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<document id="95296E4124AEFD931A40EA63B0819646" ID-CLB-Dataset="22232" ID-DOI="10.11646/zootaxa.4776.1.1" ID-GBIF-Dataset="d92dcbf9-8bc1-43a6-8be1-b553a257ca0b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3821288" ID-ZooBank="5A3743A2-9E64-45E8-920D-92C9ED9CAB5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589267646334" checkinUser="plazi" docAuthor="Serrano, Artur R. M., Capela, Rúben A., Nunes, Telmo &amp; Santos, Carmen Van-Dú- Nem Neto" docDate="2020" docId="1B3C87AA004FFFBCFF1DFE26FDC7FB05" docLanguage="en" docName="zootaxa.4776.1.1.pdf.imf" docOrigin="Zootaxa 4776 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Gnathocera cruda subsp. gracilis" docType="treatment" docVersion="7" lastPageNumber="95" masterDocId="E705FFD20011FFE2FF8AFFFAFFA4FFA6" masterDocTitle="The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes" masterLastPageNumber="130" masterPageNumber="1" pageNumber="95" updateTime="1698839254326" updateUser="ExternalLinkService">
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<mods:title id="5BC0ACC3266B85649AAA1E5128A36066">The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes</mods:title>
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<mods:namePart id="A0765D3FA796833298545F551FEAAE15">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="DEF0E0CE32353A177FC385964696B394">Capela, Rúben A.</mods:namePart>
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<mods:namePart id="2B5104314E8C5EBD2A5DF4691A5C5754">Nunes, Telmo</mods:namePart>
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<mods:namePart id="CC4A70AF67F3DF58E40D7E95CC1C36B6">Santos, Carmen Van-Dú- Nem Neto</mods:namePart>
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<treatment id="1B3C87AA004FFFBCFF1DFE26FDC7FB05" ID-DOI="http://doi.org/10.5281/zenodo.3845323" ID-GBIF-Taxon="164123809" ID-Zenodo-Dep="3845323" LSID="urn:lsid:plazi:treatment:1B3C87AA004FFFBCFF1DFE26FDC7FB05" httpUri="http://treatment.plazi.org/id/1B3C87AA004FFFBCFF1DFE26FDC7FB05" lastPageNumber="95" pageId="94" pageNumber="95">
<subSubSection id="DB8F6537004FFFBCFF1DFE26FDDEFE51" box="[151,634,476,503]" pageId="94" pageNumber="95" type="nomenclature">
<paragraph id="932A36BC004FFFBCFF1DFE26FDDEFE51" blockId="94.[151,1436,476,1187]" box="[151,634,476,503]" pageId="94" pageNumber="95">
<heading id="C86281D0004FFFBCFF1DFE26FDDEFE51" bold="true" box="[151,634,476,503]" fontSize="11" level="1" pageId="94" pageNumber="95" reason="1">
<taxonomicName id="54954D3F004FFFBCFF1DFE26FDDEFE51" authority="(Janson, 1877)" baseAuthorityName="Janson" baseAuthorityYear="1877" box="[151,634,476,503]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="cruda" subSpecies="gracilis">
<emphasis id="A1E1EAAE004FFFBCFF1DFE26FDDEFE51" bold="true" box="[151,634,476,503]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF1DFE26FE63FE51" bold="true" box="[151,455,476,503]" italics="true" pageId="94" pageNumber="95">Gnathocera cruda gracilis</emphasis>
(Janson, 1877)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537004FFFBCFF1DFDFBFEA2FDBD" box="[151,262,513,539]" pageId="94" pageNumber="95" type="description">
<paragraph id="932A36BC004FFFBCFF1DFDFBFEA2FDBD" blockId="94.[151,1436,476,1187]" box="[151,262,513,539]" pageId="94" pageNumber="95">
(
<figureCitation id="0BAE2A39004FFFBCFF2AFDFBFF5AFDBD" box="[160,254,513,539]" captionStart="FIGURE 22" captionStartId="97.[151,250,1876,1901]" captionTargetBox="[247,1340,229,1838]" captionTargetId="figure@97.[235,1352,181,1851]" captionTargetPageId="97" captionText="FIGURE 22. Cetoniinae species of Angola. a, Gnathocera cruda gracilis (dorsal view), male, MZUC; b, Gnathocera cruda leopoldi (dorsal view), male, MZUC; c and d, Gnathocera marginata (dorsal views), males, MZUC and Dundo, LNO, IICT, respectively; e, Gnathocera overlaeti (dorsal view), male, Dundo, LNO, IICT; f, Gnathocera trivialis (dorsal view), male, Muita, LNO, IICT; g and h, Gnathocera trivittata ardoini (dorsal views), male and female, respectively, Dundo, LNO, IICT; i, Gnathocera trivittata dorsodiscolor (dorsal view), male, MZUC." figureDoi="http://doi.org/10.5281/zenodo.3821332" httpUri="https://zenodo.org/record/3821332/files/figure.png" pageId="94" pageNumber="95">Fig. 22a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537004FFFBCFF4DFDDEFB80FD55" pageId="94" pageNumber="95" type="distribution">
<paragraph id="932A36BC004FFFBCFF4DFDDEFD99FD99" blockId="94.[151,1436,476,1187]" box="[199,573,548,575]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFDDEFEFDFD99" bold="true" box="[199,345,548,575]" pageId="94" pageNumber="95">Distribution</emphasis>
: ANG, CON, DOC.
