F636BD3959123A78F7087">Hormiga, Gustavo</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="D6D04BBF4B73900877BD7066C9490EAE">text</mods:typeOfResource>
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<mods:title id="F33597160B573E2F17F8D94DBFE4978D">Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="302DC29982208CC1E35E70A4E960BA41">
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<mods:date id="29702892F7C991B663D9E5468BC98329">2009</mods:date>
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<mods:detail id="9E7531EDEEAD3D657523A6AD784ABC0E" type="pubDate">
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<mods:number id="703AE0840C56741C762B31FAD67D652C">2009-03-17</mods:number>
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<mods:detail id="99F52DBD259EBDAC8D50ADB8D806D3D0" type="volume">
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<mods:number id="2001FDD121A19350DD1A40D6F4BFEAD0">2009</mods:number>
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<mods:detail id="37D32CF255B1E1831DC98B32EC509880" type="issue">
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<mods:number id="C8EDEFBC3BFBC16A81535E41BD5DD801">317</mods:number>
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<mods:classification id="968A5BC9EA678A0343178AEC670BF0E2">journal article</mods:classification>
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<mods:identifier id="039D3E8068F81B5BCF247289E3E846FD" type="ISSN">0003-0090</mods:identifier>
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</mods:mods>
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<treatment id="A23B87F1FFC6FFD5E8DAB6CB7CDCF95A" LSID="urn:lsid:plazi:treatment:A23B87F1FFC6FFD5E8DAB6CB7CDCF95A" httpUri="http://treatment.plazi.org/id/A23B87F1FFC6FFD5E8DAB6CB7CDCF95A" lastPageNumber="53" pageId="52" pageNumber="53">
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<subSubSection id="6288656CFFC6FFD5E8DAB6CB7ED9FD7E" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="2A2D36E7FFC6FFD5E8DAB6CB7ED9FD7E" blockId="52.[192,588,711,764]" pageId="52" pageNumber="53">
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<heading id="7165818BFFC6FFD5E8DAB6CB7ED9FD7E" centered="true" fontSize="9" level="2" pageId="52" pageNumber="53" reason="2">
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<taxonomicName id="ED924D64FFC6FFD5E8DAB6CB7EB6FD5B" ID-CoL="6BY9P" authority="Dimitrov & Hormiga, 2009" authorityName="Dimitrov & Hormiga" authorityYear="2009" box="[192,437,711,734]" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="orphana" status="sp. nov.">
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<emphasis id="18E6EAF5FFC6FFD5E8DAB6CB7EB6FD5B" bold="true" box="[192,437,711,734]" italics="true" pageId="52" pageNumber="53">Cyrtognatha orphana</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="03D5578EFFC6FFD5E9DCB6C47D4FFD5B" box="[454,588,712,734]" pageId="52" pageNumber="53" rank="species">new species</taxonomicNameLabel>
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<figureCitation id="B2A92A62FFC6FFD5E929B6EA7EB2FD7E" box="[307,433,742,764]" captionStart="Fig" captionStartId="53.[116,155,1352,1371]" captionTargetBox="[99,1184,210,1322]" captionTargetId="figure-134@53.[99,1188,209,1322]" captionTargetPageId="53" captionText="Fig. 22. Cyrtognatha orphana, n. sp. Male palp: (A), retrolateral (B), prolateral (C). Vulva: lateral (D), dorsal (E). Scale bars 0.5 mm." pageId="52" pageNumber="53">Figures 22</figureCitation>
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,
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<figureCitation id="B2A92A62FFC6FFD5E9A5B6EA7ED9FD7E" box="[447,474,742,763]" captionStart="Fig" captionStartId="54.[150,188,816,835]" captionTargetBox="[131,1224,211,786]" captionTargetId="figure-343@54.[131,1224,211,786]" captionTargetPageId="54" captionText="Fig. 23. Cyrtognatha orphana, n. sp. Male habitus: dorsal (A), lateral (C), ventral (D). Male chelicerae (B)." pageId="52" pageNumber="53">23</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6288656CFFC6FFD5E886B71E7D8DFBB0" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="2A2D36E7FFC6FFD5E886B71E7D8DFBB0" blockId="52.[128,654,786,1759]" pageId="52" pageNumber="53">
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TYPE: Male
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<typeStatus id="F5298845FFC6FFD5E958B7187EAAFCAF" box="[322,425,788,810]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
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from
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<collectingCountry id="52857677FFC6FFD5EA1EB7187D4FFCAF" box="[516,588,788,810]" name="Brazil" pageId="52" pageNumber="53">Brazil</collectingCountry>
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,
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<collectingRegion id="E856F805FFC6FFD5EA78B7197FC7FCCD" country="Brazil" name="Sao Paulo" pageId="52" pageNumber="53">São Paulo</collectingRegion>
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olov., Cuv-Cucho [
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<emphasis id="18E6EAF5FFC6FFD5E9B0B73F7ECBFCCD" box="[426,456,819,840]" italics="true" pageId="52" pageNumber="53">sic</emphasis>
