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<document id="C4F96D6BB10BB11CEF22EA9D46A9E369" ID-CLB-Dataset="26067" ID-DOI="10.11646/zootaxa.4652.1.2" ID-GBIF-Dataset="25c52d53-5912-4ae6-b986-1d320eadf654" ID-ISSN="1175-5326" ID-Zenodo-Dep="3363533" ID-ZooBank="B8EA4D0C-07CC-4C8F-9B21-C97B9852AAFF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565270322117" checkinUser="plazi" docAuthor="Smith, Robin James, Zhai, Dayou &amp; Chang, Cheon Young" docDate="2019" docId="A90187A1BA52FF88128BD51748D4F96B" docLanguage="en" docName="zootaxa.4652.1.2.pdf" docOrigin="Zootaxa 4652 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ilyocypris incus Smith &amp; Zhai &amp; Chang 2019, sp. nov." docType="treatment" docVersion="11" lastPageNumber="92" masterDocId="5538FFD9BA45FF94121CD4664A60FF88" masterDocTitle="Ilyocypris (Crustacea: Ostracoda) species in North East Asian rice fields description of one new species, and redescriptions of Ilyocypris dentifera Sars 1903 and Ilyocypris japonica Okubo, 1990" masterLastPageNumber="92" masterPageNumber="56" pageNumber="56" updateTime="1698749105675" updateUser="ExternalLinkService">
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<mods:title id="232A38EE03CA13F50D7016F97AD33713">Ilyocypris (Crustacea: Ostracoda) species in North East Asian rice fields description of one new species, and redescriptions of Ilyocypris dentifera Sars 1903 and Ilyocypris japonica Okubo, 1990</mods:title>
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<mods:namePart id="4B574B0D4A17FE6ADB5148357A5BA1A7">Zhai, Dayou</mods:namePart>
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<mods:namePart id="35604704A3240E15C4BA612EE65EF8B7">Chang, Cheon Young</mods:namePart>
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<treatment id="A90187A1BA52FF88128BD51748D4F96B" ID-DOI="http://doi.org/10.5281/zenodo.5610194" ID-GBIF-Taxon="159043963" ID-Zenodo-Dep="5610194" LSID="urn:lsid:plazi:treatment:A90187A1BA52FF88128BD51748D4F96B" httpUri="http://treatment.plazi.org/id/A90187A1BA52FF88128BD51748D4F96B" lastPageId="28" pageId="23">
<subSubSection id="69B2653CBA52FF83128BD5174BDCFE03" box="[151,444,369,396]" pageId="23" type="nomenclature">
<paragraph id="211736B7BA52FF83128BD5174BDCFE03" blockId="23.[151,444,369,431]" box="[151,444,369,396]" pageId="23">
<heading id="7A5F81DBBA52FF83128BD5174BDCFE03" bold="true" box="[151,444,369,396]" fontSize="11" level="1" pageId="23" reason="1">
<emphasis id="13DCEAA5BA52FF83128BD5174BDCFE03" bold="true" box="[151,444,369,396]" pageId="23">
<taxonomicName id="E6A84D34BA52FF83128BD5174B36FE03" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[151,342,369,396]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="23" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA52FF83128BD5174B36FE03" bold="true" box="[151,342,369,396]" italics="true" pageId="23">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA52FF831342D5174BDCFE03" box="[350,444,369,395]" pageId="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA52FF83128BD5F34B1FFE27" box="[151,383,404,431]" pageId="23" type="description">
<paragraph id="211736B7BA52FF83128BD5F34B1FFE27" blockId="23.[151,444,369,431]" box="[151,383,404,431]" pageId="23">
(
<figureCitation id="B9932A32BA52FF831283D5F34A96FE27" box="[159,246,404,431]" captionStart="FIGURE 5" captionStartId="9.[151,250,1475,1500]" captionTargetBox="[151,1436,181,1446]" captionTargetId="figure@9.[151,1436,181,1449]" captionTargetPageId="9" captionText="FIGURE 5. Light microscopy photographs of Ilyocypris hemipenes. A, Ilyocypris japonica Okubo, 1990 (LBM 1430009150, sample 18; see Table 1). B, Ilyocypris japonica Okubo, 1990 (LBM 1430009151, sample 18). C, Ilyocypris mongolica Martens, 1991 (LBM 1430009161, sample 17). D, Ilyocypris dentifera Sars, 1903 (LBM 1430009119, sample 6). E, Ilyocypris angulata Sars, 1903 (LBM 1430009112, sample 20). F, Ilyocypris incus sp. nov. (DB40054, sample 14). E has been reflected on the vertical axis to facilitate comparisons." httpUri="https://zenodo.org/record/3363543/files/figure.png" pageId="23">Figs 5F</figureCitation>
,
<figureCitation id="B9932A32BA52FF83131DD5F34B44FE27" box="[257,292,405,431]" captionStart="FIGURE 6" captionStartId="10.[151,250,1729,1754]" captionTargetBox="[151,1436,345,1704]" captionTargetId="figure@10.[151,1436,345,1704]" captionTargetPageId="10" captionText="FIGURE 6. Light microscopy photographs of Ilyocypris Zenker organs. Black triangles mark the anterior ends. A, Ilyocypris japonica Okubo, 1990 (LBM 1430009144, sample 9; see Table 1). B, Ilyocypris dentifera Sars, 1903 (LBM 1430009119, sam- ple 6). C, Ilyocypris incus sp. nov. (LBM 1430009131, sample 14)." httpUri="https://zenodo.org/record/3363545/files/figure.png" pageId="23">6C</figureCitation>
,
<figureCitation id="B9932A32BA52FF83132CD5F34B17FE27" box="[304,375,405,431]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStart-3="FIGURE 17" captionStartId-0="24.[151,250,1637,1662]" captionStartId-1="25.[151,250,1839,1864]" captionStartId-2="27.[151,250,1671,1696]" captionStartId-3="29.[151,250,1759,1784]" captionTargetBox-0="[286,1301,181,1611]" captionTargetBox-1="[238,1350,181,1814]" captionTargetBox-2="[219,1372,181,1646]" captionTargetBox-3="[160,1431,705,1735]" captionTargetId-0="figure@24.[286,1301,181,1611]" captionTargetId-1="figure@25.[238,1350,181,1814]" captionTargetId-2="figure@27.[215,1372,181,1646]" captionTargetId-3="figure@29.[143,1442,705,1735]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionTargetPageId-2="27" captionTargetPageId-3="29" captionText-0="FIGURE 14. Ilyocypris incus sp. nov. A, female left valve, external view (LBM 1430009136). B, female right valve, external view (LBM 1430009136). C, female left valve, internal view (LBM 1430009136). D, female right valve, internal view (LBM 1430009136). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009136). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009136). G, female carapace, dorsal view, anterior to right (LBM 1430009137). H, male right valve, external view (LBM1430009131). I, male carapace, dorsal view, anterior to right (LBM 1430009138). J, male right valve, ventral view (LBM 1430009133). K, male right valve, detail of ventral margin (LBM 1430009133). Scale bar = 500 µm for AD, GJ, 174 µm for E &amp; F, 135 µm for K. All specimens from sample 14; see Table 1." captionText-1="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." captionText-2="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." captionText-3="FIGURE 17. Summary of the soft part characters defining the two Ilyocypris groups recognized herein. (1) Only two males of I. bradyi have been reported, but they were not found with any adult females (Karanovic &amp; Lee 2013). (2) Adult females of I. hanguk are unknown (Karanovic &amp; Lee 2013). (3, 4, 8, 9) Outlines drawn from figures in Karanovic &amp; Lee (2013). (5, 6) Outlines drawn from photograph in Zhai &amp; Xiao (2013). (7) Outline drawn from figure in Smith et al. (2011). (10, 11) Outlines drawn from photograph in Ma et al. (2018). * indicates figures reflected on the vertical axis to facilitate comparisons. Not to scale." httpUri-0="https://zenodo.org/record/3363561/files/figure.png" httpUri-1="https://zenodo.org/record/3363563/files/figure.png" httpUri-2="https://zenodo.org/record/3363565/files/figure.png" httpUri-3="https://zenodo.org/record/3363567/files/figure.png" pageId="23">1417</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA52FF83128BD5BF4E76FD03" pageId="23" type="reference_group">
<paragraph id="211736B7BA52FF83128BD5BF497EFE79" blockId="23.[151,1358,473,651]" box="[151,798,473,498]" pageId="23">
1974
<taxonomicName id="E6A84D34BA52FF8312CCD5BF4866FE7A" authority="Sars, 1903" authorityName="Sars" authorityYear="1903" box="[208,518,473,498]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="23" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="13DCEAA5BA52FF8312CCD5BF4BF6FE79" box="[208,406,473,497]" italics="true" pageId="23">Ilyocypris angulata</emphasis>
<bibRefCitation id="45394B46BA52FF831380D5BF4866FE7A" author="Sars, G. O." box="[412,518,473,498]" pageId="23" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
—Okubo: 4650, fig. 1av.
</paragraph>
<paragraph id="211736B7BA52FF83128BD59E49F6FD98" blockId="23.[151,1358,473,651]" box="[151,918,503,528]" pageId="23">
? 1991a
<taxonomicName id="E6A84D34BA52FF8312F1D59E4843FD98" authority="Sars, 1903" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[237,547,503,528]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="23" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA52FF8312F1D59E4BD4FD98" box="[237,436,504,528]" italics="true" pageId="23">Ilyocypris dentifera</emphasis>
<bibRefCitation id="45394B46BA52FF8313A6D59E4843FD98" author="Sars, G. O." box="[442,547,503,528]" pageId="23" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
—Kim &amp; Min: 317321, figs 8 &amp; 9.
</paragraph>
<paragraph id="211736B7BA52FF83128BD671494EFDA7" blockId="23.[151,1358,473,651]" box="[151,814,534,559]" pageId="23">
2004
<taxonomicName id="E6A84D34BA52FF8312CCD6714867FDA7" authority="Sars, 1903" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[208,519,534,559]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="23" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA52FF8312CCD6714BF7FDA7" box="[208,407,535,559]" italics="true" pageId="23">Ilyocypris dentifera</emphasis>
<bibRefCitation id="45394B46BA52FF831382D6714867FDA7" author="Sars, G. O." box="[414,519,534,559]" pageId="23" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
—Okubo: 1617, figs 4mq.
</paragraph>
<paragraph id="211736B7BA52FF83128BD6534E3AFDC5" blockId="23.[151,1358,473,651]" box="[151,1114,565,590]" pageId="23">
? 2012
<taxonomicName id="E6A84D34BA52FF8312FED6534878FDC6" authority="Sars, 1903" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[226,536,565,590]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="23" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA52FF8312FED6534BC8FDC5" box="[226,424,565,589]" italics="true" pageId="23">Ilyocypris dentifera</emphasis>
<bibRefCitation id="45394B46BA52FF8313B3D6534878FDC6" author="Sars, G. O." box="[431,536,565,590]" pageId="23" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
—Chang
<emphasis id="13DCEAA5BA52FF831060D65048C9FDC5" box="[636,681,565,589]" italics="true" pageId="23">et al</emphasis>
.:
<tableCitation id="6C2A030CBA52FF8310A0D6534967FDC5" box="[700,775,565,590]" captionStart="TABLE 1" captionStartId="1.[151,244,1160,1187]" captionTargetBox="[163,1412,1310,1957]" captionText="TABLE 1. Adult Ilyocypris specimens examined for this study. Initial discriminations of species were based on lateral views of carapaces, and confirmed by dissections of male specimens. Figures in brackets indicate the number of specimens with tubercles on the carapace. Note that tubercles vary from weakly to well-developed and were no- ticeable on whole specimens or valves in dorsal view using a stereo optical microscope." httpUri="http://table.plazi.org/id/75D7663FBA44FF95128BD0EE4E0FFA87" pageId="23" tableUuid="75D7663FBA44FF95128BD0EE4E0FFA87">Table 1</tableCitation>
(referring to
<bibRefCitation id="45394B46BA52FF831192D6534E2EFDC5" author="Kim, W. &amp; Min, G. S." box="[910,1102,565,590]" pageId="23" pagination="307 - 322" refId="ref21073" refString="Kim, W. &amp; Min, G. S. (1991 a) Redescription of four Recent freshwater ostracod (Crustacea: Ostracoda) species from Korea. Korean Journal of Zoology, 34, 307 - 322." type="journal article" year="1991">Kim &amp; Min 1991a</bibRefCitation>
).
</paragraph>
<paragraph id="211736B7BA52FF83128BD6324F2EFDE4" blockId="23.[151,1358,473,651]" box="[151,1358,595,620]" pageId="23">
2013
<taxonomicName id="E6A84D34BA52FF8312CCD6324865FDE4" authority="Sars, 1903" authorityName="Sars" authorityYear="1903" box="[208,517,595,620]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="23" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="13DCEAA5BA52FF8312CCD6324BF6FDE4" box="[208,406,596,620]" italics="true" pageId="23">Ilyocypris angulata</emphasis>
<bibRefCitation id="45394B46BA52FF831380D6324865FDE4" author="Sars, G. O." box="[412,517,595,620]" pageId="23" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
—Karanovic &amp; Lee: 41, 55, 59, 64, fig. 14 (in part, referring to Okubo 1975 [sic]).
</paragraph>
<paragraph id="211736B7BA52FF83128BD6154E76FD03" blockId="23.[151,1358,473,651]" box="[151,1046,626,651]" pageId="23">
? 2014
<taxonomicName id="E6A84D34BA52FF8312FED6154877FD03" authority="Sars, 1903" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[226,535,626,651]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="23" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA52FF8312FED6154BC8FD03" box="[226,424,627,651]" italics="true" pageId="23">Ilyocypris dentifera</emphasis>
<bibRefCitation id="45394B46BA52FF8313B3D6154877FD03" author="Sars, G. O." box="[431,535,626,651]" pageId="23" pagination="3 - 44" refId="ref22910" refString="Sars, G. O. (1903) Fresh-water Entomostraca from China and Sumatra. Archiv for Mathematik og Naturvidenskab, 25, 3 - 44." type="journal article" year="1903">Sars, 1903</bibRefCitation>
</taxonomicName>
—Smith
<emphasis id="13DCEAA5BA52FF83106AD61548C3FD03" box="[630,675,627,651]" italics="true" pageId="23">et al</emphasis>
.: 1 (referring to
<bibRefCitation id="45394B46BA52FF831157D6144E6AFD03" author="Kim, W. &amp; Min, G. S." box="[843,1034,626,651]" pageId="23" pagination="307 - 322" refId="ref21073" refString="Kim, W. &amp; Min, G. S. (1991 a) Redescription of four Recent freshwater ostracod (Crustacea: Ostracoda) species from Korea. Korean Journal of Zoology, 34, 307 - 322." type="journal article" year="1991">Kim &amp; Min 1991a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA52FF88128BD6DF48B4FCD7" lastPageId="28" pageId="23" type="materials_examined">
<paragraph id="211736B7BA52FF83128BD6DF4829FD70" blockId="23.[151,1437,697,2045]" pageId="23">
<emphasis id="13DCEAA5BA52FF83128BD6DF4B7EFD5B" bold="true" box="[151,286,697,723]" pageId="23">Etymology.</emphasis>
From the Latin incus meaning anvil and referring to the shape of the copulatory process of the hemipenis (treated as a noun in apposition).
</paragraph>
<paragraph id="211736B7BA52FF8312DBD7664B77FB9C" blockId="23.[151,1437,697,2045]" pageId="23">
<emphasis id="13DCEAA5BA52FF8312DBD7664B22FC93" bold="true" box="[199,322,768,795]" pageId="23">Diagnosis.</emphasis>
Carapace quadrate in lateral view, covered in small pits, and approximately seven tiny spines near posterior margin. Anterior inner calcified lamellae of both valves with lists. With or without tubercles on carapace. Swimming setae of antenna very long, extending significantly beyond end of claws. Sixth limb with four segments (second endopodal segment undivided). Seventh limb with two setae on third segment (= second endopodal segment). Anterior end of Zenker organ small, slightly wider than central tube. Inner lobe of hemipenis longer than outer lobe, touching or slightly overlapping middle lobe, with bulbous end. Outer lobe nearly touching to slightly overlapping middle lobe, copulatory process well developed, with large, roughly triangular distal end with uneven distal edge.
</paragraph>
<paragraph id="211736B7BA52FF8312DBD0784BA3FBD4" blockId="23.[151,1437,697,2045]" pageId="23">
<emphasis id="13DCEAA5BA52FF8312DBD0784B0DFBB0" bold="true" box="[199,365,1053,1080]" pageId="23">Type locality.</emphasis>
Rice fields in Bomun-dong, Gyeongju-si,
<collectingRegion id="E36CF855BA52FF831169D0784E33FBB0" box="[885,1107,1054,1080]" country="South Korea" name="Gyeongbuk" pageId="23">Gyeongsangbuk-do</collectingRegion>
,
<collectingCountry id="59BF7627BA52FF83167AD0784E9CFBB1" box="[1126,1276,1054,1081]" name="South Korea" pageId="23">South Korea</collectingCountry>
,
<geoCoordinate id="449C5070BA52FF831713D07B4FF8FBB1" box="[1295,1432,1053,1081]" degrees="35.8364" direction="north" orientation="latitude" pageId="23" precision="5" value="35.8364">35.8364º N</geoCoordinate>
,
<geoCoordinate id="449C5070BA52FF83128BD0244B43FBD5" box="[151,291,1090,1117]" degrees="129.2483" direction="east" orientation="longitude" pageId="23" precision="5" value="129.2483">129.2483º E</geoCoordinate>
, altitude
<specimenCount id="37AEFD3EBA52FF831396D0244BA3FBD4" box="[394,451,1090,1116]" pageId="23" type="generic">
<quantity id="E6509B52BA52FF831396D0244BDFFBD4" box="[394,447,1090,1116]" metricMagnitude="1" metricUnit="m" metricValue="7.3" pageId="23" unit="m" value="73.0">73m</quantity>
.
</specimenCount>
</paragraph>
<paragraph id="211736B7BA52FF8312DBD0004977FB2B" blockId="23.[151,1437,697,2045]" pageId="23">
<emphasis id="13DCEAA5BA52FF8312DBD0004B18FB08" bold="true" box="[199,376,1125,1152]" pageId="23">Type material.</emphasis>
<materialsCitation id="91C03CEABA52FF83139ED0004977FB2B" ID-GBIF-Occurrence="2313474991" collectionCode="LBM" pageId="23" specimenCode="LBM 1430009132" specimenCount="1" typeStatus="holotype">
<typeStatus id="FE138815BA52FF83139ED0004B8FFB08" box="[386,495,1126,1152]" pageId="23" type="holotype">Holotype</typeStatus>
: Male (
<specimenCode id="710E9ECCBA52FF83105AD0004940FB08" box="[582,800,1126,1152]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="23">LBM 1430009132</specimenCode>
), dissected with appendages sealed in a glass slide and valves stored dry in a micropalaeontological cavity slide.
