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<document id="9EA03136616465E94E6FE3254691B664" ID-CLB-Dataset="300002" ID-DOI="10.1186/s13358-023-00290-6" ID-GBIF-Dataset="68b40265-75e3-40b4-a504-7df2829f9964" ID-ISSN="1664-2384" ID-Zenodo-Dep="12001012" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1718653334918" checkinUser="felipe" docAuthor="Weis, Robert, Schweigert, GÜnter &amp; Wittische, Julian" docDate="2023" docId="03F1AE26FF88742DFF08FF38FDA00F99" docLanguage="en" docName="SwissJPalaeontol.142.24.pdf" docOrigin="Swiss Journal of Palaeontology (24) 142 (1)" docSource="http://dx.doi.org/10.1186/s13358-023-00290-6" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Cenoceras rumelangense Weis &amp; Schweigert &amp; Wittische 2023, n. sp." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FFC8D65EFF8C742BFF9FFFD3FFEA0B79" masterDocTitle="A new giant nautilid species from the Middle Jurassic of Luxembourg and Southwest Germany" masterLastPageNumber="11" masterPageNumber="1" pageNumber="5" updateTime="1721315103231" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0206E62A56CC93F7C9D99CAB5D07B398">A new giant nautilid species from the Middle Jurassic of Luxembourg and Southwest Germany</mods:title>
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<mods:namePart id="EFB0E2A7FF893F2D3D1D38103A5DDB77">Weis, Robert</mods:namePart>
<mods:affiliation id="BCB46C0D2F3471784E0B5A9EFEF6B2B0">Musée National d’histoire Naturelle, 25 rue MÜnster, 2160 Luxembourg, Grand Duchy of Luxembourg</mods:affiliation>
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<mods:namePart id="0F1004AFBA5280880AABDDCFF815460E">Schweigert, GÜnter</mods:namePart>
<mods:affiliation id="6AF294AA0A7D2105583F7C1F48B860A3">Staatliches Museum fÜr Naturkunde Stuttgart, Rosenstein 1,</mods:affiliation>
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<mods:namePart id="1528E4D0787368CFF39DED5BD5F6FAC4">Wittische, Julian</mods:namePart>
<mods:affiliation id="E1BE15C1C653972AFC6CD7A77CACBD30">Musée National d’histoire Naturelle, 25 rue MÜnster, 2160 Luxembourg, Grand Duchy of Luxembourg</mods:affiliation>
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<mods:date id="5AAC6074ABB5F6871CF16D6636725584">2023</mods:date>
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<treatment id="03F1AE26FF88742DFF08FF38FDA00F99" ID-DOI="http://doi.org/10.5281/zenodo.12773179" ID-GBIF-Taxon="235609000" ID-Zenodo-Dep="12773179" LSID="urn:lsid:plazi:treatment:03F1AE26FF88742DFF08FF38FDA00F99" httpUri="http://treatment.plazi.org/id/03F1AE26FF88742DFF08FF38FDA00F99" lastPageId="6" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="C3424CBBFF88742FFF08FF38FD360A7A" box="[151,732,235,259]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BE71F30FF88742FFF08FF38FD360A7A" blockId="4.[151,775,235,963]" box="[151,732,235,259]" pageId="4" pageNumber="5">
<heading id="D0AFA85CFF88742FFF08FF38FD360A7A" box="[151,732,235,259]" fontSize="9" level="2" pageId="4" pageNumber="5" reason="8">
<emphasis id="B92CC322FF88742FFF08FF38FE1C0A7A" bold="true" box="[151,502,235,259]" pageId="4" pageNumber="5">
<taxonomicName id="4C5864B3FF88742FFF08FF38FE590A7A" authority="Weis &amp; Schweigert &amp; Wittische, 2023" authorityName="Weis &amp; Schweigert &amp; Wittische" authorityYear="2023" box="[151,435,235,259]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="rumelangense" status="sp. nov.">
<emphasis id="B92CC322FF88742FFF08FF38FE590A7A" bold="true" box="[151,435,235,259]" italics="true" pageId="4" pageNumber="5">Cenoceras rumelangense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7E59FF88742FFE26FF38FE1C0A7A" box="[441,502,235,259]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
[Weis &amp; Schweigert]
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3424CBBFF88742FFF08FED8FEB20A5A" box="[151,344,267,291]" pageId="4" pageNumber="5" type="description">
<paragraph id="8BE71F30FF88742FFF08FED8FEB20A5A" blockId="4.[151,775,235,963]" box="[151,344,267,291]" pageId="4" pageNumber="5">
<figureCitation id="136303B5FF88742FFF08FED8FF100A5A" box="[151,250,267,291]" captionStart="Fig" captionStartId="4.[167,198,1848,1868]" captionTargetBox="[176,751,1099,1836]" captionTargetId="figure-400@4.[176,751,1099,1836]" captionTargetPageId="4" captionText="Fig. 5 Cenoceras rumelangense n.sp., holoytpe (specimen MNHNL BM790),‘Marnes sableuses d’Audun-le-Tiche’, Humphriesianum Zone, Berenskaul,Rumelange, Luxembourg" figureDoi="http://doi.org/10.5281/zenodo.12001084" httpUri="https://zenodo.org/record/12001084/files/figure.png" pageId="4" pageNumber="5">Figures 5</figureCitation>
,
<figureCitation id="136303B5FF88742FFE99FED8FEF90A5A" box="[262,275,267,291]" captionStart="Fig" captionStartId="4.[829,860,957,977]" captionTargetBox="[837,1412,236,944]" captionTargetId="figure-420@4.[837,1412,236,944]" captionTargetPageId="4" captionText="Fig. 6 Cenoceras rumelangense n.sp., paratype (specimen MNHNL BM383),‘Marnes sableuses d’Audun-le-Tiche’, Humphriesianum Zone, Rumelange,Luxembourg" figureDoi="http://doi.org/10.5281/zenodo.12001070" httpUri="https://zenodo.org/record/12001070/files/figure.png" pageId="4" pageNumber="5">6</figureCitation>
,
<figureCitation id="136303B5FF88742FFE80FEDFFEC60A5A" box="[287,300,268,291]" captionStart="Fig" captionStartId="5.[167,198,1042,1062]" captionTargetBox="[176,751,236,1029]" captionTargetId="figure-388@5.[176,751,236,1029]" captionTargetPageId="5" captionText="Fig. 7 Cenoceras rumelangense n.sp., paratype, inner whorl of a bigger specimen (specimen MNHNL BM783),‘Marnes sableuses d’Audun-le-Tiche’,Humphriesianum Zone,Berenskaul, Rumelange, Luxembourg" figureDoi="http://doi.org/10.5281/zenodo.12001123" httpUri="https://zenodo.org/record/12001123/files/figure.png" pageId="4" pageNumber="5">7</figureCitation>
,
<figureCitation id="136303B5FF88742FFEA7FED8FEAF0A5A" box="[312,325,267,291]" captionStart="Fig" captionStartId="5.[829,860,996,1016]" captionTargetBox="[837,1412,236,983]" captionTargetId="figure-414@5.[837,1412,236,983]" captionTargetPageId="5" captionText="Fig. 8 Cenoceras rumelangense n.sp., paratype,largest measured specimen (specimen MNHNL BM791),‘Marnes sableuses d’Audun-le-Tiche’, Humphriesianum Zone, Berenskaul, Rumelange, Luxembourg" figureDoi="http://doi.org/10.5281/zenodo.12001111" httpUri="https://zenodo.org/record/12001111/files/figure.png" pageId="4" pageNumber="5">8</figureCitation>
,
<figureCitation id="136303B5FF88742FFED4FED8FEB20A5A" box="[331,344,267,291]" captionStart="Fig" captionStartId="6.[167,198,1051,1071]" captionTargetBox="[176,751,236,1039]" captionTargetId="figure-644@6.[176,751,236,1039]" captionTargetPageId="6" captionText="Fig. 9 Cenoceras rumelangense n.sp., paratype (specimen SMNS 62245), Ostreenkalk Formation, Humphriesianum Zone, Gruibingen, Southwest Germany" figureDoi="http://doi.org/10.5281/zenodo.12001131" httpUri="https://zenodo.org/record/12001131/files/figure.png" pageId="4" pageNumber="5">9</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3424CBBFF88742FFF08FEF8FD4F097A" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="8BE71F30FF88742FFF08FEF8FDD00A3A" blockId="4.[151,775,235,963]" box="[151,570,299,323]" pageId="4" pageNumber="5">
1995
<taxonomicName id="4C5864B3FF88742FFF4DFEFFFE8D0A3A" box="[210,359,300,323]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B92CC322FF88742FFF4DFEFFFEAA0A3A" box="[210,320,300,323]" italics="true" pageId="4" pageNumber="5">Cenoceras</emphasis>
sp.
</taxonomicName>
: Dietl, p. 357, fig. 1.
</paragraph>
<paragraph id="8BE71F30FF88742FFF08FE98FE120A1A" blockId="4.[151,775,235,963]" box="[151,504,331,355]" pageId="4" pageNumber="5">
2005
<taxonomicName id="4C5864B3FF88742FFF4DFE9FFE8D0A1A" box="[210,359,332,355]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B92CC322FF88742FFF4DFE9FFEAA0A1A" box="[210,320,332,355]" italics="true" pageId="4" pageNumber="5">Cenoceras</emphasis>
sp.
</taxonomicName>
: Gross, p. 58.
</paragraph>
<paragraph id="8BE71F30FF88742FFF08FEB8FE690ADA" blockId="4.[151,775,235,963]" pageId="4" pageNumber="5">
? 2008
<taxonomicName id="4C5864B3FF88742FFF78FEBFFEBF0AFA" authorityName="Hyatt" authorityYear="1883" box="[231,341,364,387]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
<emphasis id="B92CC322FF88742FFF78FEBFFEBF0AFA" box="[231,341,364,387]" italics="true" pageId="4" pageNumber="5">Cenoceras</emphasis>
</taxonomicName>
gr.
<taxonomicName id="4C5864B3FF88742FFE1DFEB8FE3E0AFA" baseAuthorityName="Sowerby" baseAuthorityYear="1816" box="[386,468,363,387]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="obesum">
<emphasis id="B92CC322FF88742FFE1DFEB8FE3E0AFA" box="[386,468,363,387]" italics="true" pageId="4" pageNumber="5">obesum</emphasis>
</taxonomicName>
: Rulleau, pl. 22, fig. 2 [only specimen EPF Baj74].
</paragraph>
<paragraph id="8BE71F30FF88742FFF08FE78FDD10ABA" blockId="4.[151,775,235,963]" box="[151,571,427,451]" pageId="4" pageNumber="5">
2013
<taxonomicName id="4C5864B3FF88742FFF4DFE7FFE8D0ABA" box="[210,359,428,451]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B92CC322FF88742FFF4DFE7FFEAA0ABA" box="[210,320,428,451]" italics="true" pageId="4" pageNumber="5">Cenoceras</emphasis>
sp.
</taxonomicName>
: Dietl, p. 20, fig. 24.
</paragraph>
<paragraph id="8BE71F30FF88742FFF08FE18FD540A9A" blockId="4.[151,775,235,963]" box="[151,702,459,483]" pageId="4" pageNumber="5">
? 2016
<taxonomicName id="4C5864B3FF88742FFF7DFE1FFE430A9A" box="[226,425,460,483]" class="Cephalopoda" family="Nautilidae" genus="Metacenoceras" isUncertain="true" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B92CC322FF88742FFF7DFE1FFE690A9A" box="[226,387,460,483]" italics="true" pageId="4" pageNumber="5">Metacenoceras</emphasis>
sp.
</taxonomicName>
: Grulke, p. 139, figs. 1&amp;2.
</paragraph>
<paragraph id="8BE71F30FF88742FFF08FE38FD4F097A" blockId="4.[151,775,235,963]" box="[151,677,491,515]" pageId="4" pageNumber="5">
2021
<taxonomicName id="4C5864B3FF88742FFF4DFE3FFE72097A" baseAuthorityName="Sowerby" baseAuthorityYear="1816" box="[210,408,491,515]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="obesum">
<emphasis id="B92CC322FF88742FFF4DFE3FFE72097A" box="[210,408,491,515]" italics="true" pageId="4" pageNumber="5">Cenoceras obesum</emphasis>
</taxonomicName>
: Schweigert, p. 9, fig. 14.
</paragraph>
</subSubSection>
<subSubSection id="C3424CBBFF88742FFF08FDD8FDFC09DA" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BE71F30FF88742FFF08FDD8FDFC09DA" blockId="4.[151,775,235,963]" pageId="4" pageNumber="5">
<emphasis id="B92CC322FF88742FFF08FDD8FE6D095A" bold="true" box="[151,391,523,547]" pageId="4" pageNumber="5">Derivation of name</emphasis>
: The species is named after the town of Rumelange, a municipality in the southernmost part of
<collectingCountry id="F34F5FA0FF88742FFF6EFD98FE6A091A" box="[241,384,587,611]" name="Luxembourg" pageId="4" pageNumber="5">Luxembourg</collectingCountry>
that is widely known for its rich fossil findings from Toarcian to Bajocian rocks, including the herein described nautilids.
</paragraph>
</subSubSection>
<subSubSection id="C3424CBBFF88742FFF08FD78FAD60E7C" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BE71F30FF88742FFF08FD78FF01099A" blockId="4.[151,775,235,963]" pageId="4" pageNumber="5">
<emphasis id="B92CC322FF88742FFF08FD78FEEC09BA" bold="true" box="[151,262,683,707]" pageId="4" pageNumber="5">
<typeStatus id="54E3A192FF88742FFF08FD78FEEC09BA" box="[151,262,683,707]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
</emphasis>
:
<materialsCitation id="3B30156DFF88742FFE80FD78FCEC09BA" ID-GBIF-Occurrence="4906811301" box="[287,774,683,707]" collectionCode="MNHNL" pageId="4" pageNumber="5" specimenCode="BM790" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E3A192FF88742FFE80FD78FE6D09BA" box="[287,391,683,707]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
is specimen
<collectionCode id="ED4987F5FF88742FFDA2FD7FFD4809BA" box="[573,674,684,707]" pageId="4" pageNumber="5">MNHNL</collectionCode>
<specimenCode id="DBFEB74BFF88742FFD29FD78FCEC09BA" box="[694,774,683,707]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM790</specimenCode>
</materialsCitation>
(
<figureCitation id="136303B5FF88742FFF00FD18FF37099A" box="[159,221,715,739]" captionStart="Fig" captionStartId="4.[167,198,1848,1868]" captionTargetBox="[176,751,1099,1836]" captionTargetId="figure-400@4.[176,751,1099,1836]" captionTargetPageId="4" captionText="Fig. 5 Cenoceras rumelangense n.sp., holoytpe (specimen MNHNL BM790),‘Marnes sableuses d’Audun-le-Tiche’, Humphriesianum Zone, Berenskaul,Rumelange, Luxembourg" figureDoi="http://doi.org/10.5281/zenodo.12001084" httpUri="https://zenodo.org/record/12001084/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BE71F30FF88742FFF08FD38FD5508BA" blockId="4.[151,775,235,963]" pageId="4" pageNumber="5">
<materialsCitation id="3B30156DFF88742FFF08FD38FEB508FA" ID-GBIF-Occurrence="4906811302" collectionCode="MNHNL" location="Humphriesianum Zone of Rumelange" pageId="4" pageNumber="5" specimenCode="BM232, BM237, BM278, BM280, BM370, BM372, BM383, BM641, BM783, BM784, BM791, BM792, BM793" specimenCount="17" typeStatus="paratype">
<emphasis id="B92CC322FF88742FFF08FD38FEE7087A" bold="true" box="[151,269,747,771]" pageId="4" pageNumber="5">
<typeStatus id="54E3A192FF88742FFF08FD38FEE7087A" box="[151,269,747,771]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D5ED4B9FF88742FFEBEFD38FE54087A" box="[289,446,747,771]" count="17" pageId="4" pageNumber="5" type="generic">17 specimens</specimenCount>
from the
<location id="8E8749EBFF88742FFDDEFD38FE9B085A" LSID="urn:lsid:plazi:treatment:03F1AE26FF88742DFF08FF38FDA00F99:8E8749EBFF88742FFDDEFD38FE9B085A" name="Humphriesianum Zone of Rumelange" pageId="4" pageNumber="5">Humphriesianum Zone of Rumelange</location>
(
<collectionCode id="ED4987F5FF88742FFE1FFCD8FE00085A" box="[384,490,779,803]" pageId="4" pageNumber="5">MNHNL</collectionCode>
:
<specimenCode id="DBFEB74BFF88742FFE6BFCD8FDAD085A" box="[500,583,779,803]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM232</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFDCDFCD8FD4F085A" box="[594,677,779,803]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM237</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFD2FFCD8FCE9085A" box="[688,771,779,803]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM278</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFF08FCF8FF00083A" box="[151,234,811,835]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM280</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFE9DFCF8FEBC083A" box="[258,342,811,835]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM370</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFEF1FCF8FE2B083A" box="[366,449,811,835]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM372</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFE46FCF8FDC6083A" box="[473,556,811,835]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM383</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFDDBFCF8FD72083A" box="[580,664,811,835]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM641</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFD2FFCF8FCE8083A" box="[688,770,811,835]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM783</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFF08FC98FF00081A" box="[151,234,843,867]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM784</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFE9AFC98FEB2081A" box="[261,344,843,867]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM791</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFEEDFC98FE2F081A" box="[370,453,843,867]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM792</specimenCode>
,
<specimenCode id="DBFEB74BFF88742FFE40FC98FDD9081A" box="[479,563,843,867]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="4" pageNumber="5">BM793</specimenCode>
; MEP1; MNM1; Geolor 1 &amp; 2) and
</materialsCitation>
<materialsCitation id="3B30156DFF88742FFEF9FCBFFD5108BA" ID-GBIF-Occurrence="4906811303" collectionCode="SMNS" location="Gruibingen" municipality="Middle Jurassic Ostreenkalk Formation" pageId="4" pageNumber="5" specimenCode="SMNS 62245" specimenCount="1" stateProvince="Baden-Wuerttemberg" typeStatus="paratype">
<specimenCount id="9D5ED4B9FF88742FFEF9FCBFFDEA08FA" box="[358,512,875,899]" count="1" pageId="4" pageNumber="5" type="generic">one specimen</specimenCount>
from the
<collectingMunicipality id="6B83854AFF88742FFDF1FCB8FE2E08DA" pageId="4" pageNumber="5">Middle Jurassic Ostreenkalk Formation</collectingMunicipality>
(Humphriesianum Zone) of
<location id="8E8749EBFF88742FFF08FC78FEF008BA" LSID="urn:lsid:plazi:treatment:03F1AE26FF88742DFF08FF38FDA00F99:8E8749EBFF88742FFF08FC78FEF008BA" box="[151,282,939,963]" municipality="Middle Jurassic Ostreenkalk Formation" name="Gruibingen" pageId="4" pageNumber="5" stateProvince="Baden-Wuerttemberg">Gruibingen</location>
,
<collectingRegion id="499CD1D2FF88742FFEBAFC78FDE708BA" box="[293,525,939,963]" country="Germany" name="Baden-Wuerttemberg" pageId="4" pageNumber="5">Baden-Württemberg</collectingRegion>
(
<specimenCode id="DBFEB74BFF88742FFD84FC78FD5908BA" box="[539,691,939,963]" collectionCode="SMNS" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="4" pageNumber="5">SMNS 62245</specimenCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF274FB8FF88742FFCA2FC6EFBF90F7E" ID-DOI="http://doi.org/10.5281/zenodo.12001070" ID-Zenodo-Dep="12001070" httpUri="https://zenodo.org/record/12001070/files/figure.png" pageId="4" pageNumber="5" startId="4.[829,860,957,977]" targetBox="[837,1412,236,944]" targetPageId="4" targetType="figure">
<paragraph id="8BE71F30FF88742FFCA2FC6EFBF90F7E" blockId="4.[829,1413,957,1031]" pageId="4" pageNumber="5">
<emphasis id="B92CC322FF88742FFCA2FC6EFC8508A8" bold="true" box="[829,879,957,977]" pageId="4" pageNumber="5">Fig. 6</emphasis>
<taxonomicName id="4C5864B3FF88742FFCE5FC6EFBAF08A8" authority="Weis &amp; Schweigert &amp; Wittische, 2023" authorityName="Weis &amp; Schweigert &amp; Wittische" authorityYear="2023" box="[890,1093,957,977]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="rumelangense" status="sp. nov.">
<emphasis id="B92CC322FF88742FFCE5FC6EFBAF08A8" box="[890,1093,957,977]" italics="true" pageId="4" pageNumber="5">Cenoceras rumelangense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7E59FF88742FFBD5FC6EFB9908A8" box="[1098,1139,957,977]" pageId="4" pageNumber="5" rank="species">n.sp.</taxonomicNameLabel>
, paratype (specimen MNHNL BM383),‘Marnes sableuses d’Audun-le-Tiche’, Humphriesianum Zone, Rumelange,Luxembourg
</paragraph>
</caption>
<paragraph id="8BE71F30FF88742FFCB2FB9DFBB30FBF" blockId="4.[813,1437,1101,1926]" pageId="4" pageNumber="5">
<emphasis id="B92CC322FF88742FFCB2FB9DFC3E0F1C" bold="true" box="[813,980,1101,1126]" pageId="4" pageNumber="5">Type locality</emphasis>
: Rumelange, abandoned quarry site
<emphasis id="B92CC322FF88742FFCB2FBBDFCDF0FFC" box="[813,821,1134,1157]" italics="true" pageId="4" pageNumber="5">ʽ</emphasis>
Berenskaul-Waisskaul’ on the Luxembourgish–French border (Coordinates
<geoCoordinate id="EE6C79F7FF88742FFB83FB5DFB490FDC" box="[1052,1187,1164,1190]" degrees="49" direction="north" minutes="27" orientation="latitude" pageId="4" pageNumber="5" precision="15" seconds="12" value="49.453335">
49°27
<emphasis id="B92CC322FF88742FFBC5FB5FFB8F0FDC" box="[1114,1125,1164,1189]" italics="true" pageId="4" pageNumber="5">′</emphasis>
12
<emphasis id="B92CC322FF88742FFB1FFB5FFB640FDC" box="[1152,1166,1164,1189]" italics="true" pageId="4" pageNumber="5">ʺ</emphasis>
N
</geoCoordinate>
and
<geoCoordinate id="EE6C79F7FF88742FFB41FB5DFABE0FDC" box="[1246,1364,1164,1190]" degrees="6" direction="east" minutes="00" orientation="longitude" pageId="4" pageNumber="5" precision="15" seconds="41" value="6.011389">
6°00
<emphasis id="B92CC322FF88742FFA91FB5FFAF30FDC" box="[1294,1305,1164,1189]" italics="true" pageId="4" pageNumber="5">′</emphasis>
41
<emphasis id="B92CC322FF88742FFAAAFB5FFAA90FDC" box="[1333,1347,1164,1189]" italics="true" pageId="4" pageNumber="5">ʺ</emphasis>
E
</geoCoordinate>
; Bintz et al., 1973, p. 159) (
<figureCitation id="136303B5FF88742FFB92FB7DFBA00FBF" box="[1037,1098,1198,1222]" captionStart="Fig" captionStartId="2.[167,198,583,603]" captionTargetBox="[176,750,236,570]" captionTargetId="figure-735@2.[176,750,236,570]" captionTargetPageId="2" captionText="Fig. 3 Stratigraphic position (marked by a black arrow) and geographic location of the type locality (marked by a red circle) nearby Rumelange. Map of Southwest Luxembourg and stratigraphic log: Service géologique du Luxembourg (www.geology.lu)" figureDoi="http://doi.org/10.5281/zenodo.12001050" httpUri="https://zenodo.org/record/12001050/files/figure.png" pageId="4" pageNumber="5">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BE71F30FF88742FFCB2FB1DFAD60E7C" blockId="4.[813,1437,1101,1926]" pageId="4" pageNumber="5">
<emphasis id="B92CC322FF88742FFCB2FB1DFC3F0F9C" bold="true" box="[813,981,1229,1254]" pageId="4" pageNumber="5">Type horizon</emphasis>
: ‘Marnes sableuses d’Audun-le-Tiche’ (dom4), lower Bajocian, Humphriesianum Zone.
</paragraph>
</subSubSection>
<subSubSection id="C3424CBBFF88742FFCB2FADEFC800EDF" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BE71F30FF88742FFCB2FADEFC800EDF" blockId="4.[813,1437,1101,1926]" pageId="4" pageNumber="5">
<emphasis id="B92CC322FF88742FFCB2FADEFC4A0E5C" bold="true" box="[813,928,1293,1317]" pageId="4" pageNumber="5">Diagnosis</emphasis>
: Giant-sized, narrow-umbilicate species of
<taxonomicName id="4C5864B3FF88742FFCB2FAFDFC710E3C" authorityName="Hyatt" authorityYear="1883" box="[813,923,1326,1349]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
<emphasis id="B92CC322FF88742FFCB2FAFDFC710E3C" box="[813,923,1326,1349]" italics="true" pageId="4" pageNumber="5">Cenoceras</emphasis>
</taxonomicName>
with strongly inflated subtrapezoidal whorl section, generally higher than wide; longitudinal striation on the flattened venter and on the outer third of the flank.
</paragraph>
</subSubSection>
<subSubSection id="C3424CBBFF88742DFCB2FA7EFDA00F99" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph id="8BE71F30FF88742EFCB2FA7EFE940EA9" blockId="4.[813,1437,1101,1926]" lastBlockId="5.[151,775,1208,1904]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<emphasis id="B92CC322FF88742FFCB2FA7EFC5C0EBC" bold="true" box="[813,950,1453,1477]" pageId="4" pageNumber="5">Description</emphasis>
: The
<typeStatus id="54E3A192FF88742FFC68FA7DFBBD0EBF" box="[1015,1111,1454,1478]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
(
<figureCitation id="136303B5FF88742FFBF7FA7DFB4D0EBF" box="[1128,1191,1454,1478]" captionStart="Fig" captionStartId="4.[167,198,1848,1868]" captionTargetBox="[176,751,1099,1836]" captionTargetId="figure-400@4.[176,751,1099,1836]" captionTargetPageId="4" captionText="Fig. 5 Cenoceras rumelangense n.sp., holoytpe (specimen MNHNL BM790),‘Marnes sableuses d’Audun-le-Tiche’, Humphriesianum Zone, Berenskaul,Rumelange, Luxembourg" figureDoi="http://doi.org/10.5281/zenodo.12001084" httpUri="https://zenodo.org/record/12001084/files/figure.png" pageId="4" pageNumber="5">Fig. 5</figureCitation>
) is a steinkern with a few preserved remnants of the shell. The body-chamber occupies half of the last whorl. The last septa are more closely spaced, indicating that the specimen reached the adult age. The venter is distinctly flat and is ornamented with dense longitudinal striae that are particularly well developed on the body-chamber and which extend on the outer third of the flanks. The section of the body-chamber is markedly subtrapezoidal, compressed (compression index is 0.81 on the body-chamber and 0.77 on the phragmocone); the compression is more marked in younger whorls and less strong on the body-chamber but does not affect the lateral dimensions. The flanks are slightly rounded on the inner third (located towards the umbilicus) and distinctly flattened to slightly concave on the outer two thirds. The flanks are covered with fine falciform striae, and the outer third shows a reticulate ornamentation pattern. The phragmocone shows rather distant septa: the number of septa on the last whorl is
<emphasis id="B92CC322FF89742EFF08FAEAFF4F0E29" box="[151,165,1337,1360]" italics="true" pageId="5" pageNumber="6">n</emphasis>
<emphasis id="B92CC322FF89742EFF36FAEBFF510E29" bold="true" box="[169,187,1336,1360]" pageId="5" pageNumber="6">=</emphasis>
10. The lateral lobe is distinctly large and shallow (depth of the lateral lobe
<emphasis id="B92CC322FF89742EFE37FA8BFE500E09" bold="true" box="[424,442,1368,1392]" pageId="5" pageNumber="6">=</emphasis>
<quantity id="4CA0B2D5FF89742EFE22FA8BFDE60E09" box="[445,524,1368,1392]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="mm" value="10.0">10 mm</quantity>
), and is followed by an even less profound ventral lobe. The latero-ventral saddle is rather distinctive. Traces of the siphuncle are not visible in this specimen.
</paragraph>
<caption id="DF274FB8FF88742FFF38F8EBFE080CF8" ID-DOI="http://doi.org/10.5281/zenodo.12001084" ID-Zenodo-Dep="12001084" httpUri="https://zenodo.org/record/12001084/files/figure.png" pageId="4" pageNumber="5" startId="4.[167,198,1848,1868]" targetBox="[176,751,1099,1836]" targetPageId="4" targetType="figure">
<paragraph id="8BE71F30FF88742FFF38F8EBFE080CF8" blockId="4.[167,752,1848,1921]" pageId="4" pageNumber="5">
<emphasis id="B92CC322FF88742FFF38F8EBFF300C35" bold="true" box="[167,218,1848,1868]" pageId="4" pageNumber="5">Fig. 5</emphasis>
<taxonomicName id="4C5864B3FF88742FFF7BF8EBFE5A0C35" authority="Weis &amp; Schweigert &amp; Wittische, 2023" authorityName="Weis &amp; Schweigert &amp; Wittische" authorityYear="2023" box="[228,432,1848,1868]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="rumelangense" status="sp. nov.">
<emphasis id="B92CC322FF88742FFF7BF8EBFE5A0C35" box="[228,432,1848,1868]" italics="true" pageId="4" pageNumber="5">Cenoceras rumelangense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7E59FF88742FFE2AF8EBFE370C35" box="[437,477,1848,1868]" pageId="4" pageNumber="5" rank="species">n.sp.</taxonomicNameLabel>
, holoytpe (specimen MNHNL BM790),‘Marnes sableuses d’Audun-le-Tiche’, Humphriesianum Zone, Berenskaul,Rumelange, Luxembourg
</paragraph>
</caption>
<caption id="DF274FB8FF89742EFCA2FC37FC400F31" ID-DOI="http://doi.org/10.5281/zenodo.12001111" ID-Zenodo-Dep="12001111" httpUri="https://zenodo.org/record/12001111/files/figure.png" pageId="5" pageNumber="6" startId="5.[829,860,996,1016]" targetBox="[837,1412,236,983]" targetPageId="5" targetType="figure">
<paragraph id="8BE71F30FF89742EFCA2FC37FC400F31" blockId="5.[829,1391,996,1096]" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFCA2FC37FC850881" bold="true" box="[829,879,996,1016]" pageId="5" pageNumber="6">Fig. 8</emphasis>
<taxonomicName id="4C5864B3FF89742EFCE5FC37FBAF0881" authority="Weis &amp; Schweigert &amp; Wittische, 2023" authorityName="Weis &amp; Schweigert &amp; Wittische" authorityYear="2023" box="[890,1093,996,1016]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="rumelangense" status="sp. nov.">
<emphasis id="B92CC322FF89742EFCE5FC37FBAF0881" box="[890,1093,996,1016]" italics="true" pageId="5" pageNumber="6">Cenoceras rumelangense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7E59FF89742EFBD5FC37FB990881" box="[1098,1139,996,1016]" pageId="5" pageNumber="6" rank="species">n.sp.</taxonomicNameLabel>
, paratype,largest measured specimen (specimen MNHNL BM791),‘Marnes sableuses d’Audun-le-Tiche’, Humphriesianum Zone, Berenskaul, Rumelange, Luxembourg
</paragraph>
</caption>
<caption id="DF274FB8FF89742EFF38FBC1FEFE0F0F" ID-DOI="http://doi.org/10.5281/zenodo.12001123" ID-Zenodo-Dep="12001123" httpUri="https://zenodo.org/record/12001123/files/figure.png" pageId="5" pageNumber="6" startId="5.[167,198,1042,1062]" targetBox="[176,751,236,1029]" targetPageId="5" targetType="figure">
<paragraph id="8BE71F30FF89742EFF38FBC1FEFE0F0F" blockId="5.[167,741,1042,1142]" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFF38FBC1FF300F5F" bold="true" box="[167,218,1042,1062]" pageId="5" pageNumber="6">Fig. 7</emphasis>
<taxonomicName id="4C5864B3FF89742EFF7BFBC1FE5A0F5F" authority="Weis &amp; Schweigert &amp; Wittische, 2023" authorityName="Weis &amp; Schweigert &amp; Wittische" authorityYear="2023" box="[228,432,1042,1062]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="rumelangense" status="sp. nov.">
<emphasis id="B92CC322FF89742EFF7BFBC1FE5A0F5F" box="[228,432,1042,1062]" italics="true" pageId="5" pageNumber="6">Cenoceras rumelangense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F7E59FF89742EFE2AFBC1FE370F5F" box="[437,477,1042,1062]" pageId="5" pageNumber="6" rank="species">n.sp.</taxonomicNameLabel>
, paratype, inner whorl of a bigger specimen (specimen MNHNL BM783),‘Marnes sableuses d’Audun-le-Tiche’,Humphriesianum Zone,Berenskaul, Rumelange, Luxembourg
</paragraph>
</caption>
<paragraph id="8BE71F30FF89742EFF08FA0BFED60D49" blockId="5.[151,775,1208,1904]" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFF08FA0BFEE40E89" bold="true" box="[151,270,1496,1520]" pageId="5" pageNumber="6">Umbilicus</emphasis>
: Throughout all ontogenetic stages the overhanging umbilicus remains open, with a well-rounded umbilical edge.
</paragraph>
<paragraph id="8BE71F30FF89742EFF08F9EBFE490DA9" blockId="5.[151,775,1208,1904]" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFF08F9EBFF0E0D29" bold="true" box="[151,228,1592,1616]" pageId="5" pageNumber="6">Flanks</emphasis>
: Lower part gradually rising from the steep and overhanging umbilicus forming a well-rounded umbilical edge. Middle part of flanks form a trapezoidal section; upper part become slightly concave in the latest part of the adult body-chamber.
</paragraph>
<paragraph id="8BE71F30FF89742EFF08F90BFED80C49" blockId="5.[151,775,1208,1904]" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFF08F90BFF0E0D89" bold="true" box="[151,228,1752,1776]" pageId="5" pageNumber="6">Venter</emphasis>
: Venter is rounded only in the juvenile stage and soon becomes flattened by developing a prominent ventrolateral edge.
</paragraph>
<paragraph id="8BE71F30FF89742EFF08F8EBFAAF0E68" blockId="5.[151,775,1208,1904]" lastBlockId="5.[813,1437,1209,1905]" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFF08F8EBFEF10C29" bold="true" box="[151,283,1848,1872]" pageId="5" pageNumber="6">Suture line</emphasis>
: On the flank, the suture line forms a single, relatively shallow lateral lobe, with its deepest part located in the outer third of the flank. From the saddle at the ventrolateral edge, it turns towards mid-venter in a right angle without forming an extra ventral lobe.
</paragraph>
<paragraph id="8BE71F30FF89742EFCB2FACAFA6F0EA8" blockId="5.[813,1437,1209,1905]" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFCB2FACAFC770E48" bold="true" box="[813,925,1305,1329]" pageId="5" pageNumber="6">Ontogeny</emphasis>
: Growth pattern lacks principal changes of coiling or other modifications during ontogeny, except the inflated whorl section gradually changing from being rounded in the juvenile stage to subtrapezoidal in the medium and adult stages. In the latest stage, the chambers and suture lines are narrowing thus indicating adult size.
</paragraph>
<paragraph id="8BE71F30FF89742EFCB2FA0AFB210D88" blockId="5.[813,1437,1209,1905]" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFCB2FA0AFC0B0E88" bold="true" box="[813,993,1497,1521]" pageId="5" pageNumber="6">Ornamentation</emphasis>
: The almost uncompressed specimen from Swabia (
<figureCitation id="136303B5FF89742EFC54FA2AFBE10D68" box="[971,1035,1529,1553]" captionStart="Fig" captionStartId="6.[167,198,1051,1071]" captionTargetBox="[176,751,236,1039]" captionTargetId="figure-644@6.[176,751,236,1039]" captionTargetPageId="6" captionText="Fig. 9 Cenoceras rumelangense n.sp., paratype (specimen SMNS 62245), Ostreenkalk Formation, Humphriesianum Zone, Gruibingen, Southwest Germany" figureDoi="http://doi.org/10.5281/zenodo.12001131" httpUri="https://zenodo.org/record/12001131/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
) is the only one with a completely preserved calcitic replacement shell. The adult shell is ornamented with dense longitudinal striae both on the venter and on the outer third of the flank. The aperture does not exhibit any apertural modifications and crosses the venter in a straight way. Due to the shell preservation, the beginning of the body-chamber and the suture lines of the phragmocone cannot be traced.
</paragraph>
<paragraph id="8BE71F30FF89742DFCB2F92AFDA00F99" blockId="5.[813,1437,1209,1905]" lastBlockId="6.[151,774,1193,1249]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B92CC322FF89742EFCB2F92AFB200C68" bold="true" box="[813,1226,1785,1809]" pageId="5" pageNumber="6">Measurements and morphometrics</emphasis>
: Various measurements and parameters of
<taxonomicName id="4C5864B3FF89742EFBFFF8C9FAF10C48" authorityName="Weis &amp; Schweigert &amp; Wittische" authorityYear="2023" box="[1120,1307,1817,1841]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="rumelangense">
<emphasis id="B92CC322FF89742EFBFFF8C9FAF10C48" box="[1120,1307,1817,1841]" italics="true" pageId="5" pageNumber="6">C. rumelangense</emphasis>
</taxonomicName>
specimens are reported in
<tableCitation id="C6DA2A8BFF89742EFC74F8EAFBAA0C28" box="[1003,1088,1849,1873]" captionStart="Table 1" captionStartId="7.[151,209,238,260]" captionTargetPageId="7" captionText="Table 1 Measurements and parameters of Cenoceras rumelangense (unpublished), Metacenoceras clausum (data from Enay et al., 1994) and Cenoceras. gr. obesum sensu Rulleau (data from Rulleau, 2008)" httpUri="http://table.plazi.org/id/DF274FB8FF8B742CFF08FF3DFCE20A58" pageId="5" pageNumber="6" tableUuid="DF274FB8FF8B742CFF08FF3DFCE20A58">Table 1</tableCitation>
. We compared the data with measurements and parameters published for two coeval large-sized nautilid taxa:
<taxonomicName id="4C5864B3FF8A742DFE2BFB7AFD940FB8" baseAuthorityName="Sowerby" baseAuthorityYear="1816" box="[436,638,1193,1217]" class="Cephalopoda" family="Nautilidae" genus="Cenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="obesum">
<emphasis id="B92CC322FF8A742DFE2BFB7AFD940FB8" box="[436,638,1193,1217]" italics="true" pageId="6" pageNumber="7">Cenoceras obesum</emphasis>
</taxonomicName>
(sensu
<bibRefCitation id="EFC962C1FF8A742DFD47FB7AFEEE0F98" author="Rulleau, L." pageId="6" pageNumber="7" refId="ref9313" refString="Rulleau, L. (2008). Les Nautiles du Lias et du Dogger de la Region Lyonnaise. Lozanne: Section geo-paleo du comite d'etablissement des carrieres Lafarge." type="book" year="2008">Rulleau, 2008</bibRefCitation>
) and
<taxonomicName id="4C5864B3FF8A742DFEDDFB1AFDAF0F98" baseAuthorityName="Orbigny" baseAuthorityYear="1843" box="[322,581,1225,1249]" class="Cephalopoda" genus="Metacenoceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="clausum">
<emphasis id="B92CC322FF8A742DFEDDFB1AFDAF0F98" box="[322,581,1225,1249]" italics="true" pageId="6" pageNumber="7">Metacenoceras clausum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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