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] (
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<collectionCode id="4C83AE22FFC6FFD5E9C5B73E7D3CFCC2" box="[479,575,818,839]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="52" pageNumber="53">MNHN</collectionCode>
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9192). The label does not provide collector or date information. The label in the vial had the handwritten name ‘‘
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<taxonomicName id="ED924D64FFC6FFD5E96CB7867D74FC25" box="[374,631,906,929]" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="fusciceps">
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<emphasis id="18E6EAF5FFC6FFD5E96CB7867D74FC25" box="[374,631,906,929]" italics="true" pageId="52" pageNumber="53">Cyrtognatha fusciceps</emphasis>
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</taxonomicName>
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’’. Most probably it was chosen by its original collector, but unfortunately he/she did not describe the species. The author of this name remains unknown, and thus we chose a different species name to describe this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="6288656CFFC6FFD5E886B0377E34FB62" pageId="52" pageNumber="53" type="etymology">
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<paragraph id="2A2D36E7FFC6FFD5E886B0377E34FB62" blockId="52.[128,654,786,1759]" pageId="52" pageNumber="53">
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ETYMOLOGY: The species epithet is an adjective taken from the Latin word for
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<emphasis id="18E6EAF5FFC6FFD5E89AB0747FCDFB0A" box="[128,206,1144,1167]" italics="true" pageId="52" pageNumber="53">orphan</emphasis>
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, in reference to the lost identity of the arachnologist that first realized that these specimens belonged to an undescribed species of
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<taxonomicName id="ED924D64FFC6FFD5E8BBB0DC7E33FB62" authorityName=", Keyserling" authorityYear="1881" box="[161,304,1232,1255]" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="genus">
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<emphasis id="18E6EAF5FFC6FFD5E8BBB0DC7E33FB62" box="[161,304,1232,1255]" italics="true" pageId="52" pageNumber="53">Cyrtognatha</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="6288656CFFC6FFD5E886B0E17EDBF95A" pageId="52" pageNumber="53" type="diagnosis">
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<paragraph id="2A2D36E7FFC6FFD5E886B0E17EDBF95A" blockId="52.[128,654,786,1759]" pageId="52" pageNumber="53">
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DIAGNOSIS: This species can be distinguished easily from all other
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<taxonomicName id="ED924D64FFC6FFD5E9E4B1007FCCFAC4" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="18E6EAF5FFC6FFD5E9E4B1007D8DFAA6" box="[510,654,1292,1315]" italics="true" pageId="52" pageNumber="53">Cyrtognatha</emphasis>
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species
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</taxonomicName>
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by the conspicuously prolaterally slanted embolus (fig. 22A). It differs significantly from similar congeners (namely,
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<taxonomicName id="ED924D64FFC6FFD5EA69B16A7FD1FA1F" authorityName="Dimitrov & Hormiga" authorityYear="2009" baseAuthorityName="Chickering" baseAuthorityYear="1956" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="insolita">
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<emphasis id="18E6EAF5FFC6FFD5EA69B16A7FD1FA1F" italics="true" pageId="52" pageNumber="53">C. insolita</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="ED924D64FFC6FFD5E915B1887E72FA1F" authorityName="Dimitrov & Hormiga" authorityYear="2009" box="[271,369,1411,1434]" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="petila">
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<emphasis id="18E6EAF5FFC6FFD5E915B1887E72FA1F" box="[271,369,1411,1434]" italics="true" pageId="52" pageNumber="53">C. petila</emphasis>
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</taxonomicName>
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) in that both
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<taxonomicName id="ED924D64FFC6FFD5EA02B1887D8DFA1F" authorityName="Dimitrov & Hormiga" authorityYear="2009" baseAuthorityName="Chickering" baseAuthorityYear="1956" box="[536,654,1411,1434]" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="insolita">
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<emphasis id="18E6EAF5FFC6FFD5EA02B1887D8DFA1F" box="[536,654,1411,1434]" italics="true" pageId="52" pageNumber="53">C. insolita</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="ED924D64FFC6FFD5E8A7B1AD7E2AFA32" authorityName="Dimitrov & Hormiga" authorityYear="2009" box="[189,297,1440,1463]" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="petila">
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<emphasis id="18E6EAF5FFC6FFD5E8A7B1AD7E2AFA32" box="[189,297,1440,1463]" italics="true" pageId="52" pageNumber="53">C. petila</emphasis>
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</taxonomicName>
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have the embolus with an extensive membrane that carries small outgrowths (serrated membrane) and have a triangular metine embolic apophysis (fig. 22A, C). The basal apophysis of the conductor in
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<taxonomicName id="ED924D64FFC6FFD5E8F8B2397E6CF9CE" authorityName="Dimitrov & Hormiga" authorityYear="2009" box="[226,367,1588,1611]" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="orphana">
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<emphasis id="18E6EAF5FFC6FFD5E8F8B2397E6CF9CE" box="[226,367,1588,1611]" italics="true" pageId="52" pageNumber="53">C. orphana</emphasis>
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</taxonomicName>
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has a well-developed membranous lamina (fig. 22C), while in
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<taxonomicName id="ED924D64FFC6FFD5EA69B25F7FD1F902" authorityName="Dimitrov & Hormiga" authorityYear="2009" baseAuthorityName="Chickering" baseAuthorityYear="1956" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="insolita">
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<emphasis id="18E6EAF5FFC6FFD5EA69B25F7FD1F902" italics="true" pageId="52" pageNumber="53">C. insolita</emphasis>
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</taxonomicName>
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it has a sharpened tip (fig. 37C). In
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<taxonomicName id="ED924D64FFC6FFD5EA69B27D7FBCF921" authorityName="Dimitrov & Hormiga" authorityYear="2009" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="petila">
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<emphasis id="18E6EAF5FFC6FFD5EA69B27D7FBCF921" italics="true" pageId="52" pageNumber="53">C. petila</emphasis>
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</taxonomicName>
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the membrane of the basal apophysis of the conductor is much smaller and the tip is sharpened as in
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<taxonomicName id="ED924D64FFC6FFD5E941B2C57ED2F95A" authorityName="Dimitrov & Hormiga" authorityYear="2009" baseAuthorityName="Chickering" baseAuthorityYear="1956" box="[347,465,1736,1759]" class="Arachnida" family="Tetragnathidae" genus="Cyrtognatha" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="insolita">
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<emphasis id="18E6EAF5FFC6FFD5E941B2C57ED2F95A" box="[347,465,1736,1759]" italics="true" pageId="52" pageNumber="53">C. insolita</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="6288656CFFC6FFD5EAC0B4DF7B51F9CF" pageId="52" pageNumber="53" type="description">
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<paragraph id="2A2D36E7FFC6FFD5EAC0B4DF7B88FBDD" blockId="52.[701,1228,211,1759]" pageId="52" pageNumber="53">
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DESCRIPTION:
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<emphasis id="18E6EAF5FFC6FFD5EB92B4D87CC7FF6E" box="[904,964,212,235]" italics="true" pageId="52" pageNumber="53">Male</emphasis>
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(
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<emphasis id="18E6EAF5FFC6FFD5EBCFB4D87B36FF6E" box="[981,1077,212,235]" italics="true" pageId="52" pageNumber="53">
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<typeStatus id="F5298845FFC6FFD5EBCFB4D87B36FF6E" box="[981,1077,212,235]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
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</emphasis>
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). Habitus as in figure 23A–D. Carapace yellowish with brownish cephalic region. Fovea well marked with darker pigmentation. Abdomen cylindrical with small proximal tubercle. Distal tubercle short and extending over the spinnerets. Abdomen yellowish with numerous guanine spots concentrated dorsolaterally; dorsally and ventrally the guanine patches are absent around its longitudinal axis. Legs yellowish with remains of greenish coloration. Total length 6.02. Cephalothorax 2.32 long, 1.50 wide, 1.00 high. Abdomen 3.70 long, 0.85 wide, 1.25 high. Sternum yellowish; 0.88 long, 0.78 wide. Endites darker with brown color. Clypeus height 1.3 times one AME diameter. AME larger than the rest of the eyes. ALE and PLE close to each other on short elevations. AME diameter 0.15. Distance between AME slightly more than their diameter; AME–ALE distance 1.5 times one AME diameter. AME–PME distance at about one AME diameter. Distance between PME equal to their diameter. Chelicerae brown, very robust, and strongly divergent (fig. 23B). Cheliceral dorsal tooth very well developed, close to the fang articulation and conspicuously curved prolaterally. Femur I 5.14, 2.21 times the length of the cephalothorax. Pedipalp as in figure 22A–C. Palpal tibia length 0.25; cymbium length 0.95.