</materialsCitation>
</paragraph>
<paragraph id="211736B7BA52FF8312DBD0CB4831FB63" blockId="23.[151,1437,697,2045]" pageId="23">
<materialsCitation id="91C03CEABA52FF8312DBD0CB4831FB63" ID-GBIF-Occurrence="2313474990" collectionCode="LBM" pageId="23" specimenCode="LBM 1430009134" specimenCount="1" typeStatus="allotype">
<typeStatus id="FE138815BA52FF8312DBD0CB4B4DFB4F" box="[199,301,1197,1223]" pageId="23" type="allotype">Allotype</typeStatus>
: Female (
<specimenCode id="710E9ECCBA52FF831385D0CB4810FB4F" box="[409,624,1197,1223]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="23">LBM 1430009134</specimenCode>
), dissected with appendages sealed in a glass slide and valves stored dry in a micropalaeontological cavity slide.
</materialsCitation>
</paragraph>
<paragraph id="211736B7BA52FF8312DBD0934840FADE" blockId="23.[151,1437,697,2045]" pageId="23">
<materialsCitation id="91C03CEABA52FF8312DBD093487CFADE" ID-GBIF-Occurrence="2313474988" collectionCode="LBM" pageId="23" specimenCode="LBM 1430009131, LBM 1430009138" specimenCount="1" typeStatus="paratype">
<typeStatus id="FE138815BA52FF8312DBD0934B58FA87" box="[199,312,1269,1295]" pageId="23" type="paratype">Paratypes</typeStatus>
: Males (
<specimenCode id="710E9ECCBA52FF831382D0934813FA87" box="[414,627,1269,1295]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="23">LBM 1430009131</specimenCode>
and DB40054), dissected with appendages sealed in glass slides and valves stored dry in micropalaeontological cavity slides. Male (
<specimenCode id="710E9ECCBA52FF831142D17F4E51FABB" box="[862,1073,1305,1331]" collectionCode="LBM" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="23">LBM 1430009138</specimenCode>
) whole carapace, stored dry in a micropalaeontological cavity slide
</materialsCitation>
.
</paragraph>
<paragraph id="211736B7BA52FF8312DBD1064BDDFA49" blockId="23.[151,1437,697,2045]" pageId="23">
Females (
<collectionCode id="47B9AE72BA52FF83132FD1064B11FAF2" box="[307,369,1376,1402]" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="23">LBM</collectionCode>
1430009136 and DB40055), dissected with appendages sealed in glass slides and valves stored dry in micropalaeontological cavity slides. Female (
<collectionCode id="47B9AE72BA52FF8310C3D1E2497DFA16" box="[735,797,1412,1438]" country="Japan" httpUri="http://grbio.org/cool/pcbg-hhf5" name="Lake Biwa Museum" pageId="23">LBM</collectionCode>
1430009137), whole carapace, stored dry in a micropalaeontological cavity slide.
</paragraph>
<paragraph id="211736B7BA52FF8312DBD1AD49FDFA6D" blockId="23.[151,1437,697,2045]" box="[199,925,1483,1510]" pageId="23">
All type material collected from the
<typeStatus id="FE138815BA52FF831043D1AA48EFFA6E" box="[607,655,1484,1510]" pageId="23">type</typeStatus>
locality on
<date id="55161077BA52FF831108D1AD49F9FA6D" box="[788,921,1483,1509]" pageId="23" value="2011-07-04">4 July 2011</date>
.
</paragraph>
<paragraph id="211736B7BA52FF8312DBD1884BFAF9D8" blockId="23.[151,1437,697,2045]" pageId="23">
<emphasis id="13DCEAA5BA52FF8312DBD1884BE2F981" bold="true" box="[199,386,1518,1545]" pageId="23">Other material.</emphasis>
<bibRefCitation id="45394B46BA52FF831396D1894848F981" author="Okubo, I." box="[394,552,1519,1545]" pageId="23" pagination="46 - 50" refId="ref22322" refString="Okubo, I. (1974) Freshwater Ostracoda from Japan, XI. Ilyocypris angulata Sars, 1902. The Research Bulletin of Okayama Shujitsu Junior College, 4, 46 - 50." type="journal article" year="1974">Okubo (1974)</bibRefCitation>
stated that some specimens of the Japanese “
<taxonomicName id="E6A84D34BA52FF831636D1964EC6F981" authorityName="Sars" authorityYear="1903" box="[1066,1190,1519,1545]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="23" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="13DCEAA5BA52FF831636D1964EC6F981" box="[1066,1190,1519,1545]" italics="true" pageId="23">I. angulata</emphasis>
</taxonomicName>
” he studied would be deposited in the collections of the National Science Museum,
<collectingRegion id="E36CF855BA52FF831153D27449F9F9A4" box="[847,921,1554,1580]" country="Japan" name="Tokyo" pageId="23">Tokyo</collectingRegion>
, but enquiries revealed that the specimens are not in their collections.
</paragraph>
<paragraph id="211736B7BA52FF8312DBD23F4B86F875" blockId="23.[151,1437,697,2045]" pageId="23">
<emphasis id="13DCEAA5BA52FF8312DBD23F4B39F9FC" bold="true" box="[199,345,1625,1652]" pageId="23">Description.</emphasis>
Male carapace length
<quantity id="E6509B52BA52FF831041D23C4890F9FD" box="[605,752,1626,1653]" metricMagnitude="-1" metricUnit="m" metricValue="7.715000000000001" metricValueMax="7.970000000000001" metricValueMin="7.459999999999999" pageId="23" unit="mm" value="771.5" valueMax="797.0" valueMin="746.0">746797 µm</quantity>
, height
<quantity id="E6509B52BA52FF831150D23C49BFF9FD" box="[844,991,1626,1653]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" metricValueMax="4.36" metricValueMin="4.04" pageId="23" unit="mm" value="420.0" valueMax="436.0" valueMin="404.0">404436 µm</quantity>
, height/length 0.530.55. Female carapace length
<quantity id="E6509B52BA52FF83133CD2184BD2F910" box="[288,434,1662,1688]" metricMagnitude="-1" metricUnit="m" metricValue="8.684999999999999" metricValueMax="9.1" metricValueMin="8.27" pageId="23" unit="mm" value="868.5" valueMax="910.0" valueMin="827.0">827910 µm</quantity>
, height
<quantity id="E6509B52BA52FF831010D21B48FEF910" box="[524,670,1661,1688]" metricMagnitude="-1" metricUnit="m" metricValue="4.835" metricValueMax="5.04" metricValueMin="4.63" pageId="23" unit="mm" value="483.5" valueMax="504.0" valueMin="463.0">463504 µm</quantity>
, height/length 0.550.57. Anterior margin more inflated than posterior margin (
<figureCitation id="B9932A32BA52FF831339D2C74B18F934" box="[293,376,1697,1724]" captionStart="FIGURE 14" captionStartId="24.[151,250,1637,1662]" captionTargetBox="[286,1301,181,1611]" captionTargetId="figure@24.[286,1301,181,1611]" captionTargetPageId="24" captionText="FIGURE 14. Ilyocypris incus sp. nov. A, female left valve, external view (LBM 1430009136). B, female right valve, external view (LBM 1430009136). C, female left valve, internal view (LBM 1430009136). D, female right valve, internal view (LBM 1430009136). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009136). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009136). G, female carapace, dorsal view, anterior to right (LBM 1430009137). H, male right valve, external view (LBM1430009131). I, male carapace, dorsal view, anterior to right (LBM 1430009138). J, male right valve, ventral view (LBM 1430009133). K, male right valve, detail of ventral margin (LBM 1430009133). Scale bar = 500 µm for AD, GJ, 174 µm for E &amp; F, 135 µm for K. All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363561/files/figure.png" pageId="23">Fig. 14</figureCitation>
). Dorsal margin almost straight, sloping down posteriorly, maximum height anterior of muscle scars. Ventral margin strongly concave. Well-defined bifurcated sulcus near dorsal margin just anterior of adductor muscle scars, which form small rounded depression below posterior part of sulcus. Surface of valves densely covered with many small pits. Area near posterior margin with about seven tiny spines. Both valves with faint outer lists running near ventral margins (
<figureCitation id="B9932A32BA52FF831034D35648A5F8C3" box="[552,709,1840,1867]" captionStart="FIGURE 14" captionStartId="24.[151,250,1637,1662]" captionTargetBox="[286,1301,181,1611]" captionTargetId="figure@24.[286,1301,181,1611]" captionTargetPageId="24" captionText="FIGURE 14. Ilyocypris incus sp. nov. A, female left valve, external view (LBM 1430009136). B, female right valve, external view (LBM 1430009136). C, female left valve, internal view (LBM 1430009136). D, female right valve, internal view (LBM 1430009136). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009136). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009136). G, female carapace, dorsal view, anterior to right (LBM 1430009137). H, male right valve, external view (LBM1430009131). I, male carapace, dorsal view, anterior to right (LBM 1430009138). J, male right valve, ventral view (LBM 1430009133). K, male right valve, detail of ventral margin (LBM 1430009133). Scale bar = 500 µm for AD, GJ, 174 µm for E &amp; F, 135 µm for K. All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363561/files/figure.png" pageId="23">Fig. 14J &amp; K</figureCitation>
). Tubercles present in some specimens, one each side of sulcus (posterior one more developed), one slight one in mid position of sulcus, one in postero-ventral region and one in central ventral region. Inner calcified lamella narrow, especially posteriorly. Anterior inner calcified lamellae of both valves with lists. Postero-ventral region of calcified inner lamella of left valve with shallow depression containing 23 marginal ripplets (
<figureCitation id="B9932A32BA52FF8313A7D3D9487FF851" box="[443,543,1983,2009]" captionStart="FIGURE 14" captionStartId="24.[151,250,1637,1662]" captionTargetBox="[286,1301,181,1611]" captionTargetId="figure@24.[286,1301,181,1611]" captionTargetPageId="24" captionText="FIGURE 14. Ilyocypris incus sp. nov. A, female left valve, external view (LBM 1430009136). B, female right valve, external view (LBM 1430009136). C, female left valve, internal view (LBM 1430009136). D, female right valve, internal view (LBM 1430009136). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009136). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009136). G, female carapace, dorsal view, anterior to right (LBM 1430009137). H, male right valve, external view (LBM1430009131). I, male carapace, dorsal view, anterior to right (LBM 1430009138). J, male right valve, ventral view (LBM 1430009133). K, male right valve, detail of ventral margin (LBM 1430009133). Scale bar = 500 µm for AD, GJ, 174 µm for E &amp; F, 135 µm for K. All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363561/files/figure.png" pageId="23">Fig. 14E</figureCitation>
). Adductor muscle scars obscured. Dorsal view oval, with posterior margin more rounded than anterior margin.
</paragraph>
<caption id="75D7663FBA5DFF8C128BD2034ACAF8F2" httpUri="https://zenodo.org/record/3363561/files/figure.png" pageId="24" startId="24.[151,250,1637,1662]" targetBox="[286,1301,181,1611]" targetPageId="24">
<paragraph id="211736B7BA5DFF8C128BD2034ACAF8F2" blockId="24.[151,1437,1637,1914]" pageId="24">
<emphasis id="13DCEAA5BA5DFF8C128BD2034B42F9F6" bold="true" box="[151,290,1637,1662]" pageId="24">FIGURE 14.</emphasis>
<taxonomicName id="E6A84D34BA5DFF8C1336D2004BACF9F6" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[298,460,1638,1662]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="24" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA5DFF8C1336D2004BACF9F6" box="[298,460,1638,1662]" italics="true" pageId="24">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA5DFF8C13C9D2004845F9F5" box="[469,549,1638,1661]" pageId="24" rank="species">sp. nov.</taxonomicNameLabel>
A, female left valve, external view (LBM 1430009136). B, female right valve, external view (LBM 1430009136). C, female left valve, internal view (LBM 1430009136). D, female right valve, internal view (LBM 1430009136). E, female left valve, internal view, detail of postero-ventral margin (LBM 1430009136). F, female right valve, internal view, detail of antero-ventral margin (LBM 1430009136). G, female carapace, dorsal view, anterior to right (LBM 1430009137). H, male right valve, external view (LBM1430009131). I, male carapace, dorsal view, anterior to right (LBM 1430009138). J, male right valve, ventral view (LBM 1430009133). K, male right valve, detail of ventral margin (LBM 1430009133). Scale bar = 500 µm for AD, GJ, 174 µm for E &amp; F, 135 µm for K. All specimens from sample 14; see Table 1.
</paragraph>
</caption>
<caption id="75D7663FBA5CFF8D128BD3494B6EF850" httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="25" startId="25.[151,250,1839,1864]" targetBox="[238,1350,181,1814]" targetPageId="25">
<paragraph id="211736B7BA5CFF8D128BD3494B6EF850" blockId="25.[151,1437,1839,2008]" pageId="25">
<emphasis id="13DCEAA5BA5CFF8D128BD3494B41F8C0" bold="true" box="[151,289,1839,1864]" pageId="25">FIGURE 15.</emphasis>
<taxonomicName id="E6A84D34BA5CFF8D1335D3564BACF8C0" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[297,460,1840,1864]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="25" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA5CFF8D1335D3564BACF8C0" box="[297,460,1840,1864]" italics="true" pageId="25">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA5CFF8D13C8D3564845F8CF" box="[468,549,1840,1863]" pageId="25" rank="species">sp. nov.</taxonomicNameLabel>
A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1.
</paragraph>
</caption>
<paragraph id="211736B7BA5FFF8E12DBD4FF499AFEEF" blockId="26.[151,1437,152,1908]" pageId="26">
Antennule with seven articulated segments (
<figureCitation id="B9932A32BA5FFF8E10AAD4FF497FFF3B" box="[694,799,152,179]" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15A</figureCitation>
). First segment with two long sub-apical setae on ventral margin and one medium-length, stout seta on dorsal margin. Second segment with relatively long apical-dorsal seta. Third segment with one short apical-dorsal seta and one short apical-ventral seta. Fourth segment with two long apical-dorsal setae, and two shorter apical-ventral setae. Fifth segment with two long apical-dorsal setae, and two shorter apical-ventral setae. Sixth segment with four long setae and relatively long alpha seta. Terminal segment apically with two short setae, one long seta and relatively short aesthetasc ya.
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD517483FFD93" blockId="26.[151,1437,152,1908]" pageId="26">
Male antennae with fourth segment undivided (
<figureCitation id="B9932A32BA5FFF8E10FED5174927FE03" box="[738,839,368,395]" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15B</figureCitation>
&amp;
<figureCitation id="B9932A32BA5FFF8E1171D51749FFFE04" box="[877,927,369,396]" captionStart="FIGURE 16" captionStartId="27.[151,250,1671,1696]" captionTargetBox="[219,1372,181,1646]" captionTargetId="figure@27.[215,1372,181,1646]" captionTargetPageId="27" captionText="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="26">16B</figureCitation>
). Swimming setae very long, extending considerably beyond end of claws; shorter accompanying seta also long, reaching to mid-point of claws. G3 reduced to short seta, extending to distal end of terminal segment. Claw z1 small and slender, not reaching to end of claw G2. Female antenna with G2 claw as long as claws G1 and G3 (
<figureCitation id="B9932A32BA5FFF8E1130D5BB49F4FE70" box="[812,916,477,504]" captionStart="FIGURE 16" captionStartId="27.[151,250,1671,1696]" captionTargetBox="[219,1372,181,1646]" captionTargetId="figure@27.[215,1372,181,1646]" captionTargetPageId="27" captionText="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="26">Fig. 16A</figureCitation>
). Seta z1 stout, claw-like, shorter than setae z2 and z3, approximately half length of G2.
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD6434AB4FD7B" blockId="26.[151,1437,152,1908]" pageId="26">
Mandibular coxa small, with well-developed teeth on endite (
<figureCitation id="B9932A32BA5FFF8E1169D64349BDFDB7" box="[885,989,548,575]" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15D</figureCitation>
). Mandibular palp four-segmented (
<figureCitation id="B9932A32BA5FFF8E1773D6434AB1FDEB" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15 C</figureCitation>
). Alpha seta on first segment small and slender. Second segment with two setae on outer edge, and 3+1+beta setae on inner edge. Beta seta short, stubby and hirsute. Third segment with four setae on outer apical corner, three setae on inner apical corner, and one apical seta between other two groups. Terminal segment with three stout apical setae, two thinner and shorter apical seta and one seta protruding from area proximal of mid-length, towards outer edge.
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD69B4EBFFCD7" blockId="26.[151,1437,152,1908]" pageId="26">
Maxillula palp first segment with three sub-equal medium-length setae on outer apical corner, one mediumlength seta positioned on apical edge offset from apical corner, and one sub-apical setae in mid-width position (
<figureCitation id="B9932A32BA5FFF8E1773D7474AA8FCD7" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15E</figureCitation>
). Second segment wider than long, apically with three claw-like setae and three shorter setae.
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD70F4891FB9B" blockId="26.[151,1437,152,1908]" pageId="26">
Female fifth limb with large basis supporting two a-setae and approximately 17 setae of various lengths on inner edge (endite) (
<figureCitation id="B9932A32BA5FFF8E1320D7EB4BC0FC2F" box="[316,416,908,935]" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15F</figureCitation>
). Palp with two segments, terminating with one long and two mid-length setae. Branchial plate obscured, but with at least five rays. Male palp with long, thin proximal segment, supporting distally two setae and large grasping hook. Left and right hooks similar in morphology, gently curved and tapering to sharp tips, with short, distally very fine seta on outer edge (
<figureCitation id="B9932A32BA5FFF8E1060D79F4884FB9B" box="[636,740,1016,1043]" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15H</figureCitation>
).
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD07B4899FB4F" blockId="26.[151,1437,152,1908]" pageId="26">
Sixth limb four-segmented (
<figureCitation id="B9932A32BA5FFF8E13E3D07B4806FBBF" box="[511,614,1052,1079]" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15G</figureCitation>
). First segment (= protopodite) with short d1 seta. Second segment (= first endopodal segment) with short e seta. Third segment (= second endopodal segment) with short f seta protruding from mid-length and short apical g seta. Terminal segment (= third endopodal segment) with relatively long h1 seta, short h3 seta and well-developed h2 claw, approximately 1.9x length of anterior sclerotized margin of first endopodal segment (marked with black triangle on
<figureCitation id="B9932A32BA5FFF8E1099D0CB488CFB4F" box="[645,748,1196,1223]" captionStart="FIGURE 15" captionStartId="25.[151,250,1839,1864]" captionTargetBox="[238,1350,181,1814]" captionTargetId="figure@25.[238,1350,181,1814]" captionTargetPageId="25" captionText="FIGURE 15. Ilyocypris incus sp. nov. A, antennule, male (DB40054). B, antenna, male (LBM 1430009132). C, mandibular palp, female (LBM 1430009136). D, mandibular coxa, female (LBM 1430009134). E, maxillula palp and endites (setae on endites not drawn), female (LBM 1430009136). F, fifth limb, female (DB40055). G, sixth limb, male (LBM 1430009132), arrow and dashed line indicate length of first endopodal segment. H, fifth limb palp, male (DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363563/files/figure.png" pageId="26">Fig. 15G</figureCitation>
).
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD0B74EE4FABB" blockId="26.[151,1437,152,1908]" pageId="26">
Seventh limb with four segments (
<figureCitation id="B9932A32BA5FFF8E1057D0B748D2FB64" box="[587,690,1233,1260]" captionStart="FIGURE 16" captionStartId="27.[151,250,1671,1696]" captionTargetBox="[219,1372,181,1646]" captionTargetId="figure@27.[215,1372,181,1646]" captionTargetPageId="27" captionText="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="26">Fig. 16C</figureCitation>
). First segment with seta d1. Second segment with seta e reaching to approximately mid-length of third segment. Fourth segment with two setae on anterior edge (f and g), distal-most one longest. Fourth segment small, with seta h3 slightly longer than h1, and seta h2 short.
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD15B4BE9FAF3" blockId="26.[151,1437,152,1908]" pageId="26">
Caudal ramus typical of genus, ramus distinctly curved distal of mid-length (
<figureCitation id="B9932A32BA5FFF8E1604D15B4E1FFAD0" box="[1048,1151,1341,1368]" captionStart="FIGURE 16" captionStartId="27.[151,250,1671,1696]" captionTargetBox="[219,1372,181,1646]" captionTargetId="figure@27.[215,1372,181,1646]" captionTargetPageId="27" captionText="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="26">Fig. 16D</figureCitation>
). Claw Gp slightly longer than Ga, seta sp long.
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD1E34F10FA4B" blockId="26.[151,1437,152,1908]" pageId="26">
Female reproductive organ with large, rounded genital lobe anteriorly, and smaller rounded lobe posteriorly, just anterior of base of caudal ramus (
<figureCitation id="B9932A32BA5FFF8E1012D1CF4814FA4C" box="[526,628,1449,1476]" captionStart="FIGURE 16" captionStartId="27.[151,250,1671,1696]" captionTargetBox="[219,1372,181,1646]" captionTargetId="figure@27.[215,1372,181,1646]" captionTargetPageId="27" captionText="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="26">Fig. 16E</figureCitation>
). Organ delicate and easily distorted or damaged during dissections.
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD1AB4988F9F0" blockId="26.[151,1437,152,1908]" pageId="26">
Hemipenes, outer lobe unevenly rounded distally, nearly touching to slightly overlapping middle lobe (
<figureCitation id="B9932A32BA5FFF8E175AD1AB4FF8FA6F" box="[1350,1432,1484,1511]" captionStart="FIGURE 5" captionStartId="9.[151,250,1475,1500]" captionTargetBox="[151,1436,181,1446]" captionTargetId="figure@9.[151,1436,181,1449]" captionTargetPageId="9" captionText="FIGURE 5. Light microscopy photographs of Ilyocypris hemipenes. A, Ilyocypris japonica Okubo, 1990 (LBM 1430009150, sample 18; see Table 1). B, Ilyocypris japonica Okubo, 1990 (LBM 1430009151, sample 18). C, Ilyocypris mongolica Martens, 1991 (LBM 1430009161, sample 17). D, Ilyocypris dentifera Sars, 1903 (LBM 1430009119, sample 6). E, Ilyocypris angulata Sars, 1903 (LBM 1430009112, sample 20). F, Ilyocypris incus sp. nov. (DB40054, sample 14). E has been reflected on the vertical axis to facilitate comparisons." httpUri="https://zenodo.org/record/3363543/files/figure.png" pageId="26">Fig. 5F</figureCitation>
,
<figureCitation id="B9932A32BA5FFF8E128BD1974B62F983" box="[151,258,1521,1547]" captionStart="FIGURE 16" captionStartId="27.[151,250,1671,1696]" captionTargetBox="[219,1372,181,1646]" captionTargetId="figure@27.[215,1372,181,1646]" captionTargetPageId="27" captionText="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="26">16F &amp; H</figureCitation>
). Middle lobe triangular, inner corner sharply angled, outer edge more rounded. Inner lobe longer than outer lobe, with inner edge slightly curving along most of length, before smoothly curving to bulbous distal end; bulbous end projecting towards middle lobe, sometimes slightly overlapping. Copulatory process well developed, with large, roughly triangular distal end with uneven distal edge (
<figureCitation id="B9932A32BA5FFF8E116FD23B49BBF9F0" box="[883,987,1629,1656]" captionStart="FIGURE 16" captionStartId="27.[151,250,1671,1696]" captionTargetBox="[219,1372,181,1646]" captionTargetId="figure@27.[215,1372,181,1646]" captionTargetPageId="27" captionText="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="26">Fig. 16G</figureCitation>
).
</paragraph>
<paragraph id="211736B7BA5FFF8E12EBD2E74A9BF937" blockId="26.[151,1437,152,1908]" pageId="26">
Zenker organ with 1819 internal rosettes (
<figureCitation id="B9932A32BA5FFF8E10C6D2E74952F914" box="[730,818,1665,1692]" captionStart="FIGURE 6" captionStartId="10.[151,250,1729,1754]" captionTargetBox="[151,1436,345,1704]" captionTargetId="figure@10.[151,1436,345,1704]" captionTargetPageId="10" captionText="FIGURE 6. Light microscopy photographs of Ilyocypris Zenker organs. Black triangles mark the anterior ends. A, Ilyocypris japonica Okubo, 1990 (LBM 1430009144, sample 9; see Table 1). B, Ilyocypris dentifera Sars, 1903 (LBM 1430009119, sam- ple 6). C, Ilyocypris incus sp. nov. (LBM 1430009131, sample 14)." httpUri="https://zenodo.org/record/3363545/files/figure.png" pageId="26">Fig. 6C</figureCitation>
). Proximal end oval and small, slightly wider than central tube.
</paragraph>
<paragraph id="211736B7BA5FFF8E12DBD2AE4F56F8FC" blockId="26.[151,1437,152,1908]" pageId="26">
<emphasis id="13DCEAA5BA5FFF8E12DBD2AE4BBAF96B" bold="true" box="[199,474,1736,1763]" pageId="26">Intraspecific variation.</emphasis>
Some specimens have tubercles developed on the carapace to varying degrees, while others have no obvious tubercles.
<bibRefCitation id="45394B46BA5FFF8E1013D28B48C6F88F" author="Okubo, I." box="[527,678,1773,1799]" pageId="26" pagination="46 - 50" refId="ref22322" refString="Okubo, I. (1974) Freshwater Ostracoda from Japan, XI. Ilyocypris angulata Sars, 1902. The Research Bulletin of Okayama Shujitsu Junior College, 4, 46 - 50." type="journal article" year="1974">Okubo (1974</bibRefCitation>
; as
<taxonomicName id="E6A84D34BA5FFF8E10D2D2884928F88F" authorityName="Sars" authorityYear="1903" box="[718,840,1773,1799]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="26" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="13DCEAA5BA5FFF8E10D2D2884928F88F" box="[718,840,1773,1799]" italics="true" pageId="26">I. angulata</emphasis>
</taxonomicName>
) figured specimens with five tubercles on each valve. Seta sa on the caudal ramus is variable in length, with that of some specimens up to 157% longer compared with the holotypes. The inner lobe of the hemipenis is rather variable in appearance distally, sometimes with a slight gap between it and the middle lobe, while in other specimens slightly overlapping the middle lobe (
<figureCitation id="B9932A32BA5FFF8E16DDD33F4F4AF8FC" box="[1217,1322,1881,1908]" captionStart="FIGURE 16" captionStartId="27.[151,250,1671,1696]" captionTargetBox="[219,1372,181,1646]" captionTargetId="figure@27.[215,1372,181,1646]" captionTargetPageId="27" captionText="FIGURE 16. Ilyocypris incus sp. nov. A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1." httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="26">Fig. 16H</figureCitation>
).
</paragraph>
<caption id="75D7663FBA5EFF8F128BD2E14EE9F8B8" httpUri="https://zenodo.org/record/3363565/files/figure.png" pageId="27" startId="27.[151,250,1671,1696]" targetBox="[219,1372,181,1646]" targetPageId="27">
<paragraph id="211736B7BA5EFF8F128BD2E14EE9F8B8" blockId="27.[151,1436,1671,1840]" pageId="27">
<emphasis id="13DCEAA5BA5EFF8F128BD2E14B44F928" bold="true" box="[151,292,1671,1696]" pageId="27">FIGURE 16.</emphasis>
<taxonomicName id="E6A84D34BA5EFF8F1332D2EE4BB2F928" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[302,466,1672,1696]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="27" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA5EFF8F1332D2EE4BB2F928" box="[302,466,1672,1696]" italics="true" pageId="27">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA5EFF8F13C1D2EF4850F928" box="[477,560,1673,1696]" pageId="27" rank="species">sp. nov.</taxonomicNameLabel>
A, detail of female antenna (LBM 1430009134). B, detail of male antenna (LBM 1430009132). C, seventh limb, male (LBM 1430009132). D, caudal ramus, male (DB40054). E, outline of female reproductive organ, arrow indicates anterior (LBM 1430009134). F, hemipenis (LBM 1430009132). G, copulatory process of hemipenis (LBM 1430009132). H, outlines of the hemipenes of four specimens (left to right LBM 1430009132, LBM 1430009135, LBM 1430009133 (reflected on the vertical axis), DB40054). All specimens from sample 14; see Table 1.
</paragraph>
</caption>
<paragraph id="211736B7BA5EFF8812DBD3E54B3DFF5F" blockId="27.[151,1437,1922,2021]" lastBlockId="28.[151,1437,153,1763]" lastPageId="28" pageId="27">
<emphasis id="13DCEAA5BA5EFF8F12DBD3E54B5BF815" bold="true" box="[199,315,1923,1949]" pageId="27">Remarks.</emphasis>
This species is very similar to
<taxonomicName id="E6A84D34BA5EFF8F108DD3E5496DF814" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[657,781,1922,1948]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="27" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA5EFF8F108DD3E5496DF814" box="[657,781,1922,1948]" italics="true" pageId="27">I. dentifera</emphasis>
</taxonomicName>
,
<taxonomicName id="E6A84D34BA5EFF8F1106D3E549E3F814" authorityName="Karanovic &amp; Lee" authorityYear="2013" box="[794,899,1922,1948]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="27" phylum="Arthropoda" rank="species" species="hanguk">
<emphasis id="13DCEAA5BA5EFF8F1106D3E549E3F814" box="[794,899,1922,1948]" italics="true" pageId="27">I. hanguk</emphasis>
</taxonomicName>
,
<emphasis id="13DCEAA5BA5EFF8F118CD3E549B1F814" box="[912,977,1922,1948]" italics="true" pageId="27">
<taxonomicName id="E6A84D34BA5EFF8F118CD3E549ADF814" authorityName="Ma &amp; Yu" authorityYear="2018" box="[912,973,1922,1948]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="27" phylum="Arthropoda" rank="species" species="dui">I. dui</taxonomicName>
,
</emphasis>
and
<taxonomicName id="E6A84D34BA5EFF8F161BD3E54E3DF814" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[1031,1117,1922,1948]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="27" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="13DCEAA5BA5EFF8F161BD3E54E3DF814" box="[1031,1117,1922,1948]" italics="true" pageId="27">I. gibba</emphasis>
</taxonomicName>
. The valves are more round- ed posteriorly than those of
<taxonomicName id="E6A84D34BA5EFF8F13D7D3C14827F848" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[459,583,1958,1984]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="27" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA5EFF8F13D7D3C14827F848" box="[459,583,1958,1984]" italics="true" pageId="27">I. dentifera</emphasis>
</taxonomicName>
, which tends to have a more truncated posterior. The hemipenes demonstrate several differences, including the shapes of the outer and inner lobes, and the copulatory process. Our specimens are a close match to the incorrect records of
<taxonomicName id="E6A84D34BA59FF881048D4FC48AEFF3B" authorityName="Sars" authorityYear="1903" box="[596,718,153,179]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="13DCEAA5BA59FF881048D4FC48AEFF3B" box="[596,718,153,179]" italics="true" pageId="28">I. angulata</emphasis>
</taxonomicName>
and
<taxonomicName id="E6A84D34BA59FF88111DD4FC491CFF3B" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[769,892,153,179]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA59FF88111DD4FC491CFF3B" box="[769,892,153,179]" italics="true" pageId="28">I. dentifera</emphasis>
</taxonomicName>
figured by
<bibRefCitation id="45394B46BA59FF8811E6D4FF4EF0FF3B" author="Okubo, I." box="[1018,1168,153,179]" pageId="28" pagination="46 - 50" refId="ref22322" refString="Okubo, I. (1974) Freshwater Ostracoda from Japan, XI. Ilyocypris angulata Sars, 1902. The Research Bulletin of Okayama Shujitsu Junior College, 4, 46 - 50." type="journal article" year="1974">Okubo (1974</bibRefCitation>
;
<bibRefCitation id="45394B46BA59FF881687D4FF4EB7FF3B" author="Okubo I." box="[1179,1239,153,179]" pageId="28" refId="ref22386" refString="Okubo I. (2004) Nihon tansui san kaimijinko rui ni tsuite. Kabushikigaisha Sanmon Insatsusho, Okayama, 72 pp. [In Japanese, privately published]" type="book" year="2004">2004</bibRefCitation>
), and we consider them conspecific.
</paragraph>
<paragraph id="211736B7BA59FF8812DBD4874EFCFE5B" blockId="28.[151,1437,153,1763]" pageId="28">
The misidentified
<taxonomicName id="E6A84D34BA59FF881388D4874810FF73" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[404,624,225,251]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA59FF881388D4874810FF73" box="[404,624,225,251]" italics="true" pageId="28">Ilyocypris dentifera</emphasis>
</taxonomicName>
of
<bibRefCitation id="45394B46BA59FF88108AD4874918FF73" author="Kim, W. &amp; Min, G. S." box="[662,888,225,251]" pageId="28" pagination="307 - 322" refId="ref21073" refString="Kim, W. &amp; Min, G. S. (1991 a) Redescription of four Recent freshwater ostracod (Crustacea: Ostracoda) species from Korea. Korean Journal of Zoology, 34, 307 - 322." type="journal article" year="1991">Kim &amp; Min (1991a)</bibRefCitation>
comes close to this species, but the outer lobe of the hemipenis has a slightly different shape and the claw of their figured sixth limb is robustly serrated. The outer lobe of the hemipenis does vary somewhat in our material, and so the hemipenis morphology recorded by Kim &amp; Min (op. cit.) could represent the same species. The robust serration on the sixth limb is unusual for
<taxonomicName id="E6A84D34BA59FF8816EBD52B4F07FEEF" authorityName="Zenker" box="[1271,1383,333,359]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="genus">
<emphasis id="13DCEAA5BA59FF8816EBD52B4F07FEEF" box="[1271,1383,333,359]" italics="true" pageId="28">Ilyocypris</emphasis>
</taxonomicName>
species generally, and maybe a result of Kim &amp; Mins drawing style; other species figured by them also have robust, rather exaggerated serration on the sixth limbs (
<bibRefCitation id="45394B46BA59FF8810B2D5F349E3FE27" author="Kim, W. &amp; Min, G. S." box="[686,899,405,431]" pageId="28" pagination="307 - 322" refId="ref21073" refString="Kim, W. &amp; Min, G. S. (1991 a) Redescription of four Recent freshwater ostracod (Crustacea: Ostracoda) species from Korea. Korean Journal of Zoology, 34, 307 - 322." type="journal article" year="1991">Kim &amp; Min 1991a</bibRefCitation>
;
<bibRefCitation id="45394B46BA59FF881193D5F349B9FE38" author="Kim, W. &amp; Min, G. S." box="[911,985,405,432]" pageId="28" pagination="81 - 100" refId="ref21114" refString="Kim, W. &amp; Min, G. S. (1991 b) Redescription of Recent freshwater ostracods (Crustacea: Ostracoda) from Korea: A systematic study of four species of family Cyprididae. The Korean Journal of Systematic Zoology, 7, 81 - 100." type="journal article" year="1991">1991b</bibRefCitation>
). We consider the records of
<taxonomicName id="E6A84D34BA59FF88173CD5F04FFCFE27" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[1312,1436,405,431]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA59FF88173CD5F04FFCFE27" box="[1312,1436,405,431]" italics="true" pageId="28">I. dentifera</emphasis>
</taxonomicName>
of
<bibRefCitation id="45394B46BA59FF8812AAD5DF4BF8FE5B" author="Kim, W. &amp; Min, G. S." box="[182,408,441,467]" pageId="28" pagination="307 - 322" refId="ref21073" refString="Kim, W. &amp; Min, G. S. (1991 a) Redescription of four Recent freshwater ostracod (Crustacea: Ostracoda) species from Korea. Korean Journal of Zoology, 34, 307 - 322." type="journal article" year="1991">Kim &amp; Min (1991a)</bibRefCitation>
at least in part, as possibly conspecific with
<taxonomicName id="E6A84D34BA59FF881191D5DF4E5FFE5B" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[909,1087,441,467]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF881191D5DF4E5FFE5B" box="[909,1087,441,467]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF88165AD5DF4EFCFE5B" box="[1094,1180,441,467]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="211736B7BA59FF8812DBD5BB49B2FD47" blockId="28.[151,1437,153,1763]" pageId="28">
<taxonomicName id="E6A84D34BA59FF8812DBD5BB4BF1FE7F" authorityName="Karanovic &amp; Lee" authorityYear="2013" box="[199,401,477,503]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="hanguk">
<emphasis id="13DCEAA5BA59FF8812DBD5BB4BF1FE7F" box="[199,401,477,503]" italics="true" pageId="28">Ilyocypris hanguk</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="E6A84D34BA59FF881039D5BB48B7FE7F" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[549,727,477,503]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF881039D5BB48B7FE7F" box="[549,727,477,503]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF8810C3D5BB4958FE7F" box="[735,824,477,503]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
, but the posterior margin of the male carapace is more rounded in the latter (adult female morphology of
<taxonomicName id="E6A84D34BA59FF8810CED664495CFD93" authorityName="Karanovic &amp; Lee" authorityYear="2013" box="[722,828,513,539]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="hanguk">
<emphasis id="13DCEAA5BA59FF8810CED664495CFD93" box="[722,828,513,539]" italics="true" pageId="28">I. hanguk</emphasis>
</taxonomicName>
remains unknown). The three lobes on the hemipenis are different in shape compared with those of
<taxonomicName id="E6A84D34BA59FF88108BD6434929FDB7" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[663,841,549,575]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF88108BD6434929FDB7" box="[663,841,549,575]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF881153D64349C8FDB7" box="[847,936,549,575]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
, in particular, the middle lobe of
<taxonomicName id="E6A84D34BA59FF881705D6404FE2FDB7" authorityName="Karanovic &amp; Lee" authorityYear="2013" box="[1305,1410,549,575]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="hanguk">
<emphasis id="13DCEAA5BA59FF881705D6404FE2FDB7" box="[1305,1410,549,575]" italics="true" pageId="28">I. hanguk</emphasis>
</taxonomicName>
is proximally wider and the copulatory process is missing the large distal projections found in
<taxonomicName id="E6A84D34BA59FF88168CD62F4F21FDEB" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[1168,1345,585,611]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF88168CD62F4F21FDEB" box="[1168,1345,585,611]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF88175BD62F4FFDFDEB" box="[1351,1437,585,611]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
Other features of
<taxonomicName id="E6A84D34BA59FF881347D6084BA5FD0F" authorityName="Karanovic &amp; Lee" authorityYear="2013" box="[347,453,621,647]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="hanguk">
<emphasis id="13DCEAA5BA59FF881347D6084BA5FD0F" box="[347,453,621,647]" italics="true" pageId="28">I. hanguk</emphasis>
</taxonomicName>
that are different include the very small h1 seta on the walking leg (note that the h1 and h3 setae of the sixth limbs are mislabelled in
<bibRefCitation id="45394B46BA59FF881096D6F749EDFD23" author="Karanovic, I. &amp; Lee, W." box="[650,909,657,683]" pageId="28" pagination="39 - 71" refId="ref20665" refString="Karanovic, I. &amp; Lee, W. (2013) On the ostracod genus Ilyocypris, with description of one new species from Korea and the first report of males of I. bradyi (Crustacea: Ostracoda: Podocopida). Proceedings of the Biological Society of Washington, 126, 39 - 71. https: // doi. org / 10.2988 / 0006 - 324 X- 126.1.39" type="journal article" year="2013">Karanovic &amp; Lee 2013</bibRefCitation>
), and the large number of rosettes in the Zenker organ (approximately
<quantity id="E6509B52BA59FF88138CD6D24BA9FD47" box="[400,457,692,719]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="28" unit="in" value="25.0">25 in</quantity>
<taxonomicName id="E6A84D34BA59FF8813D3D6D04859FD47" authorityName="Karanovic &amp; Lee" authorityYear="2013" box="[463,569,693,719]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="hanguk">
<emphasis id="13DCEAA5BA59FF8813D3D6D04859FD47" box="[463,569,693,719]" italics="true" pageId="28">I. hanguk</emphasis>
</taxonomicName>
,
<quantity id="E6509B52BA59FF88105BD6D348CAFD47" box="[583,682,693,719]" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="4.826" metricValueMin="4.572" pageId="28" unit="in" value="18.5" valueMax="19.0" valueMin="18.0">1819 in</quantity>
<taxonomicName id="E6A84D34BA59FF8810ADD6D34903FD47" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[689,867,693,719]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF8810ADD6D34903FD47" box="[689,867,693,719]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF881176D6D349A2FD47" box="[874,962,693,719]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="211736B7BA59FF8812DBD6BF48B4FCD7" blockId="28.[151,1437,153,1763]" pageId="28">
Similar to
<taxonomicName id="E6A84D34BA59FF881320D6BC4BC5FD7B" authorityName="Karanovic &amp; Lee" authorityYear="2013" box="[316,421,729,755]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="hanguk">
<emphasis id="13DCEAA5BA59FF881320D6BC4BC5FD7B" box="[316,421,729,755]" italics="true" pageId="28">I. hanguk</emphasis>
</taxonomicName>
, the carapace of
<taxonomicName id="E6A84D34BA59FF88107DD6BC48FBFD7B" authorityName="Ma &amp; Yu" authorityYear="2018" box="[609,667,729,755]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dui">
<emphasis id="13DCEAA5BA59FF88107DD6BC48FBFD7B" box="[609,667,729,755]" italics="true" pageId="28">I. dui</emphasis>
</taxonomicName>
resembles that of
<taxonomicName id="E6A84D34BA59FF88117AD6BC4981FD7B" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[870,993,729,755]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA59FF88117AD6BC4981FD7B" box="[870,993,729,755]" italics="true" pageId="28">I. dentifera</emphasis>
</taxonomicName>
more than
<taxonomicName id="E6A84D34BA59FF881642D6BF4F6FFD7B" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[1118,1295,729,755]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF881642D6BF4F6FFD7B" box="[1118,1295,729,755]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF88170AD6BF4F0CFD7B" box="[1302,1388,729,755]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
The middle lobe of the hemipenis of
<taxonomicName id="E6A84D34BA59FF881014D6984824FC9F" authorityName="Ma &amp; Yu" authorityYear="2018" box="[520,580,765,791]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dui">
<emphasis id="13DCEAA5BA59FF881014D6984824FC9F" box="[520,580,765,791]" italics="true" pageId="28">I. dui</emphasis>
</taxonomicName>
is most similar to that of
<taxonomicName id="E6A84D34BA59FF881174D69849B2FC9F" authorityName="Karanovic &amp; Lee" authorityYear="2013" box="[872,978,765,791]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="hanguk">
<emphasis id="13DCEAA5BA59FF881174D69849B2FC9F" box="[872,978,765,791]" italics="true" pageId="28">I. hanguk</emphasis>
</taxonomicName>
in that it is proximally wide, which also distinguishes it from
<taxonomicName id="E6A84D34BA59FF88139AD7474858FCB3" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[390,568,801,827]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF88139AD7474858FCB3" box="[390,568,801,827]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF881023D74748F6FCB3" box="[575,662,801,827]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
The outer and inner lobes and the copulatory process are also differently shaped than those of
<taxonomicName id="E6A84D34BA59FF8813DFD7234815FCD7" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[451,629,837,863]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF8813DFD7234815FCD7" box="[451,629,837,863]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF881060D72348B4FCD7" box="[636,724,837,863]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA59FF8812DBD70F4BD1FA17" pageId="28" type="reference_group">
<paragraph id="211736B7BA59FF8812DBD70F4EF9FB2B" blockId="28.[151,1437,153,1763]" pageId="28">
<taxonomicName id="E6A84D34BA59FF8812DBD70F4B1FFC0B" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[199,383,873,899]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="13DCEAA5BA59FF8812DBD70F4B1FFC0B" box="[199,383,873,899]" italics="true" pageId="28">Ilyocypris gibba</emphasis>
</taxonomicName>
has a similar outline in lateral view to
<taxonomicName id="E6A84D34BA59FF881126D70F498DFC0B" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[826,1005,873,899]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF881126D70F498DFC0B" box="[826,1005,873,899]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF8811E8D70F4E2CFC0B" box="[1012,1100,873,899]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
and overall the differences in carapace morphology are subtle.
<taxonomicName id="E6A84D34BA59FF88100ED7EB48ABFC2F" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[530,715,909,935]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="13DCEAA5BA59FF88100ED7EB48ABFC2F" box="[530,715,909,935]" italics="true" pageId="28">Ilyocypris gibba</emphasis>
</taxonomicName>
has more ripplets (610) on the postero-ventral calcified inner lamella of the left valve than
<taxonomicName id="E6A84D34BA59FF8813FFD7D748F6FC43" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[483,662,945,971]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF8813FFD7D748F6FC43" box="[483,662,945,971]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF881082D7D74895FC43" box="[670,757,945,971]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
(23), but the numbers of ripplets are known to vary within species and are not considered to be a reliable taxonomic feature (
<bibRefCitation id="45394B46BA59FF881177D7B34E28FC67" author="Mazzini, I. &amp; Gliozzi, E. &amp; Rossetti, G. &amp; Pieri, V." box="[875,1096,981,1007]" pageId="28" pagination="1 - 14" refId="ref21849" refString="Mazzini, I., Gliozzi, E., Rossetti, G. &amp; Pieri, V. (2014) The Ilyocypris puzzle: A multidisciplinary approach to the study of phenotypic variability. International Review of Hydrobiology, 99, 1 - 14. https: // doi. org / 10.1002 / iroh. 201301729" type="journal article" year="2014">
Mazzini
<emphasis id="13DCEAA5BA59FF8811D2D7B0499EFC67" box="[974,1022,981,1007]" italics="true" pageId="28">et al</emphasis>
. 2014
</bibRefCitation>
). The pitting on the carapace is coarser and the postero-dorsal corner more angular in
<taxonomicName id="E6A84D34BA59FF8810EED79C4929FB9B" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[754,841,1017,1043]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="13DCEAA5BA59FF8810EED79C4929FB9B" box="[754,841,1017,1043]" italics="true" pageId="28">I. gibba</emphasis>
</taxonomicName>
compared with
<taxonomicName id="E6A84D34BA59FF8811E0D79F4ECEFB9B" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[1020,1198,1017,1043]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF8811E0D79F4ECEFB9B" box="[1020,1198,1017,1043]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF8816A8D79F4F6BFB9B" box="[1204,1291,1017,1043]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
The appendages are also very similar in the two species, with only the hemipenes and Zenker organs showing clear differences; the inner lobe of the hemipenis of
<taxonomicName id="E6A84D34BA59FF881004D024480FFBD3" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[536,623,1089,1115]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="13DCEAA5BA59FF881004D024480FFBD3" box="[536,623,1089,1115]" italics="true" pageId="28">I. gibba</emphasis>
</taxonomicName>
is much shorter and the outer lobe more angular than those of
<taxonomicName id="E6A84D34BA59FF881730D0274AB2FBF7" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF881730D0274AB2FBF7" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF8812C5D0034B50FBF7" box="[217,304,1125,1151]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
(
<bibRefCitation id="45394B46BA59FF881322D0034BB6FBF7" author="Meisch C." box="[318,470,1125,1151]" pageId="28" refId="ref21908" refString="Meisch C. (2000) Freshwater Ostracoda of Western and Central Europe. Susswasserfauna von Mitteleuropa 8 / 3. Spektrum Akademischer Verlag, Heidelberg, Berlin, 522 pp." type="book" year="2000">Meisch 2000</bibRefCitation>
). The shape of the copulatory process of
<taxonomicName id="E6A84D34BA59FF8811BDD0004998FBF7" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[929,1016,1125,1151]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="13DCEAA5BA59FF8811BDD0004998FBF7" box="[929,1016,1125,1151]" italics="true" pageId="28">I. gibba</emphasis>
</taxonomicName>
remains unknown. The Zenker organ of
<taxonomicName id="E6A84D34BA59FF8812AAD0EC4B6DFB2B" baseAuthorityName="Ramdohr" baseAuthorityYear="1808" box="[182,269,1161,1187]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="gibba">
<emphasis id="13DCEAA5BA59FF8812AAD0EC4B6DFB2B" box="[182,269,1161,1187]" italics="true" pageId="28">I. gibba</emphasis>
</taxonomicName>
has a more inflated proximal end compared with that of
<taxonomicName id="E6A84D34BA59FF881195D0EF4E5BFB2B" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[905,1083,1161,1187]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF881195D0EF4E5BFB2B" box="[905,1083,1161,1187]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF88165ED0EF4EF9FB2B" box="[1090,1177,1161,1187]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="211736B7BA59FF8812DBD0CB4BD1FA17" blockId="28.[151,1437,153,1763]" pageId="28">
Several species known from
<collectingCountry id="59BF7627BA59FF88100DD0CB4834FB4F" box="[529,596,1197,1223]" name="China" pageId="28">China</collectingCountry>
dont have appendages described, so comparisons with these species rely solely on the carapace morphology.
<taxonomicName id="E6A84D34BA59FF881029D0B749D8FB63" authority="Yang, 1982" authorityName="Yang" authorityYear="1982" box="[565,952,1233,1259]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="xizangensis">
<emphasis id="13DCEAA5BA59FF881029D0B7494FFB63" box="[565,815,1233,1259]" italics="true" pageId="28">Ilyocypris xizangensis</emphasis>
Yang, 1982
</taxonomicName>
(in
<bibRefCitation id="45394B46BA59FF8811F5D0B74ED8FB63" author="Huang, B. R. &amp; Yang, H. R. &amp; You, K. Y." box="[1001,1208,1233,1259]" pageId="28" pagination="326 - 348" refId="ref20287" refString="Huang, B. R., Yang, H. R. &amp; You, K. Y. (1982) Pliocene and Quaternary Ostracoda of southern and southwestern Tibet. In: Investigation Group of Qinghai-Tibet Plateau of Chinese Academy of Sciences (Eds), Paleontology of Tibet, Vol. 4. Science Press, Beijing, pp. 326 - 348. [in Chinese]" type="book chapter" year="1982">
Huang
<emphasis id="13DCEAA5BA59FF881621D0B44E17FB63" box="[1085,1143,1233,1259]" italics="true" pageId="28">et al.</emphasis>
1982
</bibRefCitation>
) is noticeably more elongate, and
<taxonomicName id="E6A84D34BA59FF88132ED093488CFA87" authority="Yang et al., 2004 a" authorityName="Yang et al." authorityYear="2004" box="[306,748,1269,1295]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="sebeiensis">
<emphasis id="13DCEAA5BA59FF88132ED093487AFA87" box="[306,538,1269,1295]" italics="true" pageId="28">Ilyocypris sebeiensis</emphasis>
<bibRefCitation id="45394B46BA59FF881002D09348BAFA87" author="Yang, F. &amp; Sun, Z. &amp; Qiao, Z. &amp; Zhang, Y." box="[542,730,1269,1295]" pageId="28" pagination="367 - 381" refId="ref24100" refString="Yang, F., Sun, Z., Qiao, Z. &amp; Zhang, Y. (2004) Revision of the diagnosis of the genus Qinghaicypris Huang, 1979 (Ostracoda) and the environmental significance of its type species. Acta Micropalaeontologica Sinica, 21, 367 - 381. [Chinese with English Summary]" type="journal article" year="2004">
Yang
<emphasis id="13DCEAA5BA59FF881042D09048EEFA87" box="[606,654,1269,1295]" italics="true" pageId="28">et al</emphasis>
., 2004
</bibRefCitation>
a
</taxonomicName>
lot less elongate compared to
<taxonomicName id="E6A84D34BA59FF881620D0934E8DFA87" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[1084,1261,1269,1295]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF881620D0934E8DFA87" box="[1084,1261,1269,1295]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF8816EFD0934F29FA87" box="[1267,1353,1269,1295]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="E6A84D34BA59FF881753D0934BA1FABB" authority="Huang, 1979" authorityName="Huang" authorityYear="1979" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="echinata">
<emphasis id="13DCEAA5BA59FF881753D0934B49FABB" italics="true" pageId="28">Ilyocypris echinata</emphasis>
<bibRefCitation id="45394B46BA59FF88132CD17F4BA1FABB" author="Huang B. R." box="[304,449,1305,1331]" pageId="28" pagination="187 - 236" refId="ref20250" refString="Huang B. R. (1979) Quaternary Ostracoda from Qinghai Lake region. Bulletin of Nanjing Institute of Geology and Palaeontology, Academia Sinica, 7, 187 - 236, with 11 plates." type="book chapter" year="1979">Huang, 1979</bibRefCitation>
</taxonomicName>
has lateral spines on the carapace, not seen in
<taxonomicName id="E6A84D34BA59FF8811DBD17F4E19FABB" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[967,1145,1305,1331]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF8811DBD17F4E19FABB" box="[967,1145,1305,1331]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF88169CD17F4EB6FABB" box="[1152,1238,1305,1331]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicName id="E6A84D34BA59FF8816C0D17F4BE1FADF" authority="Huang, 1982" authorityName="Huang" authorityYear="1982" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="microspinata">
<emphasis id="13DCEAA5BA59FF8816C0D17F4A89FADF" italics="true" pageId="28">Ilyocypris microspinata</emphasis>
Huang, 1982
</taxonomicName>
has a more sloping dorsal margin and a less inflated posterior in lateral view. The Pleistocene
<taxonomicName id="E6A84D34BA59FF88128BD1074B90FAF3" authority="Huang, 1982" authorityName="Huang" authorityYear="1982" box="[151,496,1377,1403]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="opima">
<emphasis id="13DCEAA5BA59FF88128BD1074B35FAF3" box="[151,341,1377,1403]" italics="true" pageId="28">Ilyocypris opima</emphasis>
Huang, 1982
</taxonomicName>
is much more rounded in lateral view and
<taxonomicName id="E6A84D34BA59FF8811C7D1074FFCFAF3" authority="Huang, 1982" authorityName="Huang" authorityYear="1982" box="[987,1436,1377,1403]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="crassireticulata">
<emphasis id="13DCEAA5BA59FF8811C7D1074F62FAF3" box="[987,1282,1377,1403]" italics="true" pageId="28">Ilyocypris crassireticulata</emphasis>
Huang, 1982
</taxonomicName>
has much coarser pitting.
</paragraph>
</subSubSection>
<subSubSection id="69B2653CBA59FF8812DBD1CE48D4F96B" pageId="28" type="materials_examined">
<paragraph id="211736B7BA59FF8812DBD1CE4A8EF9FF" blockId="28.[151,1437,153,1763]" pageId="28">
<emphasis id="13DCEAA5BA59FF8812DBD1CE4B01FA4B" bold="true" box="[199,353,1448,1475]" pageId="28">Distribution.</emphasis>
Currently known from
<collectingRegion id="E36CF855BA59FF88107AD1CF4924FA4B" box="[614,836,1449,1475]" country="South Korea" name="Gyeongbuk" pageId="28">Gyeongsangbuk-do</collectingRegion>
,
<collectingCountry id="59BF7627BA59FF881152D1CF49BCFA4B" box="[846,988,1449,1475]" name="South Korea" pageId="28">South Korea</collectingCountry>
, and
<collectingRegion id="E36CF855BA59FF881609D1CF4E9CFA4B" box="[1045,1276,1449,1475]" country="Japan" name="Okayama" pageId="28">Okayama Prefecture</collectingRegion>
,
<collectingCountry id="59BF7627BA59FF88171AD1CF4F26FA4B" box="[1286,1350,1449,1475]" name="Japan" pageId="28">Japan</collectingCountry>
(
<figureCitation id="B9932A32BA59FF881748D1CF4FF7FA4B" box="[1364,1431,1449,1475]" captionStart="FIGURE 9" captionStartId="13.[151,250,1588,1613]" captionTargetBox="[325,1263,199,1562]" captionTargetId="figure@13.[325,1263,199,1562]" captionTargetPageId="13" captionText="FIGURE 9. Distribution maps of Ilyocypris dentifera, I. angulata, I. japonica, I. mongolica, and Ilyocypris incus sp. nov. One additional fossil record of I. dentifera exists from Manitoba, Canada (Klassen et al. 1967), but is not shown on the map. As the records of Ilyocypris dentifera of Kim &amp; Min (1991) probably at least in part refer to different species, their records are not plotted on the map. Records of I. dentifera by Huang (1986) and Huang et al. (2006) are also not plotted as they are likely to also be misidentifications. Other reports with no figures cannot be confirmed but are plotted (see text for more details).? indicates uncertain records. Abbreviations and colours of points refer to Köppen climatic zones, following Peel et al. (2007). Data from: 1, Kaiser et al. (1973). 2, Shinde et al. (2014). 3, Kulkarni et al. (2015). 4, Zhai &amp; Zhao (2014). 5, Chen (1982; 1990). 6, Sars (1903), Chen (1982). 7, Yu et al. (2005). 8, Ma &amp; Yu (2018). 9, Yu (2014). 10, Yu et al. (2009). 11, Victor &amp; Fernando (1980). 12, Zenina &amp; Schornikov (2008). 13, Sidorov &amp; Semenchenko (2012). 14, Lee et al. (2000). 15, This study. 16, Tabuki &amp; Tsuhako (2015). 17, Okubo (1990). 18, Broodbakker (1988). 19, Okubