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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne &amp; Haring, Elisabeth" docDate="2021" docId="8D9A0FAFEA71520CA06CC1BCF4FC7D89" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Siciliaria leucophryna" docType="treatment" docVersion="4" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>De Mattia, Willy</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reier, Susanne</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haring, Elisabeth</mods:namePart>
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-12-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1077</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>175</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
<mods:identifier type="ZooBank">C28AD65A76F242CFBED7DFB3702CABCE</mods:identifier>
<mods:identifier type="Pensoft-UUID">734088641608531C8E2CC69397B000ED</mods:identifier>
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<treatment ID-GBIF-Taxon="191645971" LSID="urn:lsid:plazi:treatment:8D9A0FAFEA71520CA06CC1BCF4FC7D89" httpUri="http://treatment.plazi.org/id/8D9A0FAFEA71520CA06CC1BCF4FC7D89" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="8D9A0FAF-EA71-520C-A06C-C1BCF4FC7D89" authority="(L. Pfeiffer, 1862)" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">Siciliaria leucophryna (L. Pfeiffer, 1862)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Wagner's (1913, 1925) and Brandt's (1961) Siciliaria genital drawings A Siciliaria septemplicata, Wagner, 1913: plate 572, fig. 14 B Siciliaria grohmanniana grohmanniana, Wagner, 1925: plate 1, fig. 8 C Siciliaria calcarae calcarae, 1925: plate 3, fig. 25 D Siciliaria ferrox, Brandt, 1961: fig. 1 E overall Siciliaria / Sicania and Sicilian Mauritanica distribution F Siciliaria / Sicania and Sicilian Mauritanica distribution of taxa." figureDoi="10.3897/zookeys.1077.67081.figure1" httpUri="https://binary.pensoft.net/fig/622793" pageId="0" pageNumber="1">Figs 1.F, 11.1-11.5, 12.1-12.6, 13.12-13.14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">Clausilia leucophryna</taxonomicName>
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="1" pagination="202 - 205" refId="B76" refString="Pfeiffer, L, 1862. Diagnosen einiger neuen [sic] Landschnecken. Malakozoologische Blaetter 9: 202 - 205" title="Diagnosen einiger neuen [sic] Landschnecken." volume="9" year="1862">Pfeiffer 1862</bibRefCitation>
: 204.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryne" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryne">Siciliaria leucophryne</taxonomicName>
[sic!] -
<bibRefCitation author="Boettger, O" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B8" refString="Boettger, O, 1877. Clausilienstudien. Palaeontographica (N. F.) Suppl.3: 1-122. [4 pls. Cassel]" title="Clausilienstudien. Palaeontographica (N. F.) Suppl. 3: 1 - 122. [4 pls. Cassel]" year="1877">Boettger 1877</bibRefCitation>
: 34.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia leucophryne subsp. var. var. laudabilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="leucophryne" subSpecies="var." variety="laudabilis">Clausilia leucophryne var. laudabilis</taxonomicName>
[sic!]
<bibRefCitation author="Boettger, O" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B9" refString="Boettger, O, 1879. Gattung Clausilia Drap. In: Rossmaessler, Iconographie Land- und Suesswasser-Mollusken (1) 6(4/6): 52-153. [pls. 167-178. Wiesbaden]" title="Gattung Clausilia Drap. In: Rossmaessler, Iconographie Land- und Suesswasser-Mollusken (1) 6 (4 / 6): 52 - 153. [pls. 167 - 178. Wiesbaden]" year="1879">Boettger 1879</bibRefCitation>
: 34.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia lencophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lencophryna">Clausilia lencophryna</taxonomicName>
[sic!] -
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.51778" author="Benoit, L" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B7" refString="Benoit, L, 1881. Nuovo catalogo delle conchiglie terrestri e fluviatili della Sicilia. Atti della Reale Accademia Peloritana 3: 43-216. [Sep.: I-VI, 7-176]. https://doi.org/10.5962/bhl.title.51778" title="Nuovo catalogo delle conchiglie terrestri e fluviatili della Sicilia. Atti della Reale Accademia Peloritana 3: 43 - 216. [Sep.: I-VI, 7 - 176]." url="https://doi.org/10.5962/bhl.title.51778" year="1881">Benoit 1881</bibRefCitation>
: 108.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Westerlund" authorityYear="1892" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia confinata subsp. merens" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="confinata" subSpecies="merens">Clausilia confinata merens</taxonomicName>
<bibRefCitation author="Westerlund, CA" journalOrPublisher="zoologisch-botanischen Gesellschaft in Wien" pageId="0" pageNumber="1" pagination="25 - 48" refId="B103" refString="Westerlund, CA, 1892. Spicilegium Malacologicum. Neue Binnen-Conchylien in der palaearktischen Region. Verhandlungen der k. k. zoologisch-botanischen Gesellschaft in Wien 42: 25 - 48" title="Spicilegium Malacologicum. Neue Binnen-Conchylien in der palaearktischen Region. Verhandlungen der k. k." volume="42" year="1892">Westerlund 1892</bibRefCitation>
: 47.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia leucophryne subsp. laudabilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="leucophryne" subSpecies="laudabilis">Clausilia leucophryne laudabilis</taxonomicName>
[sic!] -
<bibRefCitation author="Westerlund, CA" journalOrPublisher="zoologisch-botanischen Gesellschaft in Wien" pageId="0" pageNumber="1" pagination="25 - 48" refId="B103" refString="Westerlund, CA, 1892. Spicilegium Malacologicum. Neue Binnen-Conchylien in der palaearktischen Region. Verhandlungen der k. k. zoologisch-botanischen Gesellschaft in Wien 42: 25 - 48" title="Spicilegium Malacologicum. Neue Binnen-Conchylien in der palaearktischen Region. Verhandlungen der k. k." volume="42" year="1892">Westerlund 1892</bibRefCitation>
: 48.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">Charpentieria leucophryna</taxonomicName>
-
<bibRefCitation DOI="https://doi.org/10.1127/arch.moll/133/2004/185" author="Beckmann, KH" journalOrPublisher="Basteria" pageId="0" pageNumber="1" refId="B4" refString="Beckmann, KH, 2004. Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133(1/2): 185-191. https://doi.org/10.1127/arch.moll/133/2004/185" title="Zur Verbreitung der endemischen nordwestsizilianischen Clausiliidae der Untergattung Charpentieria (Siciliaria) mit Beschreibung von zwei neuen Unterarten (Gastropoda: Stylommatophora: Clausiliidae). Archiv fuer Molluskenkunde 133 (1 / 2): 185 - 191." url="https://doi.org/10.1127/arch.moll/133/2004/185" year="2004">Beckmann 2004</bibRefCitation>
: 188.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">Siciliaria leucophryna</taxonomicName>
-
<bibRefCitation author="Welter-Schultes, F" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="1" refId="B101" refString="Welter-Schultes, F, 2012. European non-marine molluscs, a quide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a quide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
: 341.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">Siciliaria leucophryna</taxonomicName>
- Liberto et al. 2012: 560.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">Siciliaria leucophryna</taxonomicName>
-
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
: 7.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">Siciliaria leucophryna</taxonomicName>
-
<bibRefCitation author="Liberto, F" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="481 - 490" refId="B41" refString="Liberto, F, Colomba, MS, Reitano, A, Giglio, S, Sparacio, S, 2015. The endemic door snail of Marettimo (Egadi Islands, Sicily, Italy): Siciliaria (Siciliaria) scarificata (L. Pfeiffer, 1856) (Pulmonata Clausiliidae). Biodiversity Journal 6 (1): 481 - 490" title="The endemic door snail of Marettimo (Egadi Islands, Sicily, Italy): Siciliaria (Siciliaria) scarificata (L. Pfeiffer, 1856) (Pulmonata Clausiliidae)." volume="6" year="2015">Liberto et al. 2015</bibRefCitation>
: 489.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">Charpentieria leucophryna</taxonomicName>
-
<bibRefCitation author="De Mattia, W" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/species/171446/1326286" refId="B16" refString="De Mattia, W, 2017c. Charpentieria leucophryna. The IUCN Red List of Threatened Species. https://www.iucnredlist.org/species/171446/1326286" title="Charpentieria leucophryna. The IUCN Red List of Threatened Species." url="https://www.iucnredlist.org/species/171446/1326286" year="2017 c">De Mattia 2017c</bibRefCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Sparacio, Surdo, Viviano, Liberto &amp; Reitano" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna subsp. microinsularis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="leucophryna" subSpecies="microinsularis">Siciliaria leucophryna microinsularis</taxonomicName>
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="589 - 642" refId="B92" refString="Sparacio, I, Surdo, S, Viviano, R, Liberto, F, Reitano, A, 2021. Land molluscs from the Isola delle Femmine Nature Reserve (north-western Sicily, Italy) (Gastropoda Architaenioglossa Pulmonata). Biodiversity Journal 12 (3): 589 - 642, DOI: https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" title="Land molluscs from the Isola delle Femmine Nature Reserve (north-western Sicily, Italy) (Gastropoda Architaenioglossa Pulmonata)." url="https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" volume="12" year="2021">Sparacio et al. 2021</bibRefCitation>
: 598.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria leucophryna</emphasis>
</taxonomicName>
in the mt tree forms a separate subclade within the big
<taxonomicName lsidName="S. calcarae" pageId="0" pageNumber="1" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae</emphasis>
</taxonomicName>
clade (mean p distance 11.3%). As depicted in the combined tree (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. BI tree of based on the concatenated COI and ITS 2 data set of Siciliaria, Sicania and related genera. Colours of Siciliaria and Sicania clades correspond to those in the distribution maps (Fig. 1 E, F). BI posterior probability values (&gt; 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure6" httpUri="https://binary.pensoft.net/fig/622798" pageId="0" pageNumber="1">6</figureCitation>
), the close relationship of this species with
<taxonomicName lsidName="S. calcarae" pageId="0" pageNumber="1" rank="species" species="calcarae">
<emphasis italics="true" pageId="0" pageNumber="1">S. calcarae</emphasis>
</taxonomicName>
appears clear.
<taxonomicName authorityName="Sparacio, Surdo, Viviano, Liberto &amp; Reitano" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna subsp. microinsularis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="leucophryna" subSpecies="microinsularis">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria leucophryna microinsularis</emphasis>
</taxonomicName>
Sparacio, Surdo, Viviano, Liberto &amp; Reitano, 2021 was recently introduced from the island of Isola delle Femmine (Palermo). In the comparative notes provided by
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="589 - 642" refId="B92" refString="Sparacio, I, Surdo, S, Viviano, R, Liberto, F, Reitano, A, 2021. Land molluscs from the Isola delle Femmine Nature Reserve (north-western Sicily, Italy) (Gastropoda Architaenioglossa Pulmonata). Biodiversity Journal 12 (3): 589 - 642, DOI: https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" title="Land molluscs from the Isola delle Femmine Nature Reserve (north-western Sicily, Italy) (Gastropoda Architaenioglossa Pulmonata)." url="https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" volume="12" year="2021">Sparacio et al. (2021</bibRefCitation>
: 599), the character states reveal to fall into the variability we detected for the topotypical specimens from Grotta Conza (Palermo), both regarding shell and genital morphology. In the type locality, small specimens (
<emphasis bold="true" pageId="0" pageNumber="1">Hs</emphasis>
= 14.7 mm) and light brown colour of the shell are also known. The same applies for the connection between the AUPP and PUPP where specimens from the type locality also show a
<normalizedToken originalValue="“wide”">&quot;wide&quot;</normalizedToken>
spacing. Following the pictures provided by
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="589 - 642" refId="B92" refString="Sparacio, I, Surdo, S, Viviano, R, Liberto, F, Reitano, A, 2021. Land molluscs from the Isola delle Femmine Nature Reserve (north-western Sicily, Italy) (Gastropoda Architaenioglossa Pulmonata). Biodiversity Journal 12 (3): 589 - 642, DOI: https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" title="Land molluscs from the Isola delle Femmine Nature Reserve (north-western Sicily, Italy) (Gastropoda Architaenioglossa Pulmonata)." url="https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" volume="12" year="2021">Sparacio et al. (2021</bibRefCitation>
: figs 43, 44), the PP of
<taxonomicName lsidName="S. leucophryna subsp. microinsularis" pageId="0" pageNumber="1" rank="subSpecies" species="leucophryna" subSpecies="microinsularis">
<emphasis italics="true" pageId="0" pageNumber="1">S. leucophryna microinsularis</emphasis>
</taxonomicName>
does not appear longer if compared to the PP of the specimens from the type locality (Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Siciliaria leucophryna (L. Pfeiffer, 1862), Grotta Conza, Palermo 11.1 whole distal genital organs 11.2 internal distal part of genital organs 11.3 pseudopapilla detail. Sferracavallo, via Plauto 11.4 whole distal genital organs 11.5 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure11" httpUri="https://binary.pensoft.net/fig/622803" pageId="0" pageNumber="1">11</figureCitation>
.2, 11.3, 11.5). The variability of the sculpturing of the internal walls of the vagina refrains us to consider it a reliable taxonomic character for the whole genus
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
</taxonomicName>
. The status of
<taxonomicName lsidName="S. leucophryna subsp. microinsularis" pageId="0" pageNumber="1" rank="subSpecies" species="leucophryna" subSpecies="microinsularis">
<emphasis italics="true" pageId="0" pageNumber="1">S. leucophryna microinsularis</emphasis>
</taxonomicName>
should be further investigated morphologically as well as genetically. Anyhow,
<taxonomicName authorityName="Sparacio, Surdo, Viviano, Liberto &amp; Reitano" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna subsp. microinsularis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="leucophryna" subSpecies="microinsularis">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria leucophryna microinsularis</emphasis>
</taxonomicName>
Sparacio, Surdo, Viviano, Liberto &amp; Reitano, 2021 is here deemed as a local form, thus a junior synonym of
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria l. subsp. leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="l." subSpecies="leucophryna">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria l. leucophryna</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
The population from Via Plauto, concerning the shell, matches with the &quot;Sferracavallo form&quot; described by
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck (2013b</bibRefCitation>
: 7), showing a lower inferior lamella and the palatal edge of the clausilium plate more strongly bent upwards.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation accessionNumber="MW758902, MW757124, MW758903" collectingDate="2017-04-15" collectorName="Lab ID, W. De Mattia, J. Macor" country="Italy" elevation="150" latitude="38.187115" location="Grotta Conza" longLatPrecision="1" longitude="13.279078" municipality="Palermo" specimenCount="1" stateProvince="Sicily">
<collectingCountry name="Italy">Italy</collectingCountry>
,
<collectingRegion country="Italy" name="Sicilia">Sicily</collectingRegion>
,
<collectingMunicipality>Palermo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:8D9A0FAFEA71520CA06CC1BCF4FC7D89:F215EFCD1969E990E14A0584672B71D6" country="Italy" latitude="38.187115" longLatPrecision="1" longitude="13.279078" municipality="Palermo" name="Grotta Conza" stateProvince="Sicily">Grotta Conza</location>
,
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
asl
</elevation>
,
<geoCoordinate degrees="38" direction="north" minutes="11" orientation="latitude" precision="1" seconds="13.61" value="38.187115">38°11'13.61&quot;N</geoCoordinate>
,
<geoCoordinate degrees="13" direction="east" minutes="16" orientation="longitude" precision="1" seconds="44.68" value="13.279078">13°16'44.68&quot;E</geoCoordinate>
, [
<collectorName>Lab ID</collectorName>
55_1, COI:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758902">MW758902</accessionNumber>
, ITS2:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW757124">MW757124</accessionNumber>
;
<collectorName>Lab ID</collectorName>
55_2, COI:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758903">MW758903</accessionNumber>
],
<collectorName>W. De Mattia</collectorName>
and
<collectorName>J. Macor</collectorName>
leg.,
<collectingDate value="2017-04-15">15.iv.2017</collectingDate>
. 15 live spm, 3 dissected spm
</materialsCitation>
.
<materialsCitation accessionNumber="MW758888" collectingDate="2017-04-15" collectorName="Lab ID, W. De Mattia, J. Macor" country="Italy" county="Palermo" elevation="50" latitude="38.200367" location="Plauto" longLatPrecision="1" longitude="13.267513" municipality="Sferracavallo" specimenCount="1" stateProvince="Sicily">
<collectingCountry name="Italy">Italy</collectingCountry>
,
<collectingRegion country="Italy" name="Sicilia">Sicily</collectingRegion>
,
<collectingCounty>Palermo</collectingCounty>
,
<collectingMunicipality>Sferracavallo</collectingMunicipality>
, via
<location LSID="urn:lsid:plazi:treatment:8D9A0FAFEA71520CA06CC1BCF4FC7D89:C29AE7B69E596924D70A8101F31EABF0" country="Italy" county="Palermo" latitude="38.200367" longLatPrecision="1" longitude="13.267513" municipality="Sferracavallo" name="Plauto" stateProvince="Sicily">Plauto</location>
,
<elevation metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">
<quantity metricMagnitude="1" metricUnit="m" metricValue="5.0" unit="m" value="50.0">50 m</quantity>
asl
</elevation>
,
<geoCoordinate degrees="38" direction="north" minutes="12" orientation="latitude" precision="1" seconds="1.32" value="38.200367">38°12'1.32&quot;N</geoCoordinate>
,
<geoCoordinate degrees="13" direction="east" minutes="16" orientation="longitude" precision="1" seconds="3.05" value="13.267513">13°16'3.05&quot;E</geoCoordinate>
, [
<collectorName>Lab ID</collectorName>
43_1, COI:
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758888">MW758888</accessionNumber>
],
<collectorName>W. De Mattia</collectorName>
and
<collectorName>J. Macor</collectorName>
leg.,
<collectingDate value="2017-04-15">15.iv.2017</collectingDate>
. 7 live spm, 2 dissected spm
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="shell">
<paragraph pageId="0" pageNumber="1">Shell</paragraph>
<paragraph pageId="0" pageNumber="1">
(
<emphasis bold="true" pageId="0" pageNumber="1">
Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Siciliaria leucophryna (L. Pfeiffer, 1862), Grotta Conza, Palermo 12.1 shell 12.2 detail of the aperture 12.3 clausiliar plate double side. Sferracavallo, via Plauto 12.4 shell 12.5 detail of the aperture 12.6 detail of the columellar side of last whorl." figureDoi="10.3897/zookeys.1077.67081.figure12" httpUri="https://binary.pensoft.net/fig/622804" pageId="0" pageNumber="1">12</figureCitation>
.1-12.6, 13.12-13.14).
</emphasis>
Shell decollate; whorls ribbed; dorsal keel mostly indistinct or missing; inferior lamella moderately high; anterior upper palatal plica present, widely separated from or connected with upper palatal plica; palatal edge of clausilium plate distally slightly or not receding, palatal edge distally bent upwards (as in
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck 2013b</bibRefCitation>
) or not.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements</paragraph>
<paragraph pageId="0" pageNumber="1">
(n = 35, decollate): shell height 18.2
<normalizedToken originalValue="±">+/-</normalizedToken>
2.1, whorl width 4.3
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2, aperture height 4.5
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2, aperture width 3.4
<normalizedToken originalValue="±">+/-</normalizedToken>
0.2.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs (figs 11.1, 11.4)">
<paragraph pageId="0" pageNumber="1">
External morphology of the genital organs (Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Siciliaria leucophryna (L. Pfeiffer, 1862), Grotta Conza, Palermo 11.1 whole distal genital organs 11.2 internal distal part of genital organs 11.3 pseudopapilla detail. Sferracavallo, via Plauto 11.4 whole distal genital organs 11.5 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure11" httpUri="https://binary.pensoft.net/fig/622803" pageId="0" pageNumber="1">11</figureCitation>
.1, 11.4).
</paragraph>
<paragraph pageId="0" pageNumber="1">The FO is longer than the V (FO/V range 2.1-2.4). The VD is thin along its whole course. The FDBC of the BC+SDBC is longer than the BC+SDBC (D/BC+SDBC = 2.1). The BC+SDBC is club-like to cylindrical and longer than the V (BC+SDBC/V range 1.4-1.6), with no clear distinction between the SDBC and the BC. The apex is wide and rounded. The D is longer than the V (D/V range 2.7-2.8) and longer that the BC+SDBC (D/BC+SDBC range 1.9-2.4), thinner than the BC+SDBC and with a pointed apex. The V is short and cylindrical. The A is very large. The PC is much longer than the V (P+E/V range 2.4-3.9). The PR is long and thin. There is a clear distinction between P and E as there is a visible ER and a proximal narrowing. The E is thinner but longer than the P (E/P range 1.4-1.8), almost gradually shrinking and turning into the VD.</paragraph>
<caption doi="10.3897/zookeys.1077.67081.figure11" httpUri="https://binary.pensoft.net/fig/622803" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria leucophryna</emphasis>
</taxonomicName>
(L. Pfeiffer, 1862), Grotta Conza, Palermo
<emphasis bold="true" pageId="0" pageNumber="1">11.1</emphasis>
whole distal genital organs
<emphasis bold="true" pageId="0" pageNumber="1">11.2</emphasis>
internal distal part of genital organs
<emphasis bold="true" pageId="0" pageNumber="1">11.3</emphasis>
pseudopapilla detail. Sferracavallo, via Plauto
<emphasis bold="true" pageId="0" pageNumber="1">11.4</emphasis>
whole distal genital organs
<emphasis bold="true" pageId="0" pageNumber="1">11.5</emphasis>
internal distal part of genital organs.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1077.67081.figure12" httpUri="https://binary.pensoft.net/fig/622804" pageId="0" pageNumber="1" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 12.</emphasis>
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria leucophryna</emphasis>
</taxonomicName>
(L. Pfeiffer, 1862), Grotta Conza, Palermo
<emphasis bold="true" pageId="0" pageNumber="1">12.1</emphasis>
shell
<emphasis bold="true" pageId="0" pageNumber="1">12.2</emphasis>
detail of the aperture
<emphasis bold="true" pageId="0" pageNumber="1">12.3</emphasis>
clausiliar plate double side. Sferracavallo, via Plauto
<emphasis bold="true" pageId="0" pageNumber="1">12.4</emphasis>
shell
<emphasis bold="true" pageId="0" pageNumber="1">12.5</emphasis>
detail of the aperture
<emphasis bold="true" pageId="0" pageNumber="1">12.6</emphasis>
detail of the columellar side of last whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs (figs 11.11.2–11.2, 11.4)">
<paragraph pageId="0" pageNumber="1">
Internal morphology of the genital organs (Figs
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Siciliaria leucophryna (L. Pfeiffer, 1862), Grotta Conza, Palermo 11.1 whole distal genital organs 11.2 internal distal part of genital organs 11.3 pseudopapilla detail. Sferracavallo, via Plauto 11.4 whole distal genital organs 11.5 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure11" httpUri="https://binary.pensoft.net/fig/622803" pageId="0" pageNumber="1">11</figureCitation>
.11.2-11.2, 11.4).
</paragraph>
<paragraph pageId="0" pageNumber="1">The A shows a set of irregular fleshy folds. The P presents two big smooth longitudinal pleats that occupy almost the whole internal penial space. The fine structure of the wall is smooth. The PP is very elongated and smooth with a pointed tip. The P-E transition presents two slightly different structures. The population from the L.T. (Grotta Conza) presents a first distal ER, the PP and ELP originate from the second proximal ER. The epiphallar formula is: 1ER+2ER(PP+ELP). The V presents irregular, transverse to slightly longitudinal pleats. The population from Sferracavallo presents one ER with the PP originating from it. The ELP are not connected with the ER. The epiphallar formula is: 1ER(PP)+ELP. Distally, the E shows two main ELP that, at its mid length abruptly become finely fringed. The V presents a set of transverse-oblique smooth pleats that merge together one into another forming a kind of chevron or irregular pattern.</paragraph>
<caption doi="10.3897/zookeys.1077.67081.figure13" httpUri="https://binary.pensoft.net/fig/622805" pageId="0" pageNumber="1" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 13.</emphasis>
<taxonomicName authority="grohmanniana" authorityName="grohmanniana (Rossmaessler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. grohmanniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="grohmanniana">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana grohmanniana</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
, 1836), Monte Pellegrino, Palermo.
<emphasis bold="true" pageId="0" pageNumber="1">13.1-13.3</emphasis>
palatal plicae
<emphasis bold="true" pageId="0" pageNumber="1">13.4</emphasis>
parietal lamellae
<taxonomicName authorityName="De Mattia &amp; Reier &amp; Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria grohmanniana subsp. addaurae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="grohmanniana" subSpecies="addaurae">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria grohmanniana addaurae</emphasis>
</taxonomicName>
ssp. nov., Grotta
<normalizedToken originalValue="dell’Addaura">dell'Addaura</normalizedToken>
, Punta Priola
<emphasis bold="true" pageId="0" pageNumber="1">13.5-13.7</emphasis>
palatal plicae
<emphasis bold="true" pageId="0" pageNumber="1">13.8</emphasis>
parietal lamellae.
<taxonomicName baseAuthorityName="Philippi" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria septemplicata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="septemplicata">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria septemplicata</emphasis>
</taxonomicName>
(Philippi, 1836), Monte Gallo, Sferracavallo
<emphasis bold="true" pageId="0" pageNumber="1">13.9-13.10</emphasis>
palatal plicae
<emphasis bold="true" pageId="0" pageNumber="1">13.11</emphasis>
parietal lamellae.
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria leucophryna</emphasis>
</taxonomicName>
(L. Pfeiffer, 1862), Grotta Conza, Palermo
<emphasis bold="true" pageId="0" pageNumber="1">13.12-13.13</emphasis>
palatal plicae
<emphasis bold="true" pageId="0" pageNumber="1">13.14</emphasis>
parietal lamellae.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria leucophryna</emphasis>
</taxonomicName>
colonises limestone cliffs hiding in cracks or under stones. It is very common on isolated boulders and tree trunks and barks in the xerophilic scrub around Grotta Conza. The species is quite abundant where found. According to
<bibRefCitation author="De Mattia, W" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="1" publicationUrl="https://www.iucnredlist.org/species/171446/1326286" refId="B16" refString="De Mattia, W, 2017c. Charpentieria leucophryna. The IUCN Red List of Threatened Species. https://www.iucnredlist.org/species/171446/1326286" title="Charpentieria leucophryna. The IUCN Red List of Threatened Species." url="https://www.iucnredlist.org/species/171446/1326286" year="2017 c">De Mattia (2017c)</bibRefCitation>
the species is Endangered following the IUCN criteria B1ab(iii,v)+2ab(iii,v). The map provided by De Mattia (2017) partially misses a portion of the actual distribution area, as the western part was left out from the perimeter (Pizzo Mollica and Isola delle Femmine).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1862" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria leucophryna" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="leucophryna">
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria leucophryna</emphasis>
</taxonomicName>
is known to occur from a small area from Isola delle Femmine to Sferracavallo, including the northernmost sides of Pizzo Mollica and Pizzo Manolfo (where Grotta Conza is located) [(
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="589 - 642" refId="B92" refString="Sparacio, I, Surdo, S, Viviano, R, Liberto, F, Reitano, A, 2021. Land molluscs from the Isola delle Femmine Nature Reserve (north-western Sicily, Italy) (Gastropoda Architaenioglossa Pulmonata). Biodiversity Journal 12 (3): 589 - 642, DOI: https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" title="Land molluscs from the Isola delle Femmine Nature Reserve (north-western Sicily, Italy) (Gastropoda Architaenioglossa Pulmonata)." url="https://doi.org/10.31396/Biodiv.Jour.2021.12.3.589.624" volume="12" year="2021">Sparacio et al. (2021)</bibRefCitation>
].
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" publicationUrl="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" refId="B69" refString="Nordsieck, H, 2013b. Revisory remarks on the species of Siciliaria Vest from N. W. Sicily. http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" title="Revisory remarks on the species of Siciliaria Vest from N. W. Sicily." url="http://hnords.de/5356429d6b11adc0b/535642a1dc1365e05/index.html" year="2013 b">Nordsieck (2013b)</bibRefCitation>
reports also Monte Gallo, despite recent field collecting failed to find it there. Further field research throughout the whole Pizzo Manolfo-Pizzo Castellaccio-Cozzo San Rocco massif is required to precisely define the distribution of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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