</paragraph>
<paragraph id="932A36BC004FFFBCFF4DFDB2FCB4FDC5" blockId="94.[151,1436,476,1187]" box="[199,784,584,611]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFDB2FDCEFDC5" bold="true" box="[199,618,584,611]" pageId="94" pageNumber="95">
Distribution in
<collectingCountry id="EB82762C004FFFBCFEF7FDB2FE77FDC5" box="[381,467,584,611]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
(Provinces):
</emphasis>
1) LNO, UIG.
</paragraph>
<paragraph id="932A36BC004FFFBCFF4DFD96FE36FD21" blockId="94.[151,1436,476,1187]" box="[199,402,620,647]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFD96FE36FD21" bold="true" box="[199,402,620,647]" pageId="94" pageNumber="95">Historic records:</emphasis>
</paragraph>
<paragraph id="932A36BC004FFFBCFF4DFD6BFCC7FD0A" blockId="94.[151,1436,476,1187]" box="[199,867,657,684]" pageId="94" pageNumber="95">
<collectingCountry id="EB82762C004FFFBCFF6EFD6BFE92FD0D" box="[228,310,657,683]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
(without locality), Bembe (UIG) (
<bibRefCitation id="F7044B4D004FFFBCFD3DFD6BFCF2FD0A" author="Janson, O. E." box="[695,854,657,684]" pageId="94" pageNumber="95" pagination="247 - 264" refId="ref84162" refString="Janson, O. E. (1877 b) Notices of new or little known Cetoniidae No. 3. Cistula Entomologica, 2, 247 - 264." type="journal article" year="1877">Janson 1877b</bibRefCitation>
);
</paragraph>
<paragraph id="932A36BC004FFFBCFF4DFD4FFC76FD69" blockId="94.[151,1436,476,1187]" box="[199,978,692,719]" pageId="94" pageNumber="95">
—Dundo (LNO) (
<bibRefCitation id="F7044B4D004FFFBCFE19FD4FFDCCFD69" author="Gomes Alves, M. G." box="[403,616,692,719]" pageId="94" pageNumber="95" pagination="63 - 88" refId="ref83304" refString="Gomes Alves, M. G. (1959) Cetonideos da Africa portuguesa. Memorias da Junta de Investigacao do Ultramar, 8, 63 - 88, pls. 1 - 8." type="journal article" year="1959">Gomes Alves 1959</bibRefCitation>
sub
<taxonomicName id="54954D3F004FFFBCFD17FD4FFC62FD69" authority="Janson" authorityName="Janson" authorityYear="1877" box="[669,966,693,719]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="lurida">
<emphasis id="A1E1EAAE004FFFBCFD17FD4FFCCAFD69" box="[669,878,693,719]" italics="true" pageId="94" pageNumber="95">Gnathocera lurida</emphasis>
Janson
</taxonomicName>
);
</paragraph>
<paragraph id="932A36BC004FFFBCFF4DFD23FB80FD55" blockId="94.[151,1436,476,1187]" box="[199,1060,729,756]" pageId="94" pageNumber="95">
—Dundo, Chicapa (LNO),
<collectingCountry id="EB82762C004FFFBCFE73FD23FDEFFD55" box="[505,587,729,755]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
(LNO, without locality) (
<bibRefCitation id="F7044B4D004FFFBCFCE1FD23FBBCFD55" author="Antoine, Ph." box="[875,1048,729,755]" pageId="94" pageNumber="95" pagination="65 - 92" refId="ref80368" refString="Antoine, Ph. (2002 a) Contribution a la connaissance des Cetoniidae africains (Coleoptera, Scarabaeoidea). Coleopteres, 8 (4), 65 - 92." type="journal article" year="2002">Antoine 2002a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537004FFFBCFF4DFD06FD25FC25" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="932A36BC004FFFBCFF4DFD06FD25FC25" blockId="94.[151,1436,476,1187]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFD06FE17FCB1" bold="true" box="[199,435,764,791]" pageId="94" pageNumber="95">Material examined:</emphasis>
<materialsCitation id="23FD3CE1004FFFBCFE31FD07FBE9FCBE" ID-GBIF-Occurrence="2612165512" box="[443,1101,765,792]" country="Angola" location="Angola" pageId="94" pageNumber="95" specimenCount="2" specimenCount-male="2">
<collectingCountry id="EB82762C004FFFBCFE31FD07FDB4FCB1" box="[443,528,765,791]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
,
<specimenCount id="8593FD35004FFFBCFD96FD07FDEDFCB1" box="[540,585,765,791]" pageId="94" pageNumber="95" type="male">2 ♂</specimenCount>
, Nº 1907 (blue labels), nº 146a (white labels)
</materialsCitation>
,
<specimenCount id="8593FD35004FFFBCFBD3FD07FB27FCB1" box="[1113,1155,765,791]" pageId="94" pageNumber="95" type="female">1 ♀</specimenCount>
, Nº 1907 (blue label), nº 147a (white label),
<materialsCitation id="23FD3CE1004FFFBCFEFFFCDBFCA6FCF9" ID-GBIF-Occurrence="2612165358" collectingDate="1949-03" collectionCode="LNO" country="Angola" elevation="1470" latitude="-7.366667" longLatPrecision="1303" longitude="20.833334" pageId="94" pageNumber="95" specimenCount="1" specimenCount-male="1">
<specimenCount id="8593FD35004FFFBCFEFFFCDBFE06FC9D" box="[373,418,801,827]" pageId="94" pageNumber="95" type="male">1 ♂</specimenCount>
, Nº 1907 (blue label), nº 148a (white label), old collection, box nº IX-78, MZUC; Dundo (
<geoCoordinate id="F6A1507B004FFFBCFF2AFCBFFF53FCF9" box="[160,247,837,863]" degrees="7" direction="south" minutes="22" orientation="latitude" pageId="94" pageNumber="95" precision="925" value="-7.366667">7º 22´S</geoCoordinate>
,
<geoCoordinate id="F6A1507B004FFFBCFE8FFCBFFECFFCF9" box="[261,363,836,863]" degrees="20" direction="east" minutes="50" orientation="longitude" pageId="94" pageNumber="95" precision="925" value="20.833334">20º 50´E</geoCoordinate>
,
<quantity id="546D9B59004FFFBCFEF2FCBFFE6BFCF9" box="[376,463,837,863]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="94" pageNumber="95" unit="m" value="1470.0">
<elevation id="18B8D18F004FFFBCFEF2FCBFFE6BFCF9" box="[376,463,837,863]" metricMagnitude="3" metricUnit="m" metricValue="1.47" pageId="94" pageNumber="95" unit="m" value="1470.0">1470 m</elevation>
</quantity>
alt., 51) (
<collectionCode id="F584AE79004FFFBCFDCAFCBFFDDBFCF9" box="[576,639,837,863]" pageId="94" pageNumber="95">LNO</collectionCode>
), -.
<date id="E72B107C004FFFBCFD2AFCBFFCA6FCF9" box="[672,770,837,863]" pageId="94" pageNumber="95" value="1949-03">
<collectingDate id="F76FE994004FFFBCFD2AFCBFFCA6FCF9" box="[672,770,837,863]" pageId="94" pageNumber="95" value="1949-03">III.1949</collectingDate>
</date>
</materialsCitation>
,
<specimenCount id="8593FD35004FFFBCFC85FCBEFC99FCF9" box="[783,829,836,863]" pageId="94" pageNumber="95" type="male">5 ♂</specimenCount>
,
<specimenCount id="8593FD35004FFFBCFCC1FCBFFCD3FCF9" box="[843,887,837,863]" pageId="94" pageNumber="95" type="female">4 ♀</specimenCount>
, Nº 2132,
<materialsCitation id="23FD3CE1004FFFBCFC7EFCBFFDD9FC25" ID-GBIF-Occurrence="2612165376" country="Angola" location="Junta de Investigacoes Coloniais" pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="2">
<specimenCount id="8593FD35004FFFBCFC7EFCBFFB84FCF9" box="[1012,1056,837,863]" pageId="94" pageNumber="95" type="female">2 ♀</specimenCount>
, Nº 2151,
<location id="964A6067004FFFBCFB17FCBFFEA6FC25" LSID="urn:lsid:plazi:treatment:1B3C87AA004FFFBCFF1DFE26FDC7FB05:964A6067004FFFBCFB17FCBFFEA6FC25" country="Angola" name="Junta de Investigacoes Coloniais" pageId="94" pageNumber="95">Junta de Investigações Coloniais</location>
(Entomologia), IICT, MUHNAC
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DB8F6537004FFFBCFF4DFC77FDC7FB05" pageId="94" pageNumber="95" type="discussion">
<paragraph id="932A36BC004FFFBCFF4DFC77FDC7FB05" blockId="94.[151,1436,476,1187]" pageId="94" pageNumber="95">
<emphasis id="A1E1EAAE004FFFBCFF4DFC77FE9FFC01" bold="true" box="[199,315,909,935]" pageId="94" pageNumber="95">Remarks.</emphasis>
The status of this taxon is very controversial. Its validity was established by
<bibRefCitation id="F7044B4D004FFFBCFB11FC77FAF2FC01" author="Antoine, Ph." box="[1179,1366,909,935]" pageId="94" pageNumber="95" pagination="65 - 92" refId="ref80368" refString="Antoine, Ph. (2002 a) Contribution a la connaissance des Cetoniidae africains (Coleoptera, Scarabaeoidea). Coleopteres, 8 (4), 65 - 92." type="journal article" year="2002">Antoine (2002a)</bibRefCitation>
, a criterion followed by
<bibRefCitation id="F7044B4D004FFFBCFEE5FC4BFDC6FC6D" author="Beinhundner, G." box="[367,610,945,971]" pageId="94" pageNumber="95" refId="ref81744" refString="Beinhundner, G. (2017 a) The Cetoniinae of Africa (Coleoptera: Scarabaeidae). Privately published, Euerbach, 1199 pp." type="book" year="2017">Beinhundner (2017a)</bibRefCitation>
. However,
<bibRefCitation id="F7044B4D004FFFBCFD68FC4BFC31FC6D" author="Krajcik, M." box="[738,917,945,971]" pageId="94" pageNumber="95" refId="ref84860" refString="Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI." type="book" year="2008">Krajcik (2008a)</bibRefCitation>
as well as
<bibRefCitation id="F7044B4D004FFFBCFB99FC4BFAB1FC6D" author="Schoolmeesters, P." box="[1043,1301,945,971]" pageId="94" pageNumber="95" pagination="2017" refId="ref88502" refString="Schoolmeesters, P. (2018) Scarabs: World Scarabaeidae Database (version Jan 2017). In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. &amp; Penev, L. (Eds.), Species 2000 &amp; ITIS Catalogue of Life, 2017 Annual Checklist. Available from: http: // www. catalogueoflife. org / annual-checklist / 2017 (accessed 12 January 2020)" type="book chapter" year="2018">Schoolmeesters (2018)</bibRefCitation>
keep it as a synonym of
<taxonomicName id="54954D3F004FFFBCFEAAFC2FFDB8FC49" authority="Janson, 1877" authorityName="Janson" authorityYear="1877" box="[288,540,981,1007]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="lurida">
<emphasis id="A1E1EAAE004FFFBCFEAAFC2FFE21FC49" box="[288,389,981,1007]" italics="true" pageId="94" pageNumber="95">G. lurida</emphasis>
Janson, 1877
</taxonomicName>
[the name
<emphasis id="A1E1EAAE004FFFBCFD1FFC2FFD7FFC49" box="[661,731,981,1007]" italics="true" pageId="94" pageNumber="95">lucida</emphasis>
used by
<bibRefCitation id="F7044B4D004FFFBCFCB1FC2FFC48FC49" author="Krajcik, M." box="[827,1004,981,1007]" pageId="94" pageNumber="95" refId="ref84860" refString="Krajcik, M. (2008 a) Cetoniidae of the World. Catalogue - Part I. (Coleoptera: Scarabaeoidea). Animma. x, Supplement, 1 - 96 + I-XXXVI." type="book" year="2008">Krajcik (2008a)</bibRefCitation>
is a misspelling of
<taxonomicName id="54954D3F004FFFBCFB4AFC2FFAA0FC49" authorityName="Janson" authorityYear="1877" box="[1216,1284,981,1007]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="lurida">
<emphasis id="A1E1EAAE004FFFBCFB4AFC2FFAA0FC49" box="[1216,1284,981,1007]" italics="true" pageId="94" pageNumber="95">lurida</emphasis>
</taxonomicName>
, since
<bibRefCitation id="F7044B4D004FFFBCFADAFC2FFF54FBB2" author="Janson, O. E." pageId="94" pageNumber="95" pagination="247 - 264" refId="ref84162" refString="Janson, O. E. (1877 b) Notices of new or little known Cetoniidae No. 3. Cistula Entomologica, 2, 247 - 264." type="journal article" year="1877">Janson (1877b)</bibRefCitation>
did not describe any taxon named
<emphasis id="A1E1EAAE004FFFBCFD0AFC03FD62FBB5" box="[640,710,1017,1043]" italics="true" pageId="94" pageNumber="95">lucida</emphasis>
].
<taxonomicName id="54954D3F004FFFBCFD55FC03FBA9FBB5" baseAuthorityName="Janson" baseAuthorityYear="1877" box="[735,1037,1017,1043]" class="Insecta" family="Cetoniidae" genus="Gnathocera" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="cruda" subSpecies="gracilis">
<emphasis id="A1E1EAAE004FFFBCFD55FC03FBA9FBB5" box="[735,1037,1017,1043]" italics="true" pageId="94" pageNumber="95">Gnathocera cruda gracilis</emphasis>
</taxonomicName>
was described based on one specimen from
<collectingCountry id="EB82762C004FFFBCFE81FBE7FEC4FB91" box="[267,352,1053,1079]" name="Angola" pageId="94" pageNumber="95">Angola</collectingCountry>
, but
<bibRefCitation id="F7044B4D004FFFBCFE1DFBE7FDE7FB9E" author="Janson, O. E." box="[407,579,1053,1080]" pageId="94" pageNumber="95" pagination="247 - 264" refId="ref84162" refString="Janson, O. E. (1877 b) Notices of new or little known Cetoniidae No. 3. Cistula Entomologica, 2, 247 - 264." type="journal article" year="1877">Janson (1877b)</bibRefCitation>
referred that “there are also specimens in Mr. Higginss collection found by Monteiro, at Bembe”, a municipality within the
<collectingRegion id="5151F85E004FFFBCFD27FBBBFCF5FBFD" box="[685,849,1089,1115]" country="Angola" name="Uige" pageId="94" pageNumber="95">Uíge Province</collectingRegion>
. The adult specimens of IICT were collected by light (registered information) probably at the same time of
<taxonomicName id="54954D3F004FFFBCFD62FB9FFC67FBD9" box="[744,963,1125,1151]" class="Insecta" family="Cetoniidae" genus="Chondrorrhina" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="trivittata" subSpecies="ardoini">
<emphasis id="A1E1EAAE004FFFBCFD62FB9FFC67FBD9" box="[744,963,1125,1151]" italics="true" pageId="94" pageNumber="95">G. trivittata ardoini</emphasis>
</taxonomicName>
(see remarks under this taxon). Ecological data are not available for this subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>