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89 lines
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<document id="B98AD9BA7A82C594EB41A46E82BB2D64" ID-DOI="10.1111/j.1096-3642.2011.00731.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5442095" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630452512153" checkinUser="valdenar" docAuthor="Li, Liang & Zhou, Hong-Zhang" docDate="2011" docId="E0275714D9710E10FC0FEAD7FE71A8FF" docLanguage="en" docName="j.1096-3642.2011.00731.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00731.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Pseudohesperus Hayashi. Z 2008" docType="treatment" docVersion="4" lastPageNumber="691" masterDocId="1C1E2F6CD97D0E1DFFA5EC4DFFF6AE0C" masterDocTitle="Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="722" masterPageNumber="679" pageNumber="691" updateTime="1699275715089" updateUser="plazi">
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<mods:title id="B8555C8B81379F0B25211092D3F0AC5F">Revision and phylogenetic assessment of the rove beetle genus Pseudohesperus Hayashi, with broad reference to the subtribe Philonthina (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
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<mods:namePart id="D757294A99BF649849F8152CE3342EEE">Li, Liang</mods:namePart>
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<mods:namePart id="D884B20B12906EAFB188D0F689136E8C">Zhou, Hong-Zhang</mods:namePart>
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<mods:title id="4618F02174523DB21601FB69B5DADE8F">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="4DB622FF215E2424C0560366B12731B9">
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<mods:date id="3D28DC9C7A905ADFBDE42CDBDD71046C">2011</mods:date>
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<mods:number id="3EF7D5913FD3B833FFBFC6A7FEEA3A97">2011-10-25</mods:number>
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<mods:number id="BE2223C4A05D84C3E5EA283FD5306EC9">163</mods:number>
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<treatment id="E0275714D9710E10FC0FEAD7FE71A8FF" ID-DOI="http://doi.org/10.5281/zenodo.5492131" ID-Zenodo-Dep="5492131" LSID="urn:lsid:plazi:treatment:E0275714D9710E10FC0FEAD7FE71A8FF" httpUri="http://treatment.plazi.org/id/E0275714D9710E10FC0FEAD7FE71A8FF" lastPageId="13" lastPageNumber="691" pageId="12" pageNumber="691">
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<subSubSection id="2094B589D9710E11FC0FEAD7FAFFA8A1" box="[938,1289,1690,1711]" pageId="12" pageNumber="691" type="nomenclature">
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<paragraph id="6831E602D9710E11FC0FEAD7FAFFA8A1" blockId="12.[938,1289,1690,1711]" box="[938,1289,1690,1711]" pageId="12" pageNumber="691">
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<heading id="3379516ED9710E11FC0FEAD7FAFFA8A1" box="[938,1289,1690,1711]" centered="true" fontSize="8" level="2" pageId="12" pageNumber="691" reason="8">
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THE GENUS
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<taxonomicName id="AF8E9D81D9710E11FB9CEAD7FAFFA8A1" ID-CoL="9JMLR" authorityName="Hayashi. Z" authorityYear="2008" box="[1081,1289,1690,1711]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="691" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9710E11FB9CEAD7FAFFA8A1" box="[1081,1289,1690,1711]" italics="true" pageId="12" pageNumber="691">PSEUDOHESPERUS</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="2094B589D9710E10FC83EA8DFE71A8FF" lastPageId="13" lastPageNumber="692" pageId="12" pageNumber="691" type="materials_examined">
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<paragraph id="6831E602D9710E10FC83EA8DFE71A8FF" blockId="12.[806,1422,1728,1903]" lastBlockId="13.[164,779,1237,1779]" lastPageId="13" lastPageNumber="692" pageId="12" pageNumber="691">
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The phylogenetic analysis strongly supported the monophyly of the genus
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<taxonomicName id="AF8E9D81D9710E11FBE4EA93FB02A8FF" authorityName="Hayashi. Z" authorityYear="2008" box="[1089,1268,1758,1779]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="691" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9710E11FBE4EA93FB02A8FF" box="[1089,1268,1758,1779]" italics="true" pageId="12" pageNumber="691">Pseudohesperus</emphasis>
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</taxonomicName>
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by very high branch support scores. It also showed the sister relationship of
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<taxonomicName id="AF8E9D81D9710E11FC1FEB51FB9BA93D" authorityName="Hayashi. Z" authorityYear="2008" box="[954,1133,1820,1841]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="691" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9710E11FC1FEB51FB9BA93D" box="[954,1133,1820,1841]" italics="true" pageId="12" pageNumber="691">Pseudohesperus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="AF8E9D81D9710E11FB1FEB51FAE6A93D" box="[1210,1296,1820,1841]" class="Insecta" family="Staphylinidae" genus="Bisnius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="691" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9710E11FB1FEB51FAE6A93D" box="[1210,1296,1820,1841]" italics="true" pageId="12" pageNumber="691">Bisnius</emphasis>
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</taxonomicName>
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with low branch scores. This was consistent with the speculation of
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<bibRefCitation id="0C1F9BF3D9710E11FCDCEB14FBDEA963" author="Hayashi Y" box="[889,1064,1881,1903]" pageId="12" pageNumber="691" pagination="145 - 153" refId="ref20737" refString="Hayashi Y. 2008. Pseudohesperus, a new genus of Philonthina (Coleoptera: Staphylinidae) from East Asia, with redescription of its type species Philonthus rutiliventris Sharp. The Entomological Review of Japan 63: 145 - 153." type="journal article" year="2008">Hayashi (2008)</bibRefCitation>
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based on morphological study. The generic diagnosis of
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<taxonomicName id="AF8E9D81D9700E10FE62E898FD8CAAE6" authorityName="Hayashi. Z" authorityYear="2008" box="[455,634,1237,1258]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="692" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9700E10FE62E898FD8CAAE6" box="[455,634,1237,1258]" italics="true" pageId="13" pageNumber="692">Pseudohesperus</emphasis>
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</taxonomicName>
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provided by
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<bibRefCitation id="0C1F9BF3D9700E10FF01E8B9FEA2AB06" author="Hayashi Y" box="[164,340,1268,1290]" pageId="13" pageNumber="692" pagination="145 - 153" refId="ref20737" refString="Hayashi Y. 2008. Pseudohesperus, a new genus of Philonthina (Coleoptera: Staphylinidae) from East Asia, with redescription of its type species Philonthus rutiliventris Sharp. The Entomological Review of Japan 63: 145 - 153." type="journal article" year="2008">Hayashi (2008)</bibRefCitation>
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is not precise because of the limited material he examined and now requires a few improvements and additions: (1) apical segment of maxillary palps distinctly narrower than penultimate segment; (2) apical segment of labial palps slightly longer than penultimate segment; (3) gular sutures narrowly separated at base; (4) pronotum more or less densely punctured, but with narrow, impunctate midline; (5) apical portion of mesoventrite distinctly convex ventrad; (6) mesoventrite without transverse carina; (7) marginal carina of mesocoxal acetabula more or less straight; (8) underside of paramere of aedeagus with sensory peg setae. The evolutionary relationship of all the members of
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<taxonomicName id="AF8E9D81D9700E10FDF2EACFFCFCA89B" authorityName="Hayashi. Z" authorityYear="2008" box="[599,778,1666,1687]" class="Insecta" family="Staphylinidae" genus="Pseudohesperus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="692" phylum="Arthropoda" rank="genus">
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<emphasis id="5AFA3A10D9700E10FDF2EACFFCFCA89B" box="[599,778,1666,1687]" italics="true" pageId="13" pageNumber="692">Pseudohesperus</emphasis>
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</taxonomicName>
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was revealed by the phylogenetic analysis and can be correlated well with their zoogeographical patterns (see Zoogeography).
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</paragraph>
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