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</paragraph>
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<paragraph id="2A2D36E7FFC6FFD5EAC1B06C7B51F9CF" blockId="52.[701,1228,211,1759]" pageId="52" pageNumber="53">
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<emphasis id="18E6EAF5FFC6FFD5EAC1B06C7C2EFBF2" box="[731,813,1120,1143]" italics="true" pageId="52" pageNumber="53">Female</emphasis>
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(
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<emphasis id="18E6EAF5FFC6FFD5EB25B06D7CA1FBF3" box="[831,930,1121,1142]" italics="true" pageId="52" pageNumber="53">
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<typeStatus id="F5298845FFC6FFD5EB25B06D7CA1FBF3" box="[831,930,1121,1142]" pageId="52" pageNumber="53" type="paratype">paratype</typeStatus>
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</emphasis>
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). Habitus and pigmentation pattern as in male. Carapace, chelicerae, and endites with lighter color than in male. Legs as in male with some remains of greenish coloring. Total length 6.41. Cephalothorax 2.25 long, 1.50 wide, 1.00 high. Abdomen 4.16 long, 1.22 wide, 1.22 high. Sternum 0.87 long, 0.87 wide. Chelicerae not divergent and without dorsal tooth; fang without outgrowth. Eyes as in male but AME closer to each other and to ALE. Distance between AME less than their diameter and between them and ALE equal to their diameter. AME diameter 0.12. Clypeus height 1.25 times one AME diameter. Femur I 4.65, 2.06 times the length of the cephalothorax. Vulva as in figure 22D, E.
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</paragraph>
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</subSubSection>
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<subSubSection id="6288656CFFC6FFD5EAC0B25D7CDCF95A" pageId="52" pageNumber="53" type="distribution">
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<paragraph id="2A2D36E7FFC6FFD5EAC0B25D7B18F903" blockId="52.[701,1228,211,1759]" pageId="52" pageNumber="53">
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DISTRIBUTION: This species is known only from the
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<typeStatus id="F5298845FFC6FFD5EB37B27D7C5CF903" box="[813,863,1649,1670]" pageId="52" pageNumber="53">type</typeStatus>
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locality (fig. 5).
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</paragraph>
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<paragraph id="2A2D36E7FFC6FFD5EAC0B2817CDCF95A" blockId="52.[701,1228,211,1759]" pageId="52" pageNumber="53">
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ADDITIONAL MATERIAL EXAMINED:
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<emphasis id="18E6EAF5FFC6FFD5EC90B2837DF4F947" italics="true" pageId="52" pageNumber="53">
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<typeStatus id="F5298845FFC6FFD5EC90B2837DF3F947" pageId="52" pageNumber="53" type="paratype">Paratype</typeStatus>
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:
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</emphasis>
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<collectingCountry id="52857677FFC6FFD5EAE4B2A07C6EF944" box="[766,877,1708,1729]" name="Brazil" pageId="52" pageNumber="53">BRAZIL</collectingCountry>
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:
|
||
<collectingRegion id="E856F805FFC6FFD5EB62B2A07CEDF947" box="[888,1006,1708,1730]" country="Brazil" name="Sao Paulo" pageId="52" pageNumber="53">São Paulo</collectingRegion>
|
||
olov., Cuv-Cucho,
|
||
<specimenCount id="3C94FD6EFFC6FFD5EAA7B2C57C1CF95A" box="[701,799,1737,1759]" pageId="52" pageNumber="53" type="female">1 female</specimenCount>
|
||
(MNHN 9192).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |