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<document id="F3222850C3082C853BFF4B2693EEC9CD" ID-CLB-Dataset="22232" ID-DOI="10.11646/zootaxa.4776.1.1" ID-GBIF-Dataset="d92dcbf9-8bc1-43a6-8be1-b553a257ca0b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3821288" ID-ZooBank="5A3743A2-9E64-45E8-920D-92C9ED9CAB5A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589267646334" checkinUser="plazi" docAuthor="Serrano, Artur R. M., Capela, Rúben A., Nunes, Telmo &amp; Santos, Carmen Van-Dú- Nem Neto" docDate="2020" docId="1B3C87AA0027FFD4FF1DFD6AFE68FB4D" docLanguage="en" docName="zootaxa.4776.1.1.pdf.imf" docOrigin="Zootaxa 4776 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Coenochilus calcaratus subsp. vagus" docType="treatment" docVersion="7" lastPageNumber="55" masterDocId="E705FFD20011FFE2FF8AFFFAFFA4FFA6" masterDocTitle="The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes" masterLastPageNumber="130" masterPageNumber="1" pageNumber="55" updateTime="1698839254326" updateUser="ExternalLinkService">
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<mods:title id="28CF655353CC2953B57065CDF2C00A6D">The rose chafers (Coleoptera: Scarabaeidae: Cetoniinae) of Angola: a descriptive checklist with new records and synonymic notes</mods:title>
</mods:titleInfo>
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<mods:namePart id="C8FB868F2C510AC9BEB51F89EEF28BEB">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="37A25FD5631F6B7F46DD507C4C6A9586">Capela, Rúben A.</mods:namePart>
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<mods:namePart id="18759164FABCAD68D88F411CDDF55F3E">Nunes, Telmo</mods:namePart>
</mods:name>
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<mods:namePart id="7A5674245263F51E939827749D29D31E">Santos, Carmen Van-Dú- Nem Neto</mods:namePart>
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<mods:date id="4B0E473FA7AD9CB0D418A0C5EA3DE128">2020</mods:date>
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<mods:number id="57CC71AF4D8EB6E06F9F475A76A569D3">4776</mods:number>
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<mods:identifier id="536E82366F252FAF20DB66F4B0D30970" type="DOI">10.11646/zootaxa.4776.1.1</mods:identifier>
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<treatment id="1B3C87AA0027FFD4FF1DFD6AFE68FB4D" ID-DOI="http://doi.org/10.5281/zenodo.3845429" ID-GBIF-Taxon="164123924" ID-Zenodo-Dep="3845429" LSID="urn:lsid:plazi:treatment:1B3C87AA0027FFD4FF1DFD6AFE68FB4D" httpUri="http://treatment.plazi.org/id/1B3C87AA0027FFD4FF1DFD6AFE68FB4D" lastPageNumber="55" pageId="54" pageNumber="55">
<subSubSection id="DB8F65370027FFD4FF1DFD6AFD6CFD0D" box="[151,712,656,683]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="932A36BC0027FFD4FF1DFD6AFD6CFD0D" blockId="54.[151,1437,656,1259]" box="[151,712,656,683]" pageId="54" pageNumber="55">
<heading id="C86281D00027FFD4FF1DFD6AFD6CFD0D" bold="true" box="[151,712,656,683]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<taxonomicName id="54954D3F0027FFD4FF1DFD6AFD6CFD0D" authority="(Peringuey, 1907)" baseAuthorityName="Peringuey" baseAuthorityYear="1907" box="[151,712,656,683]" class="Insecta" family="Cetoniidae" genus="Coenochilus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="subSpecies" species="calcaratus" subSpecies="vagus">
<emphasis id="A1E1EAAE0027FFD4FF1DFD6AFD6CFD0D" bold="true" box="[151,712,656,683]" pageId="54" pageNumber="55">
<emphasis id="A1E1EAAE0027FFD4FF1DFD6AFE55FD0D" bold="true" box="[151,497,656,683]" italics="true" pageId="54" pageNumber="55">Coenochilus calcaratus vagus</emphasis>
(
<bibRefCitation id="F7044B4D0027FFD4FD8BFD6AFD64FD0D" author="Peringuey, L." box="[513,704,656,683]" pageId="54" pageNumber="55" pagination="289 - 546" refId="ref86844" refString="Peringuey, L. (1907) Descriptive catalogue of the Coleoptera of South Africa. Fam. Lucanidae. Fam. Scarabaeidae. Sub-families: Sericinae, Melolonthinae, Trichiinae, Valginae, Cetoniinae. Transactions of the South African Philosophical Society, 13, 289 - 546, pl. 47." type="journal article" year="1907">Péringuey, 1907</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DB8F65370027FFD4FF1DFD4FFEA3FD76" box="[151,263,693,720]" pageId="54" pageNumber="55" type="description">
<paragraph id="932A36BC0027FFD4FF1DFD4FFEA3FD76" blockId="54.[151,1437,656,1259]" box="[151,263,693,720]" pageId="54" pageNumber="55">
(
<figureCitation id="0BAE2A390027FFD4FF2AFD4FFF5BFD76" box="[160,255,693,720]" captionStart="FIGURE 12" captionStartId="57.[151,250,1876,1901]" captionTargetBox="[286,1346,188,1835]" captionTargetId="figure@57.[233,1355,181,1850]" captionTargetPageId="57" captionText="FIGURE 12. Cetoniinae species of Angola. a, Coenochilus basilewskyi (lateral view), female, Dundo, LNO, IICT; b, Coenochilus calcaratus vagus (dorsal view), male, Calulo, CUS, ASC; c and d, Coenochilus procerus (dorsal views), male and female, respectively, BembuaCamenhe, HUA, ASC; e, Coenochilus turbatus (dorsal view), male, Calulo, CUS, ASC; f, Coenochilus ventricosus (dorsal view), male, Cacuso, MAL, ASC; g, Plagiochilus angustatus (dorsal view), male, Carima, MAL, ASC; h, Plagiochilus angustatus: adult feeding in a tomentose herbaceus plant, Carima, MAL." figureDoi="http://doi.org/10.5281/zenodo.3821312" httpUri="https://zenodo.org/record/3821312/files/figure.png" pageId="54" pageNumber="55">Fig. 12b</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DB8F65370027FFD4FF4DFD22FC7EFC56" pageId="54" pageNumber="55" type="distribution">
<paragraph id="932A36BC0027FFD4FF4DFD22FD26FD55" blockId="54.[151,1437,656,1259]" box="[199,642,728,755]" pageId="54" pageNumber="55">
<emphasis id="A1E1EAAE0027FFD4FF4DFD22FEFDFD55" bold="true" box="[199,345,728,755]" pageId="54" pageNumber="55">Distribution</emphasis>
: ANG, DOC, MOZ, ZIM.
</paragraph>
<paragraph id="932A36BC0027FFD4FF4DFD06FCD3FCB1" blockId="54.[151,1437,656,1259]" box="[199,887,764,791]" pageId="54" pageNumber="55">
<emphasis id="A1E1EAAE0027FFD4FF4DFD06FDCEFCB1" bold="true" box="[199,618,764,791]" pageId="54" pageNumber="55">
Distribution in
<collectingCountry id="EB82762C0027FFD4FEF7FD06FE77FCB1" box="[381,467,764,791]" name="Angola" pageId="54" pageNumber="55">Angola</collectingCountry>
(Provinces):
</emphasis>
1) HUA; 2) CUS, HUI.
</paragraph>
<paragraph id="932A36BC0027FFD4FF4DFCDAFE36FC9D" blockId="54.[151,1437,656,1259]" box="[199,402,800,827]" pageId="54" pageNumber="55">
<emphasis id="A1E1EAAE0027FFD4FF4DFCDAFE36FC9D" bold="true" box="[199,402,800,827]" pageId="54" pageNumber="55">Historic records:</emphasis>
</paragraph>
<paragraph id="932A36BC0027FFD4FF4DFCBFFDFDFCF9" blockId="54.[151,1437,656,1259]" box="[199,601,836,863]" pageId="54" pageNumber="55">
—Bailundo (HUA) (
<bibRefCitation id="F7044B4D0027FFD4FE25FCBFFDE9FCF9" author="Schein, H." box="[431,589,836,863]" pageId="54" pageNumber="55" pagination="271 - 325" refId="ref88247" refString="Schein, H. (1954 a) Uber Coenochilus (Coleoptera, Cetoniidae). Eine Revision der afrikanischen Arten mit einer Bestimmungstabelle und ein Nachtrag zu meiner Revision der asiatischen Arten. Entomologische Arbeiten aus dem Museum G. Frey, 5, 271 - 325." type="journal article" year="1954">Schein 1954a</bibRefCitation>
);
</paragraph>
<paragraph id="932A36BC0027FFD4FF4DFC93FD1DFC25" blockId="54.[151,1437,656,1259]" box="[199,697,872,900]" pageId="54" pageNumber="55">
<collectingCountry id="EB82762C0027FFD4FF6EFC93FE92FC25" box="[228,310,873,899]" name="Angola" pageId="54" pageNumber="55">Angola</collectingCountry>
(without locality) (
<bibRefCitation id="F7044B4D0027FFD4FD84FC93FD08FC25" author="Ferreira, M. C." box="[526,684,872,899]" pageId="54" pageNumber="55" pagination="415 - 1317" refId="ref83177" refString="Ferreira, M. C. (1965) Catalogo dos Coleopteros de Angola. Revista de Entomologia de Mocambique, 8 (2), 415 - 1317." type="journal article" year="1965">Ferreira 1965</bibRefCitation>
);
</paragraph>
<paragraph id="932A36BC0027FFD4FF4DFC77FB1BFC0E" blockId="54.[151,1437,656,1259]" box="[199,1215,909,936]" pageId="54" pageNumber="55">
<collectingCountry id="EB82762C0027FFD4FF6EFC77FE92FC01" box="[228,310,909,935]" name="Angola" pageId="54" pageNumber="55">Angola</collectingCountry>
(without locality) (
<bibRefCitation id="F7044B4D0027FFD4FD84FC77FD50FC01" author="Sakai, K. &amp; Nagai, S." box="[526,756,909,935]" pageId="54" pageNumber="55" pagination="1 - 421" refId="ref88030" refString="Sakai, K. &amp; Nagai, S. (1998) The Cetoniine Beetles of the World. Mushi-Sha's Iconographic Series of Insects, Tokyo, Japan, 3, 1 - 421." type="journal article" year="1998">Sakai &amp; Nagai 1998</bibRefCitation>
sub
<taxonomicName id="54954D3F0027FFD4FCA3FC77FB10FC0E" authority="Westwood" authorityName="Westwood" authorityYear="1874" box="[809,1204,909,936]" class="Insecta" family="Cetoniidae" genus="Coenochilus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="A1E1EAAE0027FFD4FCA3FC77FB96FC01" box="[809,1074,909,935]" italics="true" pageId="54" pageNumber="55">Coenochilus calcaratus</emphasis>
Westwood
</taxonomicName>
).
</paragraph>
<paragraph id="932A36BC0027FFD4FF4DFC4BFC7EFC56" blockId="54.[151,1437,656,1259]" pageId="54" pageNumber="55">
<emphasis id="A1E1EAAE0027FFD4FF4DFC4BFE7AFC6D" bold="true" box="[199,478,944,971]" pageId="54" pageNumber="55">New localities reported</emphasis>
:
<materialsCitation id="23FD3CE10027FFD4FE64FC4BFC72FC56" ID-GBIF-Occurrence="2612165372" collectingDate="2013-11-11" collectingDateMax="2013-11-12" collectingDateMin="2013-11-11" collectionCode="SW, HUI" collectorName="P. Schule &amp; P. Malec" elevation="1613" latitude="-14.148055" longLatPrecision="21" longitude="14.47111" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1">
Negola (
<quantity id="546D9B590027FFD4FDDAFC4AFD05FC6D" box="[592,673,944,971]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="54" pageNumber="55" unit="km" value="3.5">3,5 km</quantity>
<collectionCode id="F584AE790027FFD4FD23FC4BFD71FC6D" box="[681,725,945,971]" pageId="54" pageNumber="55">SW</collectionCode>
) (
<geoCoordinate id="F6A1507B0027FFD4FD64FC4BFC37FC6D" box="[750,915,944,971]" degrees="14" direction="south" minutes="08" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="53" value="-14.148055">14º 08´53´´ S</geoCoordinate>
,
<geoCoordinate id="F6A1507B0027FFD4FC2BFC4BFBECFC6D" box="[929,1096,945,971]" degrees="14" direction="east" minutes="28" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="16" value="14.47111">14º 28´16´´ E</geoCoordinate>
,
<quantity id="546D9B590027FFD4FBDFFC4BFB08FC6D" box="[1109,1196,945,971]" metricMagnitude="3" metricUnit="m" metricValue="1.613" pageId="54" pageNumber="55" unit="m" value="1613.0">
<elevation id="18B8D18F0027FFD4FBDFFC4BFB08FC6D" box="[1109,1196,945,971]" metricMagnitude="3" metricUnit="m" metricValue="1.613" pageId="54" pageNumber="55" unit="m" value="1613.0">1613 m</elevation>
</quantity>
alt., 317) (
<collectionCode id="F584AE790027FFD4FAA6FC4BFAC7FC6D" box="[1324,1379,945,971]" pageId="54" pageNumber="55">HUI</collectionCode>
),
<date id="E72B107C0027FFD4FAF2FC4BFEBDFC49" pageId="54" pageNumber="55" value="2013-11-11" valueMax="2013-11-12" valueMin="2013-11-11">
<collectingDate id="F76FE9940027FFD4FAF2FC4BFEBDFC49" pageId="54" pageNumber="55" value="2013-11-11" valueMax="2013-11-12" valueMin="2013-11-11">11- 12.XI.2013</collectingDate>
</date>
,
<specimenCount id="8593FD350027FFD4FEAFFC2FFEEDFC49" box="[293,329,981,1007]" pageId="54" pageNumber="55" type="male">1♂</specimenCount>
,
<collectorName id="3E60536A0027FFD4FEDCFC2FFE18FC49" box="[342,444,981,1007]" pageId="54" pageNumber="55">P. Schüle</collectorName>
leg., PMC (
<collectorName id="3E60536A0027FFD4FDCFFC2FFD0EFC49" box="[581,682,981,1007]" pageId="54" pageNumber="55">P. Malec</collectorName>
, personal communication)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DB8F65370027FFD4FF4DFC02FCB7FB91" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="932A36BC0027FFD4FF4DFC02FCB7FB91" blockId="54.[151,1437,656,1259]" pageId="54" pageNumber="55">
<emphasis id="A1E1EAAE0027FFD4FF4DFC02FE12FBB5" bold="true" box="[199,438,1016,1043]" pageId="54" pageNumber="55">Material examined:</emphasis>
<materialsCitation id="23FD3CE10027FFD4FE4AFC03FCABFB91" ID-GBIF-Occurrence="2612165371" collectingDate="2015-12-03" collectionCode="CUS" collectorName="A. Serrano &amp; R. Capela" elevation="882" latitude="-9.944722" longLatPrecision="21" longitude="14.943055" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1">
Calulo (Faz. Belo Horizonte) (
<geoCoordinate id="F6A1507B0027FFD4FCA8FC03FC19FBB5" box="[802,957,1016,1043]" degrees="9" direction="south" minutes="56" orientation="latitude" pageId="54" pageNumber="55" precision="15" seconds="41" value="-9.944722">9º 56´41´´ S</geoCoordinate>
,
<geoCoordinate id="F6A1507B0027FFD4FC47FC03FBD3FBB5" box="[973,1143,1016,1043]" degrees="14" direction="east" minutes="56" orientation="longitude" pageId="54" pageNumber="55" precision="15" seconds="35" value="14.943055">14º 56´35´´ E</geoCoordinate>
,
<quantity id="546D9B590027FFD4FB0DFC03FB75FBB5" box="[1159,1233,1017,1043]" metricMagnitude="2" metricUnit="m" metricValue="8.82" pageId="54" pageNumber="55" unit="m" value="882.0">
<elevation id="18B8D18F0027FFD4FB0DFC03FB75FBB5" box="[1159,1233,1017,1043]" metricMagnitude="2" metricUnit="m" metricValue="8.82" pageId="54" pageNumber="55" unit="m" value="882.0">882 m</elevation>
</quantity>
alt., 128) (
<collectionCode id="F584AE790027FFD4FADEFC03FA2BFBB5" box="[1364,1423,1017,1043]" country="USA" lsid="urn:lsid:biocol.org:col:12908" name="Cusino Wildlife Research Station" pageId="54" pageNumber="55" type="Herbarium">CUS</collectionCode>
),
<date id="E72B107C0027FFD4FF1DFBE6FEB1FB91" box="[151,277,1052,1079]" pageId="54" pageNumber="55" value="2015-12-03">
<collectingDate id="F76FE9940027FFD4FF1DFBE6FEB1FB91" box="[151,277,1052,1079]" pageId="54" pageNumber="55" value="2015-12-03">3.XII.2015</collectingDate>
</date>
,
<specimenCount id="8593FD350027FFD4FEAAFBE7FEE8FB91" box="[288,332,1053,1079]" pageId="54" pageNumber="55" type="male">1 ♂</specimenCount>
, DO,
<collectorName id="3E60536A0027FFD4FE07FBE7FDA2FB91" box="[397,518,1053,1079]" pageId="54" pageNumber="55">A. Serrano</collectorName>
&amp;
<collectorName id="3E60536A0027FFD4FDA0FBE7FD3DFB91" box="[554,665,1053,1079]" pageId="54" pageNumber="55">R. Capela</collectorName>
leg., ASC
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DB8F65370027FFD4FF4DFBBBFE68FB4D" pageId="54" pageNumber="55" type="discussion">
<paragraph id="932A36BC0027FFD4FF4DFBBBFE68FB4D" blockId="54.[151,1437,656,1259]" pageId="54" pageNumber="55">
<emphasis id="A1E1EAAE0027FFD4FF4DFBBBFE9FFBFD" bold="true" box="[199,315,1089,1115]" pageId="54" pageNumber="55">Remarks.</emphasis>
A polytypic species with three subspecies within the
<taxonomicName id="54954D3F0027FFD4FC1AFBBBFB82FBFD" authorityName="Westwood" authorityYear="1874" box="[912,1062,1089,1115]" class="Insecta" family="Cetoniidae" genus="Coenochilus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calcaratus">
<emphasis id="A1E1EAAE0027FFD4FC1AFBBBFB82FBFD" box="[912,1062,1089,1115]" italics="true" pageId="54" pageNumber="55">C. calcaratus</emphasis>
</taxonomicName>
group (
<emphasis id="A1E1EAAE0027FFD4FBF4FBB8FB18FBFD" box="[1150,1212,1090,1115]" italics="true" pageId="54" pageNumber="55">sensu</emphasis>
<bibRefCitation id="F7044B4D0027FFD4FB49FBBBFAFBFBFD" author="Schein, H." box="[1219,1375,1088,1115]" pageId="54" pageNumber="55" pagination="271 - 325" refId="ref88247" refString="Schein, H. (1954 a) Uber Coenochilus (Coleoptera, Cetoniidae). Eine Revision der afrikanischen Arten mit einer Bestimmungstabelle und ein Nachtrag zu meiner Revision der asiatischen Arten. Entomologische Arbeiten aus dem Museum G. Frey, 5, 271 - 325." type="journal article" year="1954">Schein 1954a</bibRefCitation>
). The Calulo adult specimen was collected on the soil in a countryroad located in a secondary open forest. Ecological data on abundance and habitat preferences was given for the nominal subspecies for
<collectingCountry id="EB82762C0027FFD4FB82FB73FBF5FB05" box="[1032,1105,1161,1187]" name="Ghana" pageId="54" pageNumber="55">Ghana</collectingCountry>
(
<bibRefCitation id="F7044B4D0027FFD4FBD5FB73FA94FB05" author="Mudge, A. D. &amp; Orozco, J. &amp; Philips, T. K. &amp; Antoine, Ph." box="[1119,1328,1161,1187]" pageId="54" pageNumber="55" pagination="113 - 174" refId="ref86394" refString="Mudge, A. D., Orozco, J., Philips, T. K. &amp; Antoine, Ph. (2012) The cetoniine fauna of the Upper Guinean forests and savannas of Ghana (Coleoptera: Scarabaeidae: Cetoniinae). Terrestrial Arthropod Reviews, 5, 113 - 174. https: // doi. org / 10.1163 / 187498312 X 635319" type="journal article" year="2012">
Mudge
<emphasis id="A1E1EAAE0027FFD4FB3CFB70FB42FB05" box="[1206,1254,1161,1187]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 2012
</bibRefCitation>
). See also general comments in
<bibRefCitation id="F7044B4D0027FFD4FE0DFB57FD95FB61" author="Schein, H." box="[391,561,1196,1223]" pageId="54" pageNumber="55" pagination="271 - 325" refId="ref88247" refString="Schein, H. (1954 a) Uber Coenochilus (Coleoptera, Cetoniidae). Eine Revision der afrikanischen Arten mit einer Bestimmungstabelle und ein Nachtrag zu meiner Revision der asiatischen Arten. Entomologische Arbeiten aus dem Museum G. Frey, 5, 271 - 325." type="journal article" year="1954">Schein (1954a)</bibRefCitation>
and the remarks under the genus
<taxonomicName id="54954D3F0027FFD4FC20FB57FB92FB61" authorityName="Schaum" authorityYear="1841" box="[938,1078,1197,1223]" class="Insecta" family="Cetoniidae" genus="Coenochilus" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="A1E1EAAE0027FFD4FC20FB57FB92FB61" box="[938,1078,1197,1223]" italics="true" pageId="54" pageNumber="55">Coenochilus</emphasis>
</taxonomicName>
. It is a new record for the Huíla and Kwanza Sul Provinces.
</paragraph>
</subSubSection>
</treatment>
</document>