352 lines
48 KiB
XML
352 lines
48 KiB
XML
<document id="3288F2B60EA3CB6CEA2050F90A9E2480" ID-DOI="10.5281/zenodo.4650779" ID-ISSN="1638-9395" ID-Zenodo-Dep="4650779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617194601297" checkinUser="felipe" docAuthor="Kostopoulos, Dimitris S." docDate="2005" docId="9636CE11B46BFF8FFD30D988FEDEFE3E" docLanguage="en" docName="Geodiversitas.27.4.747-791.pdf" docOrigin="Geodiversitas 27 (4)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Palaeoryx majori Schlosser 1904" docType="treatment" docVersion="3" lastPageNumber="780" masterDocId="6A0FB669B474FFADFFCADD44FF88FFDB" masterDocTitle="The Bovidae (Mammalia, Artiodactyla) from the late Miocene of Akkaşdağı, Turkey" masterLastPageNumber="791" masterPageNumber="747" pageNumber="778" updateTime="1698897313800" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="E99BE0ECF2F3F220E1C9F76A3E611CE8">The Bovidae (Mammalia, Artiodactyla) from the late Miocene of Akkaşdağı, Turkey</mods:title>
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<mods:name id="0DD11BB4548018A494910B4838EB7453" type="personal">
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<mods:namePart id="E4E7A4C9CC84141AD672FBFDC2C7244C">Kostopoulos, Dimitris S.</mods:namePart>
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<mods:affiliation id="C85E974514324669BF591762F602443E">University of Thessaloniki, Museum of the Geological Department, 54124 Thessaloniki (Greece) dkostop @ geo. auth. gr</mods:affiliation>
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<mods:title id="853192EF5FBDF42635B0693B565A013F">Geodiversitas</mods:title>
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<mods:date id="7B1BFB7DCC590220F0386C1FEF440589">2005</mods:date>
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<mods:number id="782036E6FCD2D849A7F5226478BA8B33">27</mods:number>
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<treatment id="9636CE11B46BFF8FFD30D988FEDEFE3E" LSID="urn:lsid:plazi:treatment:9636CE11B46BFF8FFD30D988FEDEFE3E" httpUri="http://treatment.plazi.org/id/9636CE11B46BFF8FFD30D988FEDEFE3E" lastPageId="34" lastPageNumber="780" pageId="31" pageNumber="778">
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<subSubSection id="56852C8CB46BFFB2FD30D988FBEBFB3E" box="[762,1123,1228,1253]" pageId="31" pageNumber="778" type="nomenclature">
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<paragraph id="1E207F07B46BFFB2FD30D988FBEBFB3E" blockId="31.[762,1123,1228,1253]" box="[762,1123,1228,1253]" pageId="31" pageNumber="778">
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<heading id="4568C86BB46BFFB2FD30D988FBEBFB3E" box="[762,1123,1228,1253]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="778" reason="2">
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<taxonomicName id="D99F0484B46BFFB2FD30D988FBEBFB3E" authority="Schlosser, 1904" authorityName="Schlosser" authorityYear="1904" box="[762,1123,1228,1253]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="31" pageNumber="778" phylum="Chordata" rank="species" species="majori">
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<emphasis id="2CEBA315B46BFFB2FD30D988FC3CFB3E" bold="true" box="[762,948,1228,1253]" italics="true" pageId="31" pageNumber="778">Palaeoryx majori</emphasis>
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<bibRefCitation id="7A0E02F6B46BFFB2FC71D988FBEBFB3E" author="SCHLOSSER M." box="[955,1123,1228,1253]" pageId="31" pageNumber="778" pagination="21 - 118" refId="ref23722" refString="SCHLOSSER M. 1904. - Die fossiles Cavicornis von Samos. Beitrage zur Palaontologie und Geologie Osterreich-Ungarn 17: 21 - 118." type="journal article" year="1904">Schlosser, 1904</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="56852C8CB46BFFB2FD6ED842FB31FA9E" pageId="31" pageNumber="778" type="reference_group">
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<paragraph id="1E207F07B46BFFB2FD6ED842FB14FAC6" blockId="31.[676,1209,1286,1466]" box="[676,1180,1286,1309]" pageId="31" pageNumber="778">
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<treatmentCitationGroup id="3E8F5829B46BFFB2FD6ED842FB14FAC6" box="[676,1180,1286,1309]" pageId="31" pageNumber="778">
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<treatmentCitation id="9F3E5916B46BFFB2FD6ED842FB8FFAC6" author="SCHLOSSER M." box="[676,1031,1286,1309]" page="38" pageId="31" pageNumber="778" year="1904">
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<taxonomicName id="D99F0484B46BFFB2FD6ED842FB8FFAC6" authority="Schlosser, 1904: 38" authorityName="Schlosser" authorityPageNumber="38" authorityYear="1904" box="[676,1031,1286,1309]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="31" pageNumber="778" phylum="Chordata" rank="species" species="majori">
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<emphasis id="2CEBA315B46BFFB2FD6ED842FCCBFAC6" box="[676,835,1286,1309]" italics="true" pageId="31" pageNumber="778">Palaeoryx majori</emphasis>
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<bibRefCitation id="7A0E02F6B46BFFB2FC83D842FB8FFAC6" author="SCHLOSSER M." box="[841,1031,1286,1309]" pageId="31" pageNumber="778" pagination="21 - 118" refId="ref23722" refString="SCHLOSSER M. 1904. - Die fossiles Cavicornis von Samos. Beitrage zur Palaontologie und Geologie Osterreich-Ungarn 17: 21 - 118." type="journal article" year="1904">Schlosser, 1904: 38</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, pl. VII, fig. 5.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="1E207F07B46BFFB2FD6ED86AFB31FA9E" blockId="31.[676,1209,1286,1466]" box="[676,1209,1326,1349]" pageId="31" pageNumber="778">
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<treatmentCitationGroup id="3E8F5829B46BFFB2FD6ED86AFB31FA9E" box="[676,1209,1326,1349]" pageId="31" pageNumber="778">
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<treatmentCitation id="9F3E5916B46BFFB2FD6ED86AFB8CFA9E" author="ANDREE J." box="[676,1028,1326,1349]" page="161" pageId="31" pageNumber="778" year="1926">
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<taxonomicName id="D99F0484B46BFFB2FD6ED86AFB8CFA9E" authority="Andree, 1926: 161" authorityName="Andree" authorityPageNumber="161" authorityYear="1926" box="[676,1028,1326,1349]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="31" pageNumber="778" phylum="Chordata" rank="species" species="laticeps">
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<emphasis id="2CEBA315B46BFFB2FD6ED86AFCCFFA9E" box="[676,839,1326,1349]" italics="true" pageId="31" pageNumber="778">Palaeoryx laticeps</emphasis>
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<bibRefCitation id="7A0E02F6B46BFFB2FC87D86AFB8CFA9E" author="ANDREE J." box="[845,1028,1326,1349]" pageId="31" pageNumber="778" pagination="135 - 175" refId="ref21498" refString="ANDREE J. 1926. - Neue Cavicornier aus dem Pliocan von Samos. Palaeontographica 67 (6): 135 - 175." type="journal article" year="1926">Andree, 1926: 161</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, pl. XIII, figs 4, 6.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="56852C8CB46BFFB2FD6ED813FC7DFA61" pageId="31" pageNumber="778" type="materials_examined">
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<paragraph id="1E207F07B46BFFB2FD6ED813FC7DFA61" blockId="31.[676,1209,1286,1466]" pageId="31" pageNumber="778">MATERIAL EXAMINED. — Frontlet: AK3-130; P3-M3: AK5-643 (LM1-M3 = 60.7); P4-M3: AK5-426 (LM1-M3 = 62.0); P3-P4: AK4-191; P3: AK5-609; M2: AK5-369; m2-m3: AK4-199.</paragraph>
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</subSubSection>
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<subSubSection id="56852C8CB46BFF8FFD6ED8A8FEDEFE3E" lastPageId="34" lastPageNumber="781" pageId="31" pageNumber="778" type="description">
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<paragraph id="1E207F07B46BFFB2FD6ED8A8FCBDF9DF" blockId="31.[676,1209,1516,1701]" box="[676,821,1516,1541]" pageId="31" pageNumber="778">DESCRIPTION</paragraph>
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||
<paragraph id="1E207F07B46BFFB2FD6EDB48FD51F9FE" blockId="31.[676,1209,1516,1701]" box="[676,729,1548,1573]" pageId="31" pageNumber="778">
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<emphasis id="2CEBA315B46BFFB2FD6EDB48FD51F9FE" box="[676,729,1548,1573]" italics="true" pageId="31" pageNumber="778">Skull</emphasis>
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</paragraph>
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<paragraph id="1E207F07B46BFF8DFD6EDB68FC9DFAFE" blockId="31.[676,1209,1516,1701]" lastBlockId="32.[676,1209,1036,1317]" lastPageId="32" lastPageNumber="779" pageId="31" pageNumber="778">
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The braincase is wide (Appendix:
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<tableCitation id="531D4ABCB46BFFB2FBD6DB68FBF7F99E" box="[1052,1151,1580,1605]" captionStart="TABLE" captionStartId="42.[113,124,918,935]" captionText="TABLE 10. — Skull measurements (in mm) of Miotragocerus valenciennesi (Gaudry, 1861), Pachytragus crassicornis Schlosser, 1904 and Palaeoryx majori Schlosser, 1904 from Akkaşdağı. Abbreviations: DAP, anteroposterior diameter; DT, transverse diameter." pageId="31" pageNumber="778">Table 10</tableCitation>
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||
) and the cranial roof slightly convex (
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||
<figureCitation id="86A46382B46BFFB2FBDBDB08FBFAF9BE" box="[1041,1138,1612,1637]" captionStart="FIG" captionStartId="31.[165,176,1076,1093]" captionTargetBox="[113,1202,203,1039]" captionTargetId="figure-907@31.[113,1202,203,1039]" captionTargetPageId="31" captionText="FIG. 23. — Palaeoryx majori Schlosser, 1904 from Akkaşdağı, skull AK3-130; A, lateral view; B, dorsal view. Scale 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650859" httpUri="https://zenodo.org/record/4650859/files/figure.png" pageId="31" pageNumber="778">Fig. 23A</figureCitation>
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||
). The opisthocranium is short and moderately low. The mid-frontal suture is raised slightly and is rather open (
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<figureCitation id="86A46382B454FF8DFF76D948FE97FBFE" box="[188,287,1036,1061]" captionStart="FIG" captionStartId="31.[165,176,1076,1093]" captionTargetBox="[113,1202,203,1039]" captionTargetId="figure-907@31.[113,1202,203,1039]" captionTargetPageId="31" captionText="FIG. 23. — Palaeoryx majori Schlosser, 1904 from Akkaşdağı, skull AK3-130; A, lateral view; B, dorsal view. Scale 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650859" httpUri="https://zenodo.org/record/4650859/files/figure.png" pageId="32" pageNumber="779">Fig. 23B</figureCitation>
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||
). The frontals are not elevated between the horn-cores. Ahead of them the frontals incline smoothly, forming an angle of about 105° with the cranial roof. The orbits are rather round- ed and well projected laterally. The width of the skull at the dorsal orbital rims accounts
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<quantity id="D967D2E2B454FF8DFDE8D9E8FD0BFB1E" box="[546,643,1196,1221]" metricMagnitude="-1" metricUnit="m" metricValue="1.55" pageId="32" pageNumber="779" unit="mm" value="155.0">155 mm</quantity>
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. The supraorbital foramens are sunken into elongated and narrow pits. There are no postcornual grooves. The anterior tuberosities of the preserved basioccipital are strong, widely separated and slightly oblique comparatively to the sagittal plane. Between them and the posterior tuberosities a wide furrow runs along the basioccipital axis. The oval foramen is large, placed well in front of the anterior tuberosities. The horn-cores are elongated (≈
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<quantity id="D967D2E2B454FF8DFEE0D8A8FE01F9DE" box="[298,393,1516,1541]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="779" unit="mm" value="300.0">300 mm</quantity>
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||
along the anterior surface) and placed just above the orbits and laterally (
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||
<figureCitation id="86A46382B454FF8DFFB0DB68FF72F99E" box="[122,250,1580,1605]" captionStart="FIG" captionStartId="31.[165,176,1076,1093]" captionTargetBox="[113,1202,203,1039]" captionTargetId="figure-907@31.[113,1202,203,1039]" captionTargetPageId="31" captionText="FIG. 23. — Palaeoryx majori Schlosser, 1904 from Akkaşdağı, skull AK3-130; A, lateral view; B, dorsal view. Scale 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650859" httpUri="https://zenodo.org/record/4650859/files/figure.png" pageId="32" pageNumber="779">Fig. 23A, B</figureCitation>
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). Their basal part forms an angle of about 80° with the cranial roof. They are placed well apart at the base (internal basal distance at about
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<quantity id="D967D2E2B454FF8DFF73DBC8FE84F97E" box="[185,268,1676,1701]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="32" pageNumber="779" unit="mm" value="57.0">57 mm</quantity>
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||
) and widely divergent toward the apexes. In their uppermost part they re-curve inwards (
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<figureCitation id="86A46382B454FF8DFCD8D968FCEDFB9E" box="[786,869,1068,1093]" captionStart="FIG" captionStartId="31.[165,176,1076,1093]" captionTargetBox="[113,1202,203,1039]" captionTargetId="figure-907@31.[113,1202,203,1039]" captionTargetPageId="31" captionText="FIG. 23. — Palaeoryx majori Schlosser, 1904 from Akkaşdağı, skull AK3-130; A, lateral view; B, dorsal view. Scale 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650859" httpUri="https://zenodo.org/record/4650859/files/figure.png" pageId="32" pageNumber="779">Fig. 23</figureCitation>
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||
). In lateral view they appear strongly inclined backwards and curved posteriorly (
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<figureCitation id="86A46382B454FF8DFD3BD928FCD9FB5E" box="[753,849,1132,1157]" captionStart="FIG" captionStartId="31.[165,176,1076,1093]" captionTargetBox="[113,1202,203,1039]" captionTargetId="figure-907@31.[113,1202,203,1039]" captionTargetPageId="31" captionText="FIG. 23. — Palaeoryx majori Schlosser, 1904 from Akkaşdağı, skull AK3-130; A, lateral view; B, dorsal view. Scale 5 cm." figureDoi="http://doi.org/10.5281/zenodo.4650859" httpUri="https://zenodo.org/record/4650859/files/figure.png" pageId="32" pageNumber="779">Fig. 23A</figureCitation>
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). Thin discontinuous longitudinal grooves run along their surface. They do not bear any keel. Their cross-section is elliptical with weak mediolateral compression at the base, becoming stronger towards the tips (Appendix:
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<tableCitation id="531D4ABCB454FF8DFD6ED848FC8DFAFE" box="[676,773,1292,1317]" captionStart="TABLE" captionStartId="42.[113,124,918,935]" captionText="TABLE 10. — Skull measurements (in mm) of Miotragocerus valenciennesi (Gaudry, 1861), Pachytragus crassicornis Schlosser, 1904 and Palaeoryx majori Schlosser, 1904 from Akkaşdağı. Abbreviations: DAP, anteroposterior diameter; DT, transverse diameter." pageId="32" pageNumber="779">Table 10</tableCitation>
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||
).
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||
</paragraph>
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||
<caption id="4AE02F8FB454FF8DFFBBDED6FF53FC15" ID-DOI="http://doi.org/10.5281/zenodo.4650863" ID-Zenodo-Dep="4650863" httpUri="https://zenodo.org/record/4650863/files/figure.png" pageId="32" pageNumber="779" startId="32.[113,124,914,931]" targetBox="[113,1201,203,878]" targetPageId="32">
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||
<paragraph id="1E207F07B454FF8DFFBBDED6FF53FC15" blockId="32.[113,1209,914,974]" pageId="32" pageNumber="779">
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||
FIG. 24. — Drawings of the skull specimens of
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||
<taxonomicName id="D99F0484B454FF8DFE33DED6FC98FC78" authority="Schlosser, 1904" authorityName="Schlosser" authorityYear="1904" box="[505,784,914,931]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="32" pageNumber="779" phylum="Chordata" rank="species" species="majori">
|
||
<emphasis id="2CEBA315B454FF8DFE33DED6FD0BFC78" box="[505,643,914,931]" italics="true" pageId="32" pageNumber="779">Palaeoryx majori</emphasis>
|
||
<bibRefCitation id="7A0E02F6B454FF8DFD42DED6FC98FC78" author="SCHLOSSER M." box="[648,784,914,931]" pageId="32" pageNumber="779" pagination="21 - 118" refId="ref23722" refString="SCHLOSSER M. 1904. - Die fossiles Cavicornis von Samos. Beitrage zur Palaontologie und Geologie Osterreich-Ungarn 17: 21 - 118." type="journal article" year="1904">Schlosser, 1904</bibRefCitation>
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||
</taxonomicName>
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||
(
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||
<emphasis id="2CEBA315B454FF8DFCD0DED6FCAFFC78" bold="true" box="[794,807,914,931]" pageId="32" pageNumber="779">A</emphasis>
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||
,
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||
<emphasis id="2CEBA315B454FF8DFCF8DED6FCB7FC78" bold="true" box="[818,831,914,931]" pageId="32" pageNumber="779">B</emphasis>
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||
) and
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||
<taxonomicName id="D99F0484B454FF8DFCA5DED6FBF6FC78" authority="(Wagner, 1857)" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[879,1150,914,931]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="32" pageNumber="779" phylum="Chordata" rank="species" species="pallasi">
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||
<emphasis id="2CEBA315B454FF8DFCA5DED6FC71FC78" box="[879,1017,914,931]" italics="true" pageId="32" pageNumber="779">Palaeoryx pallasi</emphasis>
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||
(Wagner, 1857)
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||
</taxonomicName>
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||
(
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<emphasis id="2CEBA315B454FF8DFB42DED6FB1EFC78" bold="true" box="[1160,1174,914,931]" pageId="32" pageNumber="779">C</emphasis>
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||
,
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||
<emphasis id="2CEBA315B454FF8DFB6BDED6FB27FC78" bold="true" box="[1185,1199,914,931]" pageId="32" pageNumber="779">D</emphasis>
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||
);
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<emphasis id="2CEBA315B454FF8DFFBBDEE3FFF6FC63" bold="true" box="[113,126,935,952]" pageId="32" pageNumber="779">A</emphasis>
|
||
, MTLB-160 (AeMNH);
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<emphasis id="2CEBA315B454FF8DFE8BDEE3FEC6FC63" bold="true" box="[321,334,935,952]" pageId="32" pageNumber="779">B</emphasis>
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||
, PIM-121;
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<emphasis id="2CEBA315B454FF8DFE64DEE3FE34FC63" bold="true" box="[430,444,935,952]" pageId="32" pageNumber="779">C</emphasis>
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||
, MTLA-113 (AeMNH);
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<emphasis id="2CEBA315B454FF8DFDB4DEE3FD04FC63" bold="true" box="[638,652,935,952]" pageId="32" pageNumber="779">D</emphasis>
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, M10831 (BMNH) from Samos and Pikermi (approximately at the same scale).
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</paragraph>
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</caption>
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<paragraph id="1E207F07B454FF8DFD6ED808FC81FABE" blockId="32.[676,1210,1356,1701]" box="[676,777,1356,1381]" pageId="32" pageNumber="779">
|
||
<emphasis id="2CEBA315B454FF8DFD6ED808FC81FABE" box="[676,777,1356,1381]" italics="true" pageId="32" pageNumber="779">Dentition</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1E207F07B454FF8CFD6ED828FF15FE9E" blockId="32.[676,1210,1356,1701]" lastBlockId="33.[113,646,204,325]" lastPageId="33" lastPageNumber="780" pageId="32" pageNumber="779">
|
||
The few dental specimens assigned to this form, cannot give a complete idea on the tooth morphology. They differ from the previously described dentition of
|
||
<emphasis id="2CEBA315B454FF8DFCBFD888FB03FA3E" box="[885,1163,1484,1509]" italics="true" pageId="32" pageNumber="779">
|
||
Miotragoceus
|
||
<taxonomicName id="D99F0484B454FF8DFBCED888FB03FA3E" baseAuthorityName="Moya-Sola" baseAuthorityYear="1983" box="[1028,1163,1484,1509]" class="Mammalia" family="Bovidae" genus="Miotragocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="32" pageNumber="779" phylum="Chordata" rank="species" species="valenciennesi">valenciennesi</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="D99F0484B454FF8DFD6ED8A8FC3CF9DE" baseAuthorityName="Roth F Wagner" baseAuthorityYear="1854" box="[676,948,1516,1541]" class="Mammalia" family="Bovidae" genus="Tragoportax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="32" pageNumber="779" phylum="Chordata" rank="species" species="amalthea">
|
||
<emphasis id="2CEBA315B454FF8DFD6ED8A8FCABF9DE" box="[676,803,1516,1541]" italics="true" pageId="32" pageNumber="779">Tragoportax</emphasis>
|
||
aff.
|
||
<emphasis id="2CEBA315B454FF8DFC9ED8A8FC3CF9DE" box="[852,948,1516,1541]" italics="true" pageId="32" pageNumber="779">amalthea</emphasis>
|
||
</taxonomicName>
|
||
in the longer molar row and the stronger hypsodonty. The P3 is lingually bi-lobed, while both the P3, 4 bear a hypoconal spur. The upper molars are wide with well developed basal pillar, which in some specimens (mainly in M2) appears double. The lobes fuse slowly together, giving rise to a strong central islet. The m3 has a clear goat fold, a strong “ectostylid” and a well developed-rounded talonid.
|
||
</paragraph>
|
||
<paragraph id="1E207F07B455FF8CFFBBDC28FE88FE5F" blockId="33.[113,646,364,1701]" box="[113,256,364,389]" pageId="33" pageNumber="780">COMPARISON</paragraph>
|
||
<paragraph id="1E207F07B455FF8CFFBBDCC8FF46FCBE" blockId="33.[113,646,364,1701]" pageId="33" pageNumber="780">
|
||
The morphological characters of the skull AK3- 130 leave no doubts about its attribution to the genus
|
||
<taxonomicName id="D99F0484B455FF8CFF72DC88FE49FE3E" authority="Gaudry, 1861" authorityName="Gaudry" authorityYear="1861" box="[184,449,460,485]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="genus">
|
||
<emphasis id="2CEBA315B455FF8CFF72DC88FE94FE3E" box="[184,284,460,485]" italics="true" pageId="33" pageNumber="780">Palaeoryx</emphasis>
|
||
<bibRefCitation id="7A0E02F6B455FF8CFEECDC88FE49FE3E" author="GAUDRY A." box="[294,449,460,485]" pageId="33" pageNumber="780" pagination="388 - 400" refId="ref22113" refString="GAUDRY A. 1861. - Notes sur les antilopes trouvees a Pikermi (Grece). Bulletin de la Societe geologique de France 18: 388 - 400." type="journal article" year="1861">Gaudry, 1861</bibRefCitation>
|
||
</taxonomicName>
|
||
, according to the diagnoses given by Pilgrim F Hopwood (1928) and
|
||
<bibRefCitation id="7A0E02F6B455FF8CFF69DF48FEC0FDFE" author="GENTRY A. W." box="[163,328,524,549]" pageId="33" pageNumber="780" pagination="229 - 296" refId="ref22251" refString="GENTRY A. W. 1971. - The earliest goats and other antelopes from Samos Hipparion Fauna. Bulletin of the British Museum (Natural History), Geology 20: 229 - 296." type="journal article" year="1971">Gentry (1971)</bibRefCitation>
|
||
. Although early authors dis- tinguished several species into this genus (
|
||
<bibRefCitation id="7A0E02F6B455FF8CFFB0DF08FE97FDBE" author="SCHLOSSER M." box="[122,287,588,613]" pageId="33" pageNumber="780" pagination="21 - 118" refId="ref23722" refString="SCHLOSSER M. 1904. - Die fossiles Cavicornis von Samos. Beitrage zur Palaontologie und Geologie Osterreich-Ungarn 17: 21 - 118." type="journal article" year="1904">Schlosser 1904</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7A0E02F6B455FF8CFEE5DF08FE4AFDBE" author="ANDREE J." box="[303,450,588,613]" pageId="33" pageNumber="780" pagination="135 - 175" refId="ref21498" refString="ANDREE J. 1926. - Neue Cavicornier aus dem Pliocan von Samos. Palaeontographica 67 (6): 135 - 175." type="journal article" year="1926">Andree 1926</bibRefCitation>
|
||
; summarized and reviewed by Pilgrim F Hopwood 1928), later authors simplify its taxonomy and recognize a single species
|
||
<taxonomicName id="D99F0484B455FF8CFEDADFE8FD0EFD1E" authority="(Wagner, 1857)" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[272,646,684,709]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFEDADFE8FE4EFD1E" box="[272,454,684,709]" italics="true" pageId="33" pageNumber="780">Palaeoryx pallasi</emphasis>
|
||
(Wagner, 1857)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7A0E02F6B455FF8CFFB0DF88FE8CFD3E" author="GENTRY A. W." box="[122,260,716,741]" pageId="33" pageNumber="780" pagination="229 - 296" refId="ref22251" refString="GENTRY A. W. 1971. - The earliest goats and other antelopes from Samos Hipparion Fauna. Bulletin of the British Museum (Natural History), Geology 20: 229 - 296." type="journal article" year="1971">Gentry 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="7A0E02F6B455FF8CFED9DF88FE30FD3E" author="SOLOUNIAS N." box="[275,440,716,741]" pageId="33" pageNumber="780" pagination="1 - 232" refId="ref23812" refString="SOLOUNIAS N. 1981. - The Turolian fauna from the island of Samos, Greece. Contribution on Vertebrate Evolution 6: 1 - 232." type="journal article" year="1981">Solounias 1981</bibRefCitation>
|
||
; Bosscha-Erdbrink 1988).
|
||
<bibRefCitation id="7A0E02F6B455FF8CFF08DFA8FEF6FCDE" author="SOLOUNIAS N." box="[194,382,748,773]" pageId="33" pageNumber="780" pagination="1 - 232" refId="ref23812" refString="SOLOUNIAS N. 1981. - The Turolian fauna from the island of Samos, Greece. Contribution on Vertebrate Evolution 6: 1 - 232." type="journal article" year="1981">Solounias (1981)</bibRefCitation>
|
||
accepts the monospecificity of the genus, but he also recognizes four varieties based on some dental and horn-core differences.
|
||
</paragraph>
|
||
<paragraph id="1E207F07B455FF8CFFBBDE28FEDCFABE" blockId="33.[113,646,364,1701]" pageId="33" pageNumber="780">
|
||
My observations on the
|
||
<taxonomicName id="D99F0484B455FF8CFE4BDE28FE6EFC5E" authorityName="Gaudry" authorityYear="1861" box="[385,486,876,901]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="genus">
|
||
<emphasis id="2CEBA315B455FF8CFE4BDE28FE6EFC5E" box="[385,486,876,901]" italics="true" pageId="33" pageNumber="780">Palaeoryx</emphasis>
|
||
</taxonomicName>
|
||
material from Pikermi (BMNH, MNHN) and Samos (MGL, BMNH, AMNH, PIM, AeMNH) allow a slightly different approach, recognizing
|
||
<taxonomicName id="D99F0484B455FF8CFDDDDE88FEEFFBDE" authority="Schlosser, 1904" authorityName="Schlosser" authorityYear="1904" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="majori">
|
||
<emphasis id="2CEBA315B455FF8CFDDDDE88FF3EFBDE" italics="true" pageId="33" pageNumber="780">Palaeoryx majori</emphasis>
|
||
<bibRefCitation id="7A0E02F6B455FF8CFF74DEA8FEEFFBDE" author="SCHLOSSER M." box="[190,359,1004,1029]" pageId="33" pageNumber="780" pagination="21 - 118" refId="ref23722" refString="SCHLOSSER M. 1904. - Die fossiles Cavicornis von Samos. Beitrage zur Palaontologie und Geologie Osterreich-Ungarn 17: 21 - 118." type="journal article" year="1904">Schlosser, 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
as a valid species. Indeed, the differences between
|
||
<taxonomicName id="D99F0484B455FF8CFE79D948FD0EFBFE" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[435,646,1036,1061]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFE79D948FD0EFBFE" box="[435,646,1036,1061]" italics="true" pageId="33" pageNumber="780">Palaeoryx pallasi</emphasis>
|
||
</taxonomicName>
|
||
(W a g n e r, 1 8 5 7) a n d
|
||
<emphasis id="2CEBA315B455FF8CFE4AD968FD0EFB9E" box="[384,646,1068,1093]" italics="true" pageId="33" pageNumber="780">P a l a e o r y x w o o d w a r d i</emphasis>
|
||
Pilgrim F Hopwood, 1928 do not merit a species value, since they do not exceed population variability observed in other late Miocene bovids. It seems that these two forms are correctly regard- ed as conspecific, meaning that
|
||
<taxonomicName id="D99F0484B455FF8CFE31D988FDE1FB3E" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[507,617,1228,1253]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFE31D988FD82FB3E" box="[507,522,1228,1253]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFDD4D988FDE1FB3E" box="[542,617,1228,1253]" italics="true" pageId="33" pageNumber="780">pallasi</emphasis>
|
||
</taxonomicName>
|
||
is equally present in Pikermi and Samos. Nevertheless, the fossil record of Samos offers strong evidences on the occurrence of a second species on the island.
|
||
</paragraph>
|
||
<paragraph id="1E207F07B455FF8CFFBBD828FC36FD3E" blockId="33.[113,646,364,1701]" lastBlockId="33.[676,1209,204,1701]" pageId="33" pageNumber="780">
|
||
Recent excavations in Samos (
|
||
<bibRefCitation id="7A0E02F6B455FF8CFE23D828FF21FA7E" author="KOUFOS G. D. & SYRIDES G. E. & KOSTOPOULOS D. S. & KOLIADIMOU K. K. & SYLVESTROU I. A. & SEITANIDES G. & VLACHOU T. D." pageId="33" pageNumber="780" pagination="877 - 885" refId="ref23152" refString="KOUFOS G. D., SYRIDES G. E., KOSTOPOULOS D. S., KOLIADIMOU K. K., SYLVESTROU I. A., SEITANIDES G. CH. F VLACHOU T. D. 1997. - New excavations in the Neogene mammalian localities of Mytilinii, Samos island, Greece. Geodiversitas 19 (4): 877 - 885." type="journal article" year="1997">
|
||
Koufos
|
||
<emphasis id="2CEBA315B455FF8CFD80D828FDF5FA5E" box="[586,637,1388,1413]" italics="true" pageId="33" pageNumber="780">et al</emphasis>
|
||
. 1997
|
||
</bibRefCitation>
|
||
) brought into the light two skull specimens of
|
||
<taxonomicName id="D99F0484B455FF8CFF45D8E8FF7CFA1E" authorityName="Gaudry" authorityYear="1861" box="[143,244,1452,1477]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="genus">
|
||
<emphasis id="2CEBA315B455FF8CFF45D8E8FF7CFA1E" box="[143,244,1452,1477]" italics="true" pageId="33" pageNumber="780">Palaeoryx</emphasis>
|
||
</taxonomicName>
|
||
from the middle Turolian fossiliferous horizons, labeled MTLA-113 and MTLB- 160 (AeMNH) (
|
||
<figureCitation id="86A46382B455FF8CFEF9D8A8FE0EF9DE" box="[307,390,1516,1541]" captionStart="FIG" captionStartId="32.[113,124,914,931]" captionTargetBox="[113,1201,203,878]" captionTargetId="figure-1329@32.[113,1201,230,878]" captionTargetPageId="32" captionText="FIG. 24. — Drawings of the skull specimens of Palaeoryx majori Schlosser, 1904 (A, B) and Palaeoryx pallasi (Wagner, 1857) (C, D); A, MTLB-160 (AeMNH); B, PIM-121; C, MTLA-113 (AeMNH); D, M10831 (BMNH) from Samos and Pikermi (approximately at the same scale)." figureDoi="http://doi.org/10.5281/zenodo.4650863" httpUri="https://zenodo.org/record/4650863/files/figure.png" pageId="33" pageNumber="780">Fig. 24</figureCitation>
|
||
). Although similar in their general morphology, these two skulls differ in a large set of secondary characters summarized in the appendix (
|
||
<tableCitation id="531D4ABCB455FF8CFEF1DB08FE2BF9BE" box="[315,419,1612,1637]" captionStart="TABLE" captionStartId="44.[113,124,203,220]" captionText="TABLE 14. — Differentiate skull characters between Palaeoryx pallasi (Wagner, 1857) and Palaeoryx majori Schlosser, 1904 from Samos." pageId="33" pageNumber="780">Table 14</tableCitation>
|
||
). These differences obviously exceed those of intraspecific variability. The cranial and dental characters of MTLA- 113 are very close to those of other specimens from Samos and Pikermi attributed to
|
||
<taxonomicName id="D99F0484B455FF8CFB9EDDA8FD66FEFE" baseAuthorityName="Wagner" baseAuthorityYear="1857" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFB9EDDA8FD66FEFE" italics="true" pageId="33" pageNumber="780">Palaeoryx pallasi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="86A46382B455FF8CFCC9DC48FCD0FEFE" box="[771,856,268,293]" captionStart="FIG" captionStartId="32.[113,124,914,931]" captionTargetBox="[113,1201,203,878]" captionTargetId="figure-1329@32.[113,1201,230,878]" captionTargetPageId="32" captionText="FIG. 24. — Drawings of the skull specimens of Palaeoryx majori Schlosser, 1904 (A, B) and Palaeoryx pallasi (Wagner, 1857) (C, D); A, MTLB-160 (AeMNH); B, PIM-121; C, MTLA-113 (AeMNH); D, M10831 (BMNH) from Samos and Pikermi (approximately at the same scale)." figureDoi="http://doi.org/10.5281/zenodo.4650863" httpUri="https://zenodo.org/record/4650863/files/figure.png" pageId="33" pageNumber="780">Fig. 24</figureCitation>
|
||
). MTLB-160 seems however identical to the specimen figured by
|
||
<bibRefCitation id="7A0E02F6B455FF8CFB9BDC68FD6EFEBE" author="SCHLOSSER M." pageId="33" pageNumber="780" pagination="21 - 118" refId="ref23722" refString="SCHLOSSER M. 1904. - Die fossiles Cavicornis von Samos. Beitrage zur Palaontologie und Geologie Osterreich-Ungarn 17: 21 - 118." type="journal article" year="1904">Schlosser (1904</bibRefCitation>
|
||
: pl. VII, fig. 5) under the name
|
||
<taxonomicName id="D99F0484B455FF8CFB9EDC08FD7AFE5E" authorityName="Schlosser" authorityYear="1904" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="majori">
|
||
<emphasis id="2CEBA315B455FF8CFB9EDC08FD7AFE5E" italics="true" pageId="33" pageNumber="780">Palaeoryx majori</emphasis>
|
||
</taxonomicName>
|
||
. Most of the characters observed in MTLB-160 reappear in the
|
||
<typeStatus id="C124C1A5B455FF8CFBE8DCC8FB18FE7E" box="[1058,1168,396,421]" pageId="33" pageNumber="780" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="D99F0484B455FF8CFD6EDCE8FB9FFE1E" authority="Andree, 1926" authorityName="Andree" authorityYear="1926" box="[676,1047,428,453]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="laticeps">
|
||
<emphasis id="2CEBA315B455FF8CFD6EDCE8FCE5FE1E" box="[676,877,428,453]" italics="true" pageId="33" pageNumber="780">Palaeoryx laticeps</emphasis>
|
||
<bibRefCitation id="7A0E02F6B455FF8CFCBDDCE8FB9FFE1E" author="ANDREE J." box="[887,1047,428,453]" pageId="33" pageNumber="780" pagination="135 - 175" refId="ref21498" refString="ANDREE J. 1926. - Neue Cavicornier aus dem Pliocan von Samos. Palaeontographica 67 (6): 135 - 175." type="journal article" year="1926">Andree, 1926</bibRefCitation>
|
||
</taxonomicName>
|
||
stored in the Münster collection with the serial number PIM- 121 (
|
||
<figureCitation id="86A46382B455FF8CFD2EDCA8FCBFFDDE" box="[740,823,492,517]" captionStart="FIG" captionStartId="32.[113,124,914,931]" captionTargetBox="[113,1201,203,878]" captionTargetId="figure-1329@32.[113,1201,230,878]" captionTargetPageId="32" captionText="FIG. 24. — Drawings of the skull specimens of Palaeoryx majori Schlosser, 1904 (A, B) and Palaeoryx pallasi (Wagner, 1857) (C, D); A, MTLB-160 (AeMNH); B, PIM-121; C, MTLA-113 (AeMNH); D, M10831 (BMNH) from Samos and Pikermi (approximately at the same scale)." figureDoi="http://doi.org/10.5281/zenodo.4650863" httpUri="https://zenodo.org/record/4650863/files/figure.png" pageId="33" pageNumber="780">Fig. 24</figureCitation>
|
||
). The braincase structure of this specimen also differs from that of another specimen in the same collection PIM-118, attributed to
|
||
<taxonomicName id="D99F0484B455FF8CFD0EDF08FCFFFDBE" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[708,887,588,613]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFD0EDF08FCFFFDBE" box="[708,887,588,613]" italics="true" pageId="33" pageNumber="780">Palaeoryx pallasi</emphasis>
|
||
</taxonomicName>
|
||
. NHMW A4779 (
|
||
<bibRefCitation id="7A0E02F6B455FF8CFB85DF08FD54FD5E" author="SOLOUNIAS N." pageId="33" pageNumber="780" pagination="1 - 232" refId="ref23812" refString="SOLOUNIAS N. 1981. - The Turolian fauna from the island of Samos, Greece. Contribution on Vertebrate Evolution 6: 1 - 232." type="journal article" year="1981">Solounias 1981</bibRefCitation>
|
||
: fig. 61D, E) looks very similar to the specimen MTLB-160 and to the type skull of
|
||
<taxonomicName id="D99F0484B455FF8CFD6EDFE8FC6AFD1E" authority="Schlosser, 1904" authorityName="Schlosser" authorityYear="1904" box="[676,994,684,709]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="majori">
|
||
<emphasis id="2CEBA315B455FF8CFD6EDFE8FD3BFD1E" box="[676,691,684,709]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFD0DDFE8FC9DFD1E" box="[711,789,684,709]" italics="true" pageId="33" pageNumber="780">majori</emphasis>
|
||
<bibRefCitation id="7A0E02F6B455FF8CFCEADFE8FC6AFD1E" author="SCHLOSSER M." box="[800,994,684,709]" pageId="33" pageNumber="780" pagination="21 - 118" refId="ref23722" refString="SCHLOSSER M. 1904. - Die fossiles Cavicornis von Samos. Beitrage zur Palaontologie und Geologie Osterreich-Ungarn 17: 21 - 118." type="journal article" year="1904">Schlosser, 1904</bibRefCitation>
|
||
</taxonomicName>
|
||
and it should be included into this species.
|
||
</paragraph>
|
||
<paragraph id="1E207F07B455FF8CFD6EDFA8FC83FC3E" blockId="33.[676,1209,204,1701]" pageId="33" pageNumber="780">
|
||
Most of the features distinguishing
|
||
<taxonomicName id="D99F0484B455FF8CFB84DFA8FB31FCDE" authorityName="Schlosser" authorityYear="1904" box="[1102,1209,748,773]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="majori">
|
||
<emphasis id="2CEBA315B455FF8CFB84DFA8FBD5FCDE" box="[1102,1117,748,773]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFBA5DFA8FB31FCDE" box="[1135,1209,748,773]" italics="true" pageId="33" pageNumber="780">majori</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="D99F0484B455FF8CFD29DE48FCCEFCFE" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[739,838,780,805]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFD29DE48FD7AFCFE" box="[739,754,780,805]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFCC9DE48FCCEFCFE" box="[771,838,780,805]" italics="true" pageId="33" pageNumber="780">pallasi</emphasis>
|
||
</taxonomicName>
|
||
can also be observed in the skull AK3-130, which therefore is placed in the former species. The single significant difference that can be observed concerns the setting of the horncores on the frontals, which appears wider in the Akkaşdagwı form, reflecting probably intraspecific variation.
|
||
</paragraph>
|
||
<paragraph id="1E207F07B455FF8CFD6EDEA8FC83F9DE" blockId="33.[676,1209,204,1701]" pageId="33" pageNumber="780">
|
||
The dentition of
|
||
<taxonomicName id="D99F0484B455FF8CFC4ADEA8FC79FBDE" authorityName="Schlosser" authorityYear="1904" box="[896,1009,1004,1029]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="majori">
|
||
<emphasis id="2CEBA315B455FF8CFC4ADEA8FC07FBDE" box="[896,911,1004,1029]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFC69DEA8FC79FBDE" box="[931,1009,1004,1029]" italics="true" pageId="33" pageNumber="780">majori</emphasis>
|
||
</taxonomicName>
|
||
is not certainly known.
|
||
<bibRefCitation id="7A0E02F6B455FF8CFD37D948FC39FBFE" author="SCHLOSSER M." box="[765,945,1036,1061]" pageId="33" pageNumber="780" pagination="21 - 118" refId="ref23722" refString="SCHLOSSER M. 1904. - Die fossiles Cavicornis von Samos. Beitrage zur Palaontologie und Geologie Osterreich-Ungarn 17: 21 - 118." type="journal article" year="1904">Schlosser (1904)</bibRefCitation>
|
||
ascribed to his new species some isolated dentitions that however could belong to
|
||
<taxonomicName id="D99F0484B455FF8CFCD8D908FCF9FBBE" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[786,881,1100,1125]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFCD8D908FCA9FBBE" box="[786,801,1100,1125]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFCE5D908FCF9FBBE" box="[815,881,1100,1125]" italics="true" pageId="33" pageNumber="780">pallasi</emphasis>
|
||
</taxonomicName>
|
||
as well. Since both species are present in Samos, the dental characters of each form need a more careful approach. The Akkaşdagwı data are also poor to solve the issue. Judging from the Akkaşdagwı toothrows it seems that
|
||
<taxonomicName id="D99F0484B455FF8CFD14D9A8FCC5FADE" authorityName="Schlosser" authorityYear="1904" box="[734,845,1260,1285]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="majori">
|
||
<emphasis id="2CEBA315B455FF8CFD14D9A8FD65FADE" box="[734,749,1260,1285]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFCCBD9A8FCC5FADE" box="[769,845,1260,1285]" italics="true" pageId="33" pageNumber="780">majori</emphasis>
|
||
</taxonomicName>
|
||
has a shorter molar row than
|
||
<taxonomicName id="D99F0484B455FF8CFD6ED848FC84FAFE" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[676,780,1292,1317]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFD6ED848FD3BFAFE" box="[676,691,1292,1317]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFD0FD848FC84FAFE" box="[709,780,1292,1317]" italics="true" pageId="33" pageNumber="780">pallasi</emphasis>
|
||
</taxonomicName>
|
||
. The available M1-M3 lengths from Akkaşdagwı range from 60.7 to 62 being smaller than that of MGL S200, MGL S199, PIK 2456, PIK 2459, BMNH M10831 (range: 65.5-69.5). Similar values occur on the specimens MGL S327 (
|
||
<quantity id="D967D2E2B455FF8CFD23D8E8FCC4FA1E" box="[745,844,1452,1477]" metricMagnitude="-2" metricUnit="m" metricValue="6.18" pageId="33" pageNumber="780" unit="mm" value="61.8">61.8 mm</quantity>
|
||
) and BMNH M10832 (
|
||
<quantity id="D967D2E2B455FF8CFB85D8E8FB3AFA1E" box="[1103,1202,1452,1477]" metricMagnitude="-2" metricUnit="m" metricValue="6.13" pageId="33" pageNumber="780" unit="mm" value="61.3">61.3 mm</quantity>
|
||
) that, however, cannot be distinguished from
|
||
<taxonomicName id="D99F0484B455FF8CFD6ED8A8FC8CF9DE" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[676,772,1516,1541]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFD6ED8A8FD3BF9DE" box="[676,691,1516,1541]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFD08D8A8FC8CF9DE" box="[706,772,1516,1541]" italics="true" pageId="33" pageNumber="780">pallasi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1E207F07B455FF8FFD6EDB48FEDEFE3E" blockId="33.[676,1209,204,1701]" lastBlockId="34.[113,646,204,485]" lastPageId="34" lastPageNumber="781" pageId="33" pageNumber="780">
|
||
The coexistence of two
|
||
<taxonomicName id="D99F0484B455FF8CFC78DB48FB90F9FE" authorityName="Gaudry" authorityYear="1861" box="[946,1048,1548,1573]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="genus">
|
||
<emphasis id="2CEBA315B455FF8CFC78DB48FB90F9FE" box="[946,1048,1548,1573]" italics="true" pageId="33" pageNumber="780">Palaeoryx</emphasis>
|
||
</taxonomicName>
|
||
species is also manifested in Halmyropotamos,
|
||
<collectingCountry id="66883F97B455FF8CFBAEDB68FB31F99E" box="[1124,1209,1580,1605]" name="Greece" pageId="33" pageNumber="780">Greece</collectingCountry>
|
||
(
|
||
<bibRefCitation id="7A0E02F6B455FF8CFD65DB08FCE6F9BE" author="MELENTIS J." box="[687,878,1612,1637]" pageId="33" pageNumber="780" pagination="283 - 411" refId="ref23386" refString="MELENTIS J. 1967. - Studien uber fossile vertebraten Griechenlands. 19. Die Pikermifauna von Halmyropotamos (Euboa, Griechenland). I Teil: Odontologie und Kraniologie. Annales geologiques des Pays helleniques 19: 283 - 411." type="journal article" year="1967">Melentis 1967</bibRefCitation>
|
||
), where both
|
||
<taxonomicName id="D99F0484B455FF8CFBF4DB08FB31F9BE" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[1086,1209,1612,1637]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B455FF8CFBF4DB08FBC5F9BE" box="[1086,1101,1612,1637]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFBA0DB08FB31F9BE" box="[1130,1209,1612,1637]" italics="true" pageId="33" pageNumber="780">pallasi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="2CEBA315B455FF8CFD67DB28FD35F95E" box="[685,701,1644,1669]" italics="true" pageId="33" pageNumber="780">=</emphasis>
|
||
<taxonomicName id="D99F0484B455FF8CFD0CDB28FCD2F95E" box="[710,858,1644,1669]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="2CEBA315B455FF8CFD0CDB28FD5DF95E" box="[710,725,1644,1669]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFD2CDB28FCD2F95E" box="[742,858,1644,1669]" italics="true" pageId="33" pageNumber="780">woodwardi</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="D99F0484B455FF8CFC54DB28FB8BF95E" authorityName="Schlosser" authorityYear="1904" box="[926,1027,1644,1669]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="33" pageNumber="780" phylum="Chordata" rank="species" species="majori">
|
||
<emphasis id="2CEBA315B455FF8CFC54DB28FC25F95E" box="[926,941,1644,1669]" italics="true" pageId="33" pageNumber="780">P</emphasis>
|
||
.
|
||
<emphasis id="2CEBA315B455FF8CFC74DB28FB8BF95E" box="[958,1027,1644,1669]" italics="true" pageId="33" pageNumber="780">majori</emphasis>
|
||
</taxonomicName>
|
||
are present. On the contrary, the presence of the genus in
|
||
<collectingCountry id="66883F97B455FF8CFBA0DBC8FB31F97E" box="[1130,1209,1676,1701]" name="Turkey" pageId="33" pageNumber="780">Turkey</collectingCountry>
|
||
is badly documented.
|
||
<bibRefCitation id="7A0E02F6B456FF8FFEAFDD88FD8AFF3E" author="KOHLER M." box="[357,514,204,229]" pageId="34" pageNumber="781" pagination="133 - 246" refId="ref22967" refString="KOHLER M. 1987. - Boviden des turkischen Miozans (Kanozoikum und Braunkohlen der Turkei). Paleontologia i Evolucio 21: 133 - 246." type="journal article" year="1987">Köhler (1987)</bibRefCitation>
|
||
refers some isolated teeth from Kayadibi and Eski Bayırköy and a few horn-core fragments and a mandible from Mahmutgazi to
|
||
<taxonomicName id="D99F0484B456FF8FFEA8DC68FD9AFE9E" baseAuthorityName="Wagner" baseAuthorityYear="1857" box="[354,530,300,325]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="34" pageNumber="781" phylum="Chordata" rank="species" species="pallasi">
|
||
<emphasis id="2CEBA315B456FF8FFEA8DC68FD9AFE9E" box="[354,530,300,325]" italics="true" pageId="34" pageNumber="781">Palaeoryx pallasi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="7A0E02F6B456FF8FFDE8DC68FF4EFEBE" author="BOUVRAIN G." pageId="34" pageNumber="781" pagination="175 - 209" refId="ref21836" refString="BOUVRAIN G. 1994 a. - Les gisements de mammiferes du Miocene superieur de Kemiklitepe, Turquie: 9. Bovidae. Bulletin du Museum national d'Histoire naturelle 4 e ser., sect. C, 16 (1): 175 - 209." type="journal article" year="1994">Bouvrain (1994a)</bibRefCitation>
|
||
describes from KTA some dentitions and one horn-core as x
|
||
<taxonomicName id="D99F0484B456FF8FFEF0DC28FE16FE5E" authorityName="Gaudry" authorityYear="1861" box="[314,414,364,389]" class="Mammalia" family="Bovidae" genus="Palaeoryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Artiodactyla" pageId="34" pageNumber="781" phylum="Chordata" rank="genus">
|
||
<emphasis id="2CEBA315B456FF8FFEF0DC28FE16FE5E" box="[314,414,364,389]" italics="true" pageId="34" pageNumber="781">Palaeoryx</emphasis>
|
||
</taxonomicName>
|
||
sp. The morphology of the described toothrows is quite similar to that from Akkaşdagwı, certifying the presence of the genus in this locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |