treatments-xml/data/84/08/2A/84082A4BC02DB51547F03DE5EC88FCC7.xml

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<mods:title id="644E75E3EECA0C7D937E37AB9C106653">New Bryodelphax species (Heterotardigrada: Echiniscidae) from Western Borneo (Sarawak), with new molecular data for the genus</mods:title>
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<taxonomicName id="CBA1E0DEC02DB51047F03DE5EF9BFDD4" ID-CoL="cd44a8a2-1c37-416b-9363-302edc066b25" authority="Gąsiorek, 2018" authorityName="Gąsiorek" authorityYear="2018" box="[238,486,989,1013]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="374" phylum="Tardigrada" rank="species" species="arenosus" status="sp. nov.">
<emphasis id="3ED5474FC02DB51047F03DE5EF9BFDD4" bold="true" box="[238,486,989,1013]" italics="true" pageId="3" pageNumber="374">Bryodelphax arenosus</emphasis>
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<paragraph id="0C1E9B5DC02DB510462F3DC5EC4CFA34" blockId="3.[238,628,989,1045]" box="[305,561,1021,1045]" pageId="3" pageNumber="374">
(
<figureCitation id="949A87D8C02DB51046273DC5EFDCFA34" box="[313,417,1021,1045]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="3.[813,848,1315,1337]" captionStartId-1="4.[136,171,1480,1502]" captionStartId-2="5.[113,148,1973,1995]" captionTargetBox-0="[814,1451,189,1300]" captionTargetBox-1="[136,1474,189,1465]" captionTargetBox-2="[113,1451,978,1958]" captionTargetId-0="figure-768@3.[810,1454,185,1304]" captionTargetId-1="figure-247@4.[136,1474,189,1465]" captionTargetId-2="figure-10@5.[113,1451,978,1958]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Fig. 2. Schematic arrangement of plates on dorsum of Bryodelphax arenosus sp. nov. Abbreviations: c = the caudal plate; pl = lateral portion of the paired plate; sc = central portion of the scapular plate; sl = lateral portion of the scapular plate." captionText-1="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." captionText-2="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi-0="http://doi.org/10.5281/zenodo.5358947" figureDoi-1="http://doi.org/10.5281/zenodo.5358949" figureDoi-2="http://doi.org/10.5281/zenodo.5358951" httpUri-0="https://zenodo.org/record/5358947/files/figure.png" httpUri-1="https://zenodo.org/record/5358949/files/figure.png" httpUri-2="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">Figs. 2–4</figureCitation>
, 
<tableCitation id="4123AEE6C02DB51046B03DC5EC70FA34" box="[430,525,1021,1045]" captionStart="Table 2" captionStartId="5.[113,168,189,211]" captionText="Table 2. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature females of Bryodelphax arenosus sp. nov. mounted in Hoyer’s medium (N – number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD – standard deviation)." pageId="3" pageNumber="374">Tables 2</tableCitation>
, 
<tableCitation id="4123AEE6C02DB51045053DC5EC55FA34" box="[539,552,1021,1045]" captionStart="Table 3" captionStartId="6.[135,190,189,211]" captionText="Table 3. Uncorrected pairwise distance (in %) for available Bryodelphax species." pageId="3" pageNumber="374">3</tableCitation>
)
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<paragraph id="0C1E9B5DC02DB510476F3A05ECFDFBD4" blockId="3.[113,753,1085,1525]" pageId="3" pageNumber="374">
<emphasis id="3ED5474FC02DB510476F3A05EF2FFA74" bold="true" box="[113,338,1085,1109]" pageId="3" pageNumber="374">Material examined.</emphasis>
<materialsCitation id="BCC99100C02DB51046453A05EC01FBD4" ID-GBIF-Occurrence="3340080301" collectingDate="2017-08-05" collectorName="Piotr Gasiorek &amp; Artur Oczkowski" country="Poland" county="National University" elevation="106" latitude="1.7233334" location="Institute of Zoology" longLatPrecision="21" longitude="110.46388" municipality="Raffles Museum of Biodiversity Research" pageId="3" pageNumber="374" specimenCount="17" typeStatus="holotype">
All individuals were mature females. 
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(slide no. MY.813.01) and 
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(slides MY.813.01–11), six additional processed for DNA extraction and sequencing. Type locality: Bako Peninsula, lowland coastal rainforest close to the Tajor Waterfall (
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, 
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; 
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<quantity id="CB5936B8C02DB510460A3AE5EF23FAD4" box="[276,350,1245,1269]" metricMagnitude="2" metricUnit="m" metricValue="1.06" pageId="3" pageNumber="374" unit="m" value="106.0">106 m</quantity>
asl
</elevation>
), moss from a tree, coll. 
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&amp; 
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, 
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5 
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August 2017
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. 
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and 
<specimenCount id="1AA750D4C02DB51047BD3B25EF38FB17" box="[163,325,1309,1334]" pageId="3" pageNumber="374" type="generic" typeStatus="paratypes">nine paratypes</specimenCount>
deposited in the 
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and 
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, 
<location id="097ECD86C02DB51046913B05ECF4FB74" LSID="urn:lsid:plazi:treatment:84082A4BC02DB51547F03DE5EC88FCC7:097ECD86C02DB51046913B05ECF4FB74" box="[399,649,1341,1365]" country="Poland" county="National University" latitude="1.7233334" longLatPrecision="21" longitude="110.46388" municipality="Raffles Museum of Biodiversity Research" name="Jagiellonian University" pageId="3" pageNumber="374">Jagiellonian University</location>
, 
<location id="097ECD86C02DB510458A3B05EC90FB74" LSID="urn:lsid:plazi:treatment:84082A4BC02DB51547F03DE5EC88FCC7:097ECD86C02DB510458A3B05EC90FB74" box="[660,749,1341,1365]" country="Poland" county="National University" latitude="1.7233334" longLatPrecision="21" longitude="110.46388" municipality="Raffles Museum of Biodiversity Research" name="Krakow" pageId="3" pageNumber="374">Kraków</location>
, 
<collectingCountry id="74B6DBCDC02DB510476F3B65EEC1FB54" box="[113,188,1373,1397]" name="Poland" pageId="3" pageNumber="374">Poland</collectingCountry>
(slides MY.813.01–06), 
<specimenCount id="1AA750D4C02DB51046C83B65EC03FB57" box="[470,638,1373,1398]" pageId="3" pageNumber="374" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
deposited in the 
<location id="097ECD86C02DB51047A53B45EFB1FBB4" LSID="urn:lsid:plazi:treatment:84082A4BC02DB51547F03DE5EC88FCC7:097ECD86C02DB51047A53B45EFB1FBB4" box="[187,460,1405,1429]" country="Poland" county="National University" latitude="1.7233334" longLatPrecision="21" longitude="110.46388" municipality="Raffles Museum of Biodiversity Research" name="Natural History Museum" pageId="3" pageNumber="374">Natural History Museum</location>
of 
<collectingCountry id="74B6DBCDC02DB51046E93B45EC20FBB4" box="[503,605,1405,1429]" name="Denmark" pageId="3" pageNumber="374">Denmark</collectingCountry>
, 
<location id="097ECD86C02DB51045753B45EC8DFBB4" LSID="urn:lsid:plazi:treatment:84082A4BC02DB51547F03DE5EC88FCC7:097ECD86C02DB51045753B45EC8DFBB4" box="[619,752,1405,1429]" country="Poland" county="National University" latitude="1.7233334" longLatPrecision="21" longitude="110.46388" municipality="Raffles Museum of Biodiversity Research" name="Copenhagen" pageId="3" pageNumber="374">Copenhagen</location>
(slides MY.813.07–09), and last 
<specimenCount id="1AA750D4C02DB51046FD3BA6EC03FB97" box="[483,638,1438,1462]" pageId="3" pageNumber="374" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
deposited in the 
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, 
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of 
<collectingCountry id="74B6DBCDC02DB51046143BE5EF0BFBD4" box="[266,374,1501,1525]" name="Singapore" pageId="3" pageNumber="374">Singapore</collectingCountry>
(slides MY.813.10–11)
</materialsCitation>
.
</paragraph>
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<paragraph id="0C1E9B5DC02DB51044333B1BEA8BFBAE" blockId="3.[813,1452,1315,1423]" pageId="3" pageNumber="374">
Fig. 2. Schematic arrangement of plates on dorsum of 
<taxonomicName id="CBA1E0DEC02DB510422F3B1BEDF9FB74" authority="Gąsiorek, 2018" authorityName="Gąsiorek" authorityYear="2018" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="374" phylum="Tardigrada" rank="species" species="arenosus" status="sp. nov.">
<emphasis id="3ED5474FC02DB510422F3B1BEDF9FB74" italics="true" pageId="3" pageNumber="374">Bryodelphax arenosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25E6FA34C02DB51044943B78EDA5FB74" box="[906,984,1344,1365]" pageId="3" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations: c = the caudal plate; pl = lateral portion of the paired plate; sc = central portion of the scapular plate; sl = lateral portion of the scapular plate.
</paragraph>
</caption>
<subSubSection id="44BBC8D6C02DB517476F3825EACCF926" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374" type="description">
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<emphasis id="3ED5474FC02DB510476F3825EFB2F814" bold="true" box="[113,463,1565,1589]" pageId="3" pageNumber="374">Description of the new species.</emphasis>
Adults (i.e., from the third instar onwards, with developed gonopore; measurements in 
<tableCitation id="4123AEE6C02DB510476F3865EEBAF854" box="[113,199,1629,1653]" captionStart="Table 2" captionStartId="5.[113,168,189,211]" captionText="Table 2. Measurements [in µm] and sc indices [in % of the scapular plate length] of selected morphological structures of type mature females of Bryodelphax arenosus sp. nov. mounted in Hoyer’s medium (N – number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD – standard deviation)." pageId="3" pageNumber="374">Table 2</tableCitation>
).
</paragraph>
<paragraph id="0C1E9B5DC02DB517476F38A5EDB3F886" blockId="3.[113,753,1693,2101]" lastBlockId="4.[835,1475,1615,1703]" lastPageId="4" lastPageNumber="375" pageId="3" pageNumber="374">
Body yellowish, pearly opalescent; eyes absent or not visible after preparation. Both primary and secondary clavae oval and small (
<figureCitation id="949A87D8C02DB51047A738E5EF79F8D4" box="[185,260,1757,1781]" captionStart="Fig" captionStartId="3.[813,848,1315,1337]" captionTargetBox="[814,1451,189,1300]" captionTargetId="figure-768@3.[810,1454,185,1304]" captionTargetPageId="3" captionText="Fig. 2. Schematic arrangement of plates on dorsum of Bryodelphax arenosus sp. nov. Abbreviations: c = the caudal plate; pl = lateral portion of the paired plate; sc = central portion of the scapular plate; sl = lateral portion of the scapular plate." figureDoi="http://doi.org/10.5281/zenodo.5358947" httpUri="https://zenodo.org/record/5358947/files/figure.png" pageId="3" pageNumber="374">Figs. 2</figureCitation>
, 
<figureCitation id="949A87D8C02DB510461138E5EF52F8D4" box="[271,303,1757,1781]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">3C</figureCitation>
, 
<figureCitation id="949A87D8C02DB510462438E5EF22F8D4" box="[314,351,1757,1781]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4A</figureCitation>
). 
<emphasis id="3ED5474FC02DB510466E38E6EF8EF8D4" box="[368,499,1758,1781]" italics="true" pageId="3" pageNumber="374">Cirri interni</emphasis>
and 
<emphasis id="3ED5474FC02DB510453B38E6EC12F8D4" box="[549,623,1758,1781]" italics="true" pageId="3" pageNumber="374">externi</emphasis>
with poorly developed cirrophores. Cirri 
<emphasis id="3ED5474FC02DB51046B938C6EFCAF934" box="[423,439,1790,1813]" italics="true" pageId="3" pageNumber="374">A</emphasis>
long (on average reaching ⅓ of the body length, whereas 
<taxonomicName id="CBA1E0DEC02DB51046843925EC5FF914" authorityName="Thulin" authorityYear="1928" box="[410,546,1821,1845]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="3" pageNumber="374" phylum="Tardigrada" rank="genus">
<emphasis id="3ED5474FC02DB51046843925EC5FF914" box="[410,546,1821,1845]" italics="true" pageId="3" pageNumber="374">Bryodelphax</emphasis>
</taxonomicName>
typically has cirrus 
<emphasis id="3ED5474FC02DB510476F3906EEFCF974" box="[113,129,1854,1877]" italics="true" pageId="3" pageNumber="374">A</emphasis>
/body length ratio around ¼; 
<figureCitation id="949A87D8C02DB51046D43905EC6AF974" box="[458,535,1853,1877]" captionStart="Fig" captionStartId="3.[813,848,1315,1337]" captionTargetBox="[814,1451,189,1300]" captionTargetId="figure-768@3.[810,1454,185,1304]" captionTargetPageId="3" captionText="Fig. 2. Schematic arrangement of plates on dorsum of Bryodelphax arenosus sp. nov. Abbreviations: c = the caudal plate; pl = lateral portion of the paired plate; sc = central portion of the scapular plate; sl = lateral portion of the scapular plate." figureDoi="http://doi.org/10.5281/zenodo.5358947" httpUri="https://zenodo.org/record/5358947/files/figure.png" pageId="3" pageNumber="374">Figs. 2</figureCitation>
, 
<figureCitation id="949A87D8C02DB51045393905EC37F974" box="[551,586,1853,1877]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">3A</figureCitation>
). Dorsal plates with faint intra-cuticular pillars visible only under 100× magnification with immersion oil (
<figureCitation id="949A87D8C02DB51046C13945EC46F9B4" box="[479,571,1917,1941]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3C</figureCitation>
, 
<figureCitation id="949A87D8C02DB510455B3945EC28F9B4" box="[581,597,1917,1941]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4</figureCitation>
), better visible on the scapular and caudal plates (
<figureCitation id="949A87D8C02DB51046C839A5EC4CF994" box="[470,561,1949,1973]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3C</figureCitation>
, 
<figureCitation id="949A87D8C02DB510452539A5EC01F994" box="[571,636,1949,1973]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4A, C</figureCitation>
), pillars on the posterior portion of the caudal plate are also the largest (
<figureCitation id="949A87D8C02DB510476739E5EEA7F9D4" box="[121,218,2013,2037]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3C</figureCitation>
, 
<figureCitation id="949A87D8C02DB51047F739E5EF71F9D4" box="[233,268,2013,2037]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4C</figureCitation>
). Distinct dark ridges are present on lateral portions of the scapular, paired, and caudal plates, forming characteristic and stable ornamented pattern (
<figureCitation id="949A87D8C02DB51045763625EC91F614" box="[616,748,2077,2101]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3B, C</figureCitation>
, 
<figureCitation id="949A87D8C02DB51044333BFEED40FBFF" box="[813,829,1478,1502]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4</figureCitation>
). Pseudopores or pores (SEM observations are needed to verify the morphology of the cuticular depressions) minute and barely visible (visible in PCM as bright dots with blurred margins; 
<figureCitation id="949A87D8C02DB510448E381EED91F81F" box="[912,1004,1574,1598]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3C</figureCitation>
, 
<figureCitation id="949A87D8C02DB51044E9381EEA7AF81F" box="[1015,1031,1574,1598]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4</figureCitation>
), sometimes absent. Pseudopores/pores distributed unevenly, i.e., on the anterocentral portion of the scapular plate 8–21 pseudopores/pores per 100 μm 
<superScript id="FBD43615C02DB5104278385DEB13F852" attach="left" box="[1382,1390,1637,1651]" fontSize="6" pageId="3" pageNumber="374">2</superScript>
were observed (x =17, N=10; 
<figureCitation id="949A87D8C02DB510432D38BEEAF7F8BF" box="[1075,1162,1670,1694]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">Fig. 4A</figureCitation>
), on the central portion of the caudal plate: 11–18 pseudopores/pores per 100 μm 
<superScript id="FBD43615C02DB5104244389DEB1FF892" attach="left" box="[1370,1378,1701,1715]" fontSize="6" pageId="3" pageNumber="374">2</superScript>
(x =15, N=10; 
<figureCitation id="949A87D8C02DB510446938FEEDB8F8FF" box="[887,965,1734,1758]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">Fig. 4C</figureCitation>
), whereas sparse and bleak pseudopores/pores occur rarely on other, especially paired, plates. Scapular plate deeply faceted by dark ridges, with six or eight lighter rectangular areas in the central portion (
<figureCitation id="949A87D8C02DB51043F7391EEB10F91F" box="[1257,1389,1830,1854]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3A, C</figureCitation>
, 
<figureCitation id="949A87D8C02DB5104262391EEBDDF91F" box="[1404,1440,1830,1854]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4A</figureCitation>
). Scapular sutures clearly separate lateral portions from the central part, extending from the base of cirrophore 
<emphasis id="3ED5474FC02DB5104261395FEBF2F95F" box="[1407,1423,1895,1918]" italics="true" pageId="3" pageNumber="374">A</emphasis>
to the posterior margin of the plate (
<figureCitation id="949A87D8C02DB51043BE39BEEB5CF9BF" box="[1184,1313,1926,1950]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3B, C</figureCitation>
, 
<figureCitation id="949A87D8C02DB510423139BEEB2EF9BF" box="[1327,1363,1926,1950]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4A</figureCitation>
). Paired plates divided into two roughly equal anterior and posterior parts by a transverse stripe (
<figureCitation id="949A87D8C02DB510436339FEEAB1F9FF" box="[1149,1228,1990,2014]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3</figureCitation>
, 
<figureCitation id="949A87D8C02DB51043C139FEEA82F9FF" box="[1247,1279,1990,2014]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4B</figureCitation>
). Caudal plate mildly faceted in the central part by the same dark ridges as the scapular plate (
<figureCitation id="949A87D8C02DB510433C363EEA07F63F" box="[1058,1146,2054,2078]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Fig. 3A</figureCitation>
), but the faceting is fainter and often poorly visible (
<figureCitation id="949A87D8C02DB5104355361EEAD1F61F" box="[1099,1196,2086,2110]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="3" pageNumber="374">Figs. 3C</figureCitation>
, 
<figureCitation id="949A87D8C02DB51043A2361EEAA2F61F" box="[1212,1247,2086,2110]" captionStart="Fig" captionStartId="5.[113,148,1973,1995]" captionTargetBox="[113,1451,978,1958]" captionTargetId="figure-10@5.[113,1451,978,1958]" captionTargetPageId="5" captionText="Fig. 4. Close-up on various parts of dorsal armature of Bryodelphax arenosus sp. nov. A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="3" pageNumber="374">4C</figureCitation>
). Median plates 1 and 2 divided transversally into two unequal parts (
<figureCitation id="949A87D8C02AB51745AB3877ED7FF846" box="[693,770,1615,1639]" captionStart="Fig" captionStartId="3.[813,848,1315,1337]" captionTargetBox="[814,1451,189,1300]" captionTargetId="figure-768@3.[810,1454,185,1304]" captionTargetPageId="3" captionText="Fig. 2. Schematic arrangement of plates on dorsum of Bryodelphax arenosus sp. nov. Abbreviations: c = the caudal plate; pl = lateral portion of the paired plate; sc = central portion of the scapular plate; sl = lateral portion of the scapular plate." figureDoi="http://doi.org/10.5281/zenodo.5358947" httpUri="https://zenodo.org/record/5358947/files/figure.png" pageId="4" pageNumber="375">Figs. 2</figureCitation>
, 
<figureCitation id="949A87D8C02AB51747963857EEEBF8A6" box="[136,150,1647,1671]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="4" pageNumber="375">3</figureCitation>
), and additionally the anterior portion of median 2 has a transverse suture (
<figureCitation id="949A87D8C02AB517464E38B7EFE9F886" box="[336,404,1679,1703]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="4" pageNumber="375">Fig. 3</figureCitation>
). Median plate 3 large, triangular in shape, and with a roundish posterior edge (
<figureCitation id="949A87D8C02AB51745923897ECA6F8E6" box="[652,731,1711,1735]" captionStart="Fig" captionStartId="3.[813,848,1315,1337]" captionTargetBox="[814,1451,189,1300]" captionTargetId="figure-768@3.[810,1454,185,1304]" captionTargetPageId="3" captionText="Fig. 2. Schematic arrangement of plates on dorsum of Bryodelphax arenosus sp. nov. Abbreviations: c = the caudal plate; pl = lateral portion of the paired plate; sc = central portion of the scapular plate; sl = lateral portion of the scapular plate." figureDoi="http://doi.org/10.5281/zenodo.5358947" httpUri="https://zenodo.org/record/5358947/files/figure.png" pageId="4" pageNumber="375">Figs. 2</figureCitation>
, 
<figureCitation id="949A87D8C02AB51745F43897EC84F8E6" box="[746,761,1711,1735]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="4" pageNumber="375">3</figureCitation>
). Thin, non-sculptured supplementary lateral platelets present at the levels of median plates (three pairs of platelets on each body side: a pair between scapular plate and first pair of the segmental plates, a pair between paired plates, and a pair between second pair of segmental plates and caudal plate; 
<figureCitation id="949A87D8C02AB51747D03957EF35F9A6" box="[206,328,1903,1927]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="4" pageNumber="375">Fig. 3B, C</figureCitation>
). Ventral cuticle smooth, without plates or intra-cuticular pillars. Papillae on legs I and IV absent or not detectable in PCM. Large, sclerotised pedal plates IV present, with irregular margin, but never forming a dentate collar (
<figureCitation id="949A87D8C02AB51747CD39D7EF5BF626" box="[211,294,2031,2055]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="4" pageNumber="375">Fig. 3C</figureCitation>
, white arrowhead). External claws of all legs smooth, internal claws with minuscule bumps near their bases, which can be spurs (
<figureCitation id="949A87D8C02AB51743703877EABEF846" box="[1134,1219,1615,1639]" captionStart="Fig" captionStartId="4.[136,171,1480,1502]" captionTargetBox="[136,1474,189,1465]" captionTargetId="figure-247@4.[136,1474,189,1465]" captionTargetPageId="4" captionText="Fig. 3. Bryodelphax arenosus sp. nov., habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="4" pageNumber="375">Fig. 3C</figureCitation>
, insert). However, they are not measurable in PCM since barely divergent from the claw branch.
</paragraph>
<caption id="58DECBD5C02AB51747963BF0EDD6F836" ID-DOI="http://doi.org/10.5281/zenodo.5358949" ID-Zenodo-Dep="5358949" httpUri="https://zenodo.org/record/5358949/files/figure.png" pageId="4" pageNumber="375" startId="4.[136,171,1480,1502]" targetBox="[136,1474,189,1465]" targetPageId="4">
<paragraph id="0C1E9B5DC02AB51747963BF0EDD6F836" blockId="4.[136,1475,1480,1560]" pageId="4" pageNumber="375">
Fig. 3. 
<taxonomicName id="CBA1E0DEC02AB51747CD3BF0EFD3FBFF" authority="Gąsiorek, 2018" authorityName="Gąsiorek" authorityYear="2018" box="[211,430,1480,1502]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="4" pageNumber="375" phylum="Tardigrada" rank="species" species="arenosus" status="sp. nov.">
<emphasis id="3ED5474FC02AB51747CD3BF0EFD3FBFF" box="[211,430,1480,1502]" italics="true" pageId="4" pageNumber="375">Bryodelphax arenosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25E6FA34C02AB51746A93BF0EC74FBFC" box="[439,521,1480,1501]" pageId="4" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
, habitus (PCM). A, dorsal view (paratype); B, dorsolateral view (paratype); C, holotype dorsal armature in close-up (claws III inserted in the upper right corner). Empty arrowheads indicate ornamented portions of dorsal plates, whereas white arrowhead points pedal plate IV devoid of dentate collar. Scale bars in μm.
</paragraph>
</caption>
<paragraph id="0C1E9B5DC02AB517445D38F7EACCF926" blockId="4.[835,1473,1743,1799]" pageId="4" pageNumber="375">No juveniles, larvae (i.e., the second and first instars, respectively) or eggs were found.</paragraph>
</subSubSection>
<subSubSection id="44BBC8D6C02AB515445D3917EC15FC47" lastPageId="6" lastPageNumber="377" pageId="4" pageNumber="375" type="discussion">
<paragraph id="0C1E9B5DC02AB517445D3917EB11F946" blockId="4.[835,1474,1839,1895]" pageId="4" pageNumber="375">
<emphasis id="3ED5474FC02AB517445D3917EDCFF966" bold="true" box="[835,946,1839,1863]" pageId="4" pageNumber="375">Remarks.</emphasis>
All found specimens were mature females, thus it is likely that the new species is parthenogenetic.
</paragraph>
<paragraph id="0C1E9B5DC02AB515445D39B6EC15FC47" blockId="4.[835,1475,1934,2087]" lastBlockId="6.[136,775,558,614]" lastPageId="6" lastPageNumber="377" pageId="4" pageNumber="375">
<emphasis id="3ED5474FC02AB517445D39B6ED80F986" bold="true" box="[835,1021,1934,1959]" pageId="4" pageNumber="375">DNA sequences.</emphasis>
<materialsCitation id="BCC99100C02AB515431B39B7EC18FC47" ID-GBIF-Occurrence="3340080302" accessionNumber="MH414953, XX000000" collectionCode="MH" lastPageId="6" lastPageNumber="377" pageId="4" pageNumber="375" specimenCount="1">
New 18S and 28S rRNA sequences were obtained for 
<specimenCount id="1AA750D4C02AB51744D23997EA13F9E6" box="[972,1134,1967,1991]" pageId="4" pageNumber="375" type="generic">two specimens</specimenCount>
; each fragment was represented by a single haplotype and is deposited in GenBank: 
<accessionNumber id="13F206BEC02AB517445B39D7EDBCF626" box="[837,961,2031,2055]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH414953" pageId="4" pageNumber="375">
<collectionCode id="6AB00398C02AB517445B39D7ED0DF626" box="[837,880,2031,2055]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="4" pageNumber="375">MH</collectionCode>
414953
</accessionNumber>
(18S rRNA, 1112 bp long), XX000000 (28S rRNA, 688 bp long). p-distances between sequenced species of 
<taxonomicName id="CBA1E0DEC028B51547BA3C16EF51FC67" authorityName="Thulin" authorityYear="1928" box="[164,300,558,582]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="377" phylum="Tardigrada" rank="genus">
<emphasis id="3ED5474FC028B51547BA3C16EF51FC67" box="[164,300,558,582]" italics="true" pageId="6" pageNumber="377">Bryodelphax</emphasis>
</taxonomicName>
ranged from 0.8% to 2.8% (18S rRNA), and from 7.5% to 19.3% (28S rRNA) (
<tableCitation id="4123AEE6C028B515451B3C76EC26FC47" box="[517,603,590,614]" captionStart="Table 3" captionStartId="6.[135,190,189,211]" captionText="Table 3. Uncorrected pairwise distance (in %) for available Bryodelphax species." pageId="6" pageNumber="377">Table 3</tableCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="58DECBD5C02BB516476F3E85EA48FF2C" pageId="5" pageNumber="376" startId="5.[113,168,189,211]">
<paragraph id="0C1E9B5DC02BB516476F3E85EA48FF2C" blockId="5.[113,1452,189,269]" pageId="5" pageNumber="376">
Table 2. Measurements [in µm] and 
<emphasis id="3ED5474FC02BB51646FE3E86EF89FEF2" box="[480,500,190,211]" italics="true" pageId="5" pageNumber="376">sc</emphasis>
indices [in % of the scapular plate length] of selected morphological structures of type mature females of 
<taxonomicName id="CBA1E0DEC02BB51647FE3EE2EFC7FED1" authority="Gąsiorek, 2018" authorityName="Gąsiorek" authorityYear="2018" box="[224,442,218,240]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="376" phylum="Tardigrada" rank="species" species="arenosus" status="sp. nov.">
<emphasis id="3ED5474FC02BB51647FE3EE2EFC7FED1" box="[224,442,218,240]" italics="true" pageId="5" pageNumber="376">Bryodelphax arenosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25E6FA34C02BB51646DC3EE3EC6DFED1" box="[450,528,219,240]" pageId="5" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
mounted in Hoyer’s medium (N – number of specimens/structures measured, RANGE refers to the smallest and the largest structure among all measured specimens; SD – standard deviation).
</paragraph>
</caption>
<paragraph id="0C1E9B5DC02BB51645B93F07EBECFDA2" pageId="5" pageNumber="376">
<table id="7EA169FDC02B4AEC479E3F06EBEAFDA5" box="[128,1431,318,900]" gridcols="14" gridrows="7" pageId="5" pageNumber="376">
<tr id="B291991FC02B4AEC479E3F07EBEAFF75" box="[128,1431,319,340]" gridrow="0" pageId="5" pageNumber="376" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-7="1">
<th id="F140F063C02B4AEC45B93F07EC97FF75" box="[679,746,319,340]" gridcol="5" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51645B93F07EC95FF75" bold="true" box="[679,744,319,340]" pageId="5" pageNumber="376">Range</emphasis>
</th>
<th id="F140F063C02B4AEC44D93F07EA31FF75" box="[967,1100,319,340]" colspan="2" colspanRight="1" gridcol="8" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51644F53F07EA5BFF75" bold="true" box="[1003,1062,319,340]" pageId="5" pageNumber="376">Mean</emphasis>
</th>
<th id="F140F063C02B4AEC436C3F07EA96FF75" box="[1138,1259,319,340]" colspan="2" colspanRight="1" gridcol="10" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643BF3F06EABDFF75" bold="true" box="[1185,1216,318,340]" pageId="5" pageNumber="376">SD</emphasis>
</th>
<th id="F140F063C02B4AEC420C3F07EBEAFF75" box="[1298,1431,319,340]" colspan="2" colspanRight="1" gridcol="12" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51642363F07EBF8FF75" bold="true" box="[1320,1413,319,340]" pageId="5" pageNumber="376">Holotype</emphasis>
</th>
</tr>
<tr id="B291991FC02B4AEC479E3F6DEBEAFF4A" box="[128,1431,341,363]" gridrow="1" pageId="5" pageNumber="376" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="F140F063C02B4AEC479E3F6DEFE9FF4A" box="[128,404,341,363]" gridcol="0" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51647CB3F6DEF43FF4A" bold="true" box="[213,318,341,363]" pageId="5" pageNumber="376">Character</emphasis>
</th>
<td id="F140F063C02B4AEC46AD3F6DEFB6FF4A" box="[435,459,341,363]" gridcol="1" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51646A83F6EEFBAFF4A" bold="true" box="[438,455,342,363]" pageId="5" pageNumber="376">N</emphasis>
</td>
</tr>
<tr id="B291991FC02B4AEC479E3F55EBEAFFA3" box="[128,1431,365,386]" gridrow="2" pageId="5" pageNumber="376" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-7="1">
<td id="F140F063C02B4AEC45333F55EC32FFA3" box="[557,591,365,386]" gridcol="3" gridrow="2" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51645333F55EC32FFA3" bold="true" box="[557,591,365,386]" pageId="5" pageNumber="376">µm</emphasis>
</td>
<td id="F140F063C02B4AEC443F3F55ED48FFA3" box="[801,821,365,386]" gridcol="6" gridrow="2" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB516443F3F55ED48FFA3" bold="true" box="[801,821,365,386]" pageId="5" pageNumber="376">sc</emphasis>
</td>
<td id="F140F063C02B4AEC44D93F55ED8EFFA3" box="[967,1011,365,386]" gridcol="8" gridrow="2" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51644D23F55ED93FFA3" bold="true" box="[972,1006,365,386]" pageId="5" pageNumber="376">µm</emphasis>
</td>
<td id="F140F063C02B4AEC43083F55EA31FFA3" box="[1046,1100,365,386]" gridcol="9" gridrow="2" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643393F55EA46FFA3" bold="true" box="[1063,1083,365,386]" pageId="5" pageNumber="376">sc</emphasis>
</td>
<td id="F140F063C02B4AEC436C3F55EAE9FFA3" box="[1138,1172,365,386]" gridcol="10" gridrow="2" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB516436C3F55EAE9FFA3" bold="true" box="[1138,1172,365,386]" pageId="5" pageNumber="376">µm</emphasis>
</td>
<td id="F140F063C02B4AEC43DF3F55EA96FFA3" box="[1217,1259,365,386]" gridcol="11" gridrow="2" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643D23F55EA9DFFA3" bold="true" box="[1228,1248,365,386]" pageId="5" pageNumber="376">sc</emphasis>
</td>
<td id="F140F063C02B4AEC420C3F55EB40FFA3" box="[1298,1341,365,386]" gridcol="12" gridrow="2" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51642093F55EB44FFA3" bold="true" box="[1303,1337,365,386]" pageId="5" pageNumber="376">µm</emphasis>
</td>
<td id="F140F063C02B4AEC427F3F55EBEAFFA3" box="[1377,1431,365,386]" gridcol="13" gridrow="2" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB516426C3F55EBFBFFA3" bold="true" box="[1394,1414,365,386]" pageId="5" pageNumber="376">sc</emphasis>
</td>
</tr>
<tr id="B291991FC02B4AEC479E3F9BEBEAFF98" box="[128,1431,419,441]" gridrow="3" pageId="5" pageNumber="376">
<th id="F140F063C02B4AEC479E3F9BEFE9FF98" box="[128,404,419,441]" gridcol="0" gridrow="3" pageId="5" pageNumber="376">Body length</th>
<td id="F140F063C02B4AEC46AD3F9BEFB6FF98" box="[435,459,419,441]" gridcol="1" gridrow="3" pageId="5" pageNumber="376">15</td>
<td id="F140F063C02B4AEC46F53F9BEC6AFF98" box="[491,535,419,441]" gridcol="2" gridrow="3" pageId="5" pageNumber="376">76</td>
<td id="F140F063C02B4AEC45333F9BEC32FF98" box="[557,591,419,441]" gridcol="3" gridrow="3" pageId="5" pageNumber="376">–</td>
<td id="F140F063C02B4AEC457B3F9BECECFF98" box="[613,657,419,441]" gridcol="4" gridrow="3" pageId="5" pageNumber="376">95</td>
<td id="F140F063C02B4AEC45B93F9BEC97FF98" box="[679,746,419,441]" gridcol="5" gridrow="3" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51645A33F9BEC9CFF99" box="[701,737,419,440]" italics="true" pageId="5" pageNumber="376">543</emphasis>
</td>
<td id="F140F063C02B4AEC443F3F9BED48FF98" box="[801,821,419,441]" gridcol="6" gridrow="3" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB516443C3F9CED53FF98" box="[802,814,420,441]" italics="true" pageId="5" pageNumber="376">–</emphasis>
</td>
<td id="F140F063C02B4AEC44773F9BEDE2FF98" box="[873,927,419,441]" gridcol="7" gridrow="3" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB516446C3F9BEDEBFF99" box="[882,918,419,440]" italics="true" pageId="5" pageNumber="376">648</emphasis>
</td>
<td id="F140F063C02B4AEC44D93F9BED8EFF98" box="[967,1011,419,441]" gridcol="8" gridrow="3" pageId="5" pageNumber="376">84</td>
<td id="F140F063C02B4AEC43083F9BEA31FF98" box="[1046,1100,419,441]" gridcol="9" gridrow="3" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643013F9BEA3EFF99" box="[1055,1091,419,440]" italics="true" pageId="5" pageNumber="376">596</emphasis>
</td>
<td id="F140F063C02B4AEC436C3F9BEAE9FF98" box="[1138,1172,419,441]" gridcol="10" gridrow="3" pageId="5" pageNumber="376">7</td>
<td id="F140F063C02B4AEC43DF3F9BEA96FF98" box="[1217,1259,419,441]" gridcol="11" gridrow="3" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643D43F9BEA9FFF99" box="[1226,1250,419,440]" italics="true" pageId="5" pageNumber="376">39</emphasis>
</td>
<td id="F140F063C02B4AEC420C3F9BEB40FF98" box="[1298,1341,419,441]" gridcol="12" gridrow="3" pageId="5" pageNumber="376">94</td>
<td id="F140F063C02B4AEC427F3F9BEBEAFF98" box="[1377,1431,419,441]" gridcol="13" gridrow="3" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51642743F9BEBF3FF99" box="[1386,1422,419,440]" italics="true" pageId="5" pageNumber="376">648</emphasis>
</td>
</tr>
<tr id="B291991FC02B4AEC479E3FEAEBEAFFC9" box="[128,1431,466,488]" gridrow="4" pageId="5" pageNumber="376" rowspan-5="1" rowspan-7="1">
<th id="F140F063C02B4AEC479E3FEAEFE9FFC9" box="[128,404,466,488]" gridcol="0" gridrow="4" pageId="5" pageNumber="376">Scapular plate length</th>
<td id="F140F063C02B4AEC46AD3FEAEFB6FFC9" box="[435,459,466,488]" gridcol="1" gridrow="4" pageId="5" pageNumber="376">14</td>
<td id="F140F063C02B4AEC46F53FEAEC6AFFC9" box="[491,535,466,488]" gridcol="2" gridrow="4" pageId="5" pageNumber="376">12.5</td>
<td id="F140F063C02B4AEC45333FEAEC32FFC9" box="[557,591,466,488]" gridcol="3" gridrow="4" pageId="5" pageNumber="376">–</td>
<td id="F140F063C02B4AEC457B3FEAECECFFC9" box="[613,657,466,488]" gridcol="4" gridrow="4" pageId="5" pageNumber="376">15.5</td>
<td id="F140F063C02B4AEC443F3FEAED48FFC9" box="[801,821,466,488]" gridcol="6" gridrow="4" pageId="5" pageNumber="376">–</td>
<td id="F140F063C02B4AEC44D93FEAED8EFFC9" box="[967,1011,466,488]" gridcol="8" gridrow="4" pageId="5" pageNumber="376">14.2</td>
<td id="F140F063C02B4AEC43083FEAEA31FFC9" box="[1046,1100,466,488]" gridcol="9" gridrow="4" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643353FEBEA4AFFC9" box="[1067,1079,467,488]" italics="true" pageId="5" pageNumber="376">–</emphasis>
</td>
<td id="F140F063C02B4AEC436C3FEAEAE9FFC9" box="[1138,1172,466,488]" gridcol="10" gridrow="4" pageId="5" pageNumber="376">0.8</td>
<td id="F140F063C02B4AEC43DF3FEAEA96FFC9" box="[1217,1259,466,488]" gridcol="11" gridrow="4" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643CE3FEBEAA1FFC9" box="[1232,1244,467,488]" italics="true" pageId="5" pageNumber="376">–</emphasis>
</td>
<td id="F140F063C02B4AEC420C3FEAEB40FFC9" box="[1298,1341,466,488]" gridcol="12" gridrow="4" pageId="5" pageNumber="376">14.5</td>
<td id="F140F063C02B4AEC427F3FEAEBEAFFC9" box="[1377,1431,466,488]" gridcol="13" gridrow="4" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51642683FEBEBFFFFC9" box="[1398,1410,467,488]" italics="true" pageId="5" pageNumber="376">–</emphasis>
</td>
</tr>
<tr id="B291991FC02B4AEC479E3C39EBEAFCF7" box="[128,1431,513,726]" gridrow="5" pageId="5" pageNumber="376">
<th id="F140F063C02B4AEC479E3C39EFE9FCF7" box="[128,404,513,726]" gridcol="0" gridrow="5" pageId="5" pageNumber="376">
Head appendages lengths Cirrus internus Cephalic papilla Cirrus externus Clava Cirrus 
<emphasis id="3ED5474FC02BB51647F43C98EE84FC94" box="[234,249,672,693]" italics="true" pageId="5" pageNumber="376">A</emphasis>
Cirrus 
<emphasis id="3ED5474FC02BB51647C73CF8EE95FCF4" box="[217,232,704,725]" italics="true" pageId="5" pageNumber="376">A</emphasis>
/ body length ratio
</th>
<td id="F140F063C02B4AEC46AD3C39EFB6FCF7" box="[435,459,513,726]" gridcol="1" gridrow="5" pageId="5" pageNumber="376">13 10 13 13 12 12</td>
<td id="F140F063C02B4AEC46F53C39EC6AFCF7" box="[491,535,513,726]" gridcol="2" gridrow="5" pageId="5" pageNumber="376">3.2 2.0 5.4 1.6 20.9 27%</td>
<td id="F140F063C02B4AEC45333C39EC32FCF7" box="[557,591,513,726]" gridcol="3" gridrow="5" pageId="5" pageNumber="376">– – – – – –</td>
<td id="F140F063C02B4AEC457B3C39ECECFCF7" box="[613,657,513,726]" gridcol="4" gridrow="5" pageId="5" pageNumber="376">4.7 3.1 8.1 2.6 30.8 36%</td>
<td id="F140F063C02B4AEC45B93C39EC97FCF7" box="[679,746,513,726]" gridcol="5" gridrow="5" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51645A43C19EC97FC94" italics="true" pageId="5" pageNumber="376">21.3 14.8 34.8 11.9 154.8</emphasis>
</td>
<td id="F140F063C02B4AEC443F3C39ED48FCF7" box="[801,821,513,726]" gridcol="6" gridrow="5" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB516443C3C1AED53FCF4" box="[802,814,546,725]" italics="true" pageId="5" pageNumber="376">– – – – – –</emphasis>
</td>
<td id="F140F063C02B4AEC44773C39EDE2FCF7" box="[873,927,513,726]" gridcol="7" gridrow="5" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51644713C19EDE2FC94" italics="true" pageId="5" pageNumber="376">33.1 24.8 60.4 18.4 212.4</emphasis>
</td>
<td id="F140F063C02B4AEC44D93C39ED8EFCF7" box="[967,1011,513,726]" gridcol="8" gridrow="5" pageId="5" pageNumber="376">3.9 2.6 6.5 2.2 26.8 32%</td>
<td id="F140F063C02B4AEC43083C39EA31FCF7" box="[1046,1100,513,726]" gridcol="9" gridrow="5" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643023C19EA4AFCF4" italics="true" pageId="5" pageNumber="376">27.6 18.3 46.0 15.8 189.1 –</emphasis>
</td>
<td id="F140F063C02B4AEC436C3C39EAE9FCF7" box="[1138,1172,513,726]" gridcol="10" gridrow="5" pageId="5" pageNumber="376">0.5 0.3 0.9 0.3 2.8 2%</td>
<td id="F140F063C02B4AEC43DF3C39EA96FCF7" box="[1217,1259,513,726]" gridcol="11" gridrow="5" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643D93C19EAA1FCF4" italics="true" pageId="5" pageNumber="376">4.0 2.8 7.3 2.1 17.4 –</emphasis>
</td>
<td id="F140F063C02B4AEC420C3C39EB40FCF7" box="[1298,1341,513,726]" gridcol="12" gridrow="5" pageId="5" pageNumber="376">3.4 2.7 7.5 2.6 30.8 33%</td>
<td id="F140F063C02B4AEC427F3C39EBEAFCF7" box="[1377,1431,513,726]" gridcol="13" gridrow="5" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51642793C19EBECFC17" box="[1383,1425,545,566]" italics="true" pageId="5" pageNumber="376">23.4</emphasis>
18.6 
<emphasis id="3ED5474FC02BB51642793C59EBFFFCF4" italics="true" pageId="5" pageNumber="376">51.7 17.9 212.4 –</emphasis>
</td>
</tr>
<tr id="B291991FC02B4AEC479E3CD7EBEAFDA5" box="[128,1431,751,900]" gridrow="6" pageId="5" pageNumber="376">
<th id="F140F063C02B4AEC479E3CD7EFE9FDA5" box="[128,404,751,900]" gridcol="0" gridrow="6" pageId="5" pageNumber="376">Claw branch lengths Claw 1 Claw 2 Claw 3 Claw 4</th>
<td id="F140F063C02B4AEC46AD3CD7EFB6FDA5" box="[435,459,751,900]" gridcol="1" gridrow="6" pageId="5" pageNumber="376">15 15 15 15</td>
<td id="F140F063C02B4AEC46F53CD7EC6AFDA5" box="[491,535,751,900]" gridcol="2" gridrow="6" pageId="5" pageNumber="376">3.7 3.3 3.4 3.7</td>
<td id="F140F063C02B4AEC45333CD7EC32FDA5" box="[557,591,751,900]" gridcol="3" gridrow="6" pageId="5" pageNumber="376">– – – –</td>
<td id="F140F063C02B4AEC457B3CD7ECECFDA5" box="[613,657,751,900]" gridcol="4" gridrow="6" pageId="5" pageNumber="376">5.5 4.8 5.0 5.3</td>
<td id="F140F063C02B4AEC45B93CD7EC97FDA5" box="[679,746,751,900]" gridcol="5" gridrow="6" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51645A43D37EC99FDA2" box="[698,740,783,899]" italics="true" pageId="5" pageNumber="376">27.4 25.7 25.0 26.7</emphasis>
</td>
<td id="F140F063C02B4AEC443F3CD7ED48FDA5" box="[801,821,751,900]" gridcol="6" gridrow="6" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB516443C3D37ED53FDA5" box="[802,814,783,900]" italics="true" pageId="5" pageNumber="376">– – – –</emphasis>
</td>
<td id="F140F063C02B4AEC44773CD7EDE2FDA5" box="[873,927,751,900]" gridcol="7" gridrow="6" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51644713D36EDE4FDA2" box="[879,921,782,899]" italics="true" pageId="5" pageNumber="376">36.6 31.5 35.2 37.1</emphasis>
</td>
<td id="F140F063C02B4AEC44D93CD7ED8EFDA5" box="[967,1011,751,900]" gridcol="8" gridrow="6" pageId="5" pageNumber="376">4.4 4.1 4.2 4.4</td>
<td id="F140F063C02B4AEC43083CD7EA31FDA5" box="[1046,1100,751,900]" gridcol="9" gridrow="6" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643023D37EA3BFDA2" box="[1052,1094,783,899]" italics="true" pageId="5" pageNumber="376">31.3 28.4 29.2 31.4</emphasis>
</td>
<td id="F140F063C02B4AEC436C3CD7EAE9FDA5" box="[1138,1172,751,900]" gridcol="10" gridrow="6" pageId="5" pageNumber="376">0.5 0.4 0.4 0.5</td>
<td id="F140F063C02B4AEC43DF3CD7EA96FDA5" box="[1217,1259,751,900]" gridcol="11" gridrow="6" pageId="5" pageNumber="376">
<emphasis id="3ED5474FC02BB51643D93D37EA98FDA2" box="[1223,1253,783,899]" italics="true" pageId="5" pageNumber="376">3.2 2.0 2.7 2.9</emphasis>
</td>
<td id="F140F063C02B4AEC420C3CD7EB40FDA5" box="[1298,1341,751,900]" gridcol="12" gridrow="6" pageId="5" pageNumber="376">4.4 4.5 4.5 4.9</td>
<td id="F140F063C02B4AEC427F3CD7EBEAFDA5" box="[1377,1431,751,900]" gridcol="13" gridrow="6" pageId="5" pageNumber="376">30.3 31.0 31.0 33.8</td>
</tr>
</table>
</paragraph>
<caption id="58DECBD5C02BB516476F398DEA05F9C9" ID-DOI="http://doi.org/10.5281/zenodo.5358951" ID-Zenodo-Dep="5358951" httpUri="https://zenodo.org/record/5358951/files/figure.png" pageId="5" pageNumber="376" startId="5.[113,148,1973,1995]" targetBox="[113,1451,978,1958]" targetPageId="5">
<paragraph id="0C1E9B5DC02BB516476F398DEA05F9C9" blockId="5.[113,1451,1973,2024]" pageId="5" pageNumber="376">
Fig. 4. Close-up on various parts of dorsal armature of 
<taxonomicName id="CBA1E0DEC02BB516458E398DED14F9EA" authority="Gąsiorek, 2018" authorityName="Gąsiorek" authorityYear="2018" box="[656,873,1973,1995]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="376" phylum="Tardigrada" rank="species" species="arenosus" status="sp. nov.">
<emphasis id="3ED5474FC02BB516458E398DED14F9EA" box="[656,873,1973,1995]" italics="true" pageId="5" pageNumber="376">Bryodelphax arenosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="25E6FA34C02BB516446E398EEDC3F9EA" box="[880,958,1974,1995]" pageId="5" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
A, faceting of sc and ornamented pattern of sl; B, ornamented pattern of pl; C, faceting of c. Abbreviations are the same as for Figure 1. Scale bars in μm.
</paragraph>
</caption>
<caption id="58DECBD5C028B51547993E85EDD8FEF2" box="[135,933,189,211]" pageId="6" pageNumber="377" startId="6.[135,190,189,211]">
<paragraph id="0C1E9B5DC028B51547993E85EDD8FEF2" blockId="6.[135,933,189,211]" box="[135,933,189,211]" pageId="6" pageNumber="377">
Table 3. Uncorrected pairwise distance (in %) for available 
<taxonomicName id="CBA1E0DEC028B51545C83E85EDD8FEF2" box="[726,933,189,211]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="377" phylum="Tardigrada" rank="species" species="undetermined">
<emphasis id="3ED5474FC028B51545C83E85ED2CFEF3" box="[726,849,189,210]" italics="true" pageId="6" pageNumber="377">Bryodelphax</emphasis>
species.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="0C1E9B5DC028B51545C43F3DEB05FFFE" pageId="6" pageNumber="377">
<table id="7EA169FDC0284AEC47883F3DEB05FFC1" box="[150,1400,261,480]" gridcols="6" gridrows="7" pageId="6" pageNumber="377">
<tr id="B291991FC0284AEC47883F3DEB05FF3A" box="[150,1400,261,283]" gridrow="0" pageId="6" pageNumber="377" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<th id="F140F063C0284AEC45C43F3DED3AFF3A" box="[730,839,261,283]" gridcol="2" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="3ED5474FC028B51545C43F3DED3AFF3B" bold="true" box="[730,839,261,283]" pageId="6" pageNumber="377">18S rRNA</emphasis>
</th>
<th id="F140F063C0284AEC43603F3DEB05FF3A" box="[1150,1400,261,283]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="3ED5474FC028B51543DB3F3DEB4CFF3B" bold="true" box="[1221,1329,261,283]" pageId="6" pageNumber="377">28S rRNA</emphasis>
</th>
</tr>
<tr id="B291991FC0284AEC47883F24EB05FF13" box="[150,1400,284,306]" gridrow="1" pageId="6" pageNumber="377" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="F140F063C0284AEC47883F24EFDDFF13" box="[150,416,284,306]" gridcol="0" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="3ED5474FC028B51547883F24EE9CFF13" bold="true" box="[150,225,284,306]" pageId="6" pageNumber="377">Species</emphasis>
</th>
</tr>
<tr id="B291991FC0284AEC47883F0CEB05FF6B" box="[150,1400,308,330]" gridrow="2" pageId="6" pageNumber="377" rowspan-0="1">
<td id="F140F063C0284AEC452F3F0CEC1AFF6B" box="[561,615,308,330]" gridcol="1" gridrow="2" pageId="6" pageNumber="377">(1)</td>
<td id="F140F063C0284AEC45C43F0CED3AFF6B" box="[730,839,308,330]" gridcol="2" gridrow="2" pageId="6" pageNumber="377">(2)</td>
<td id="F140F063C0284AEC44A73F0CED92FF6B" box="[953,1007,308,330]" gridcol="3" gridrow="2" pageId="6" pageNumber="377">(3)</td>
<td id="F140F063C0284AEC43603F0CEAC9FF6B" box="[1150,1204,308,330]" gridcol="4" gridrow="2" pageId="6" pageNumber="377">(2)</td>
<td id="F140F063C0284AEC425C3F0CEB05FF6B" box="[1346,1400,308,330]" gridcol="5" gridrow="2" pageId="6" pageNumber="377">(3)</td>
</tr>
<tr id="B291991FC0284AEC47883F52EB05FFA1" box="[150,1400,362,384]" gridrow="3" pageId="6" pageNumber="377">
<th id="F140F063C0284AEC47883F52EB05FFA1" box="[150,1400,362,384]" colspan="6" colspanRight="5" gridcol="0" gridrow="3" pageId="6" pageNumber="377">
(1) 
<taxonomicName id="CBA1E0DEC028B51547A43F52EFECFF5E" authorityName="Thulin" authorityYear="1928" box="[186,401,362,383]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="377" phylum="Tardigrada" rank="species" species="parvulus">
<emphasis id="3ED5474FC028B51547A43F52EFECFF5E" box="[186,401,362,383]" italics="true" pageId="6" pageNumber="377">Bryodelphax parvulus</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="B291991FC0284AEC47883FB2EB05FF81" box="[150,1400,394,416]" gridrow="4" pageId="6" pageNumber="377" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="F140F063C0284AEC47883FB2EFDDFF81" box="[150,416,394,416]" gridcol="0" gridrow="4" pageId="6" pageNumber="377">
(2) 
<taxonomicName id="CBA1E0DEC028B51547A43FB2EFDDFFBE" box="[186,416,394,415]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="377" phylum="Tardigrada" rank="species" species="maculatus">
<emphasis id="3ED5474FC028B51547A43FB2EFDDFFBE" box="[186,416,394,415]" italics="true" pageId="6" pageNumber="377">Bryodelphax maculatus</emphasis>
</taxonomicName>
</th>
<td id="F140F063C0284AEC452F3FB2EC1AFF81" box="[561,615,394,416]" gridcol="1" gridrow="4" pageId="6" pageNumber="377">0.008</td>
</tr>
<tr id="B291991FC0284AEC47883F92EB05FFE1" box="[150,1400,426,448]" gridrow="5" pageId="6" pageNumber="377" rowspan-3="1" rowspan-5="1">
<th id="F140F063C0284AEC47883F92EFDDFFE1" box="[150,416,426,448]" gridcol="0" gridrow="5" pageId="6" pageNumber="377">
(3) 
<taxonomicName id="CBA1E0DEC028B51547A43F92EFE9FF9E" authorityName="Gąsiorek" authorityYear="2018" box="[186,404,426,447]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="377" phylum="Tardigrada" rank="species" species="arenosus">
<emphasis id="3ED5474FC028B51547A43F92EFE9FF9E" box="[186,404,426,447]" italics="true" pageId="6" pageNumber="377">Bryodelphax arenosus</emphasis>
</taxonomicName>
</th>
<td id="F140F063C0284AEC452F3F92EC1AFFE1" box="[561,615,426,448]" gridcol="1" gridrow="5" pageId="6" pageNumber="377">0.028</td>
<td id="F140F063C0284AEC45C43F92ED3AFFE1" box="[730,839,426,448]" gridcol="2" gridrow="5" pageId="6" pageNumber="377">0.021</td>
<td id="F140F063C0284AEC43603F92EAC9FFE1" box="[1150,1204,426,448]" gridcol="4" gridrow="5" pageId="6" pageNumber="377">0.193</td>
</tr>
<tr id="B291991FC0284AEC47883FF1EB05FFC1" box="[150,1400,457,480]" gridrow="6" pageId="6" pageNumber="377">
<th id="F140F063C0284AEC47883FF1EFDDFFC1" box="[150,416,457,480]" gridcol="0" gridrow="6" pageId="6" pageNumber="377">
(4) 
<taxonomicName id="CBA1E0DEC028B51547A43FF1EFE8FFFF" authorityName="Gasiorek &amp; Degma" authorityYear="2018" box="[186,405,457,478]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="377" phylum="Tardigrada" rank="species" species="instabilis">
<emphasis id="3ED5474FC028B51547A43FF1EFE8FFFF" box="[186,405,457,478]" italics="true" pageId="6" pageNumber="377">Bryodelphax instabilis</emphasis>
</taxonomicName>
</th>
<td id="F140F063C0284AEC452F3FF1EC1AFFC1" box="[561,615,457,480]" gridcol="1" gridrow="6" pageId="6" pageNumber="377">0.011</td>
<td id="F140F063C0284AEC45C43FF1ED3AFFC1" box="[730,839,457,480]" gridcol="2" gridrow="6" pageId="6" pageNumber="377">0.005</td>
<td id="F140F063C0284AEC44A73FF1ED92FFC1" box="[953,1007,457,480]" gridcol="3" gridrow="6" pageId="6" pageNumber="377">0.025</td>
<td id="F140F063C0284AEC43603FF1EAC9FFC1" box="[1150,1204,457,480]" gridcol="4" gridrow="6" pageId="6" pageNumber="377">0.075</td>
<td id="F140F063C0284AEC425C3FF1EB05FFC1" box="[1346,1400,457,480]" gridcol="5" gridrow="6" pageId="6" pageNumber="377">0.155</td>
</tr>
</table>
</paragraph>
<subSubSection id="44BBC8D6C028B51547963CB6EC88FCC7" pageId="6" pageNumber="377" type="etymology">
<paragraph id="0C1E9B5DC028B51547963CB6EC88FCC7" blockId="6.[136,776,654,742]" pageId="6" pageNumber="377">
<emphasis id="3ED5474FC028B51547963CB6EF75FC87" bold="true" box="[136,264,654,678]" pageId="6" pageNumber="377">Etymology.</emphasis>
From Latin 
<emphasis id="3ED5474FC028B515469B3CB6EC7CFC87" box="[389,513,654,678]" italics="true" pageId="6" pageNumber="377">
(h) 
<taxonomicName id="CBA1E0DEC028B51546BF3CB6EC7CFC87" authorityName="Gąsiorek" authorityYear="2018" box="[417,513,654,678]" class="Heterotardigrada" family="Echiniscidae" genus="Bryodelphax" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="377" phylum="Tardigrada" rank="species" species="arenosus">arenosus</taxonomicName>
</emphasis>
= sandy, inhabiting sand. The new species was found in coastal lowland rainforest with peculiarly sandy soil (arenosols), atypical for this biome.
</paragraph>
</subSubSection>
</treatment>
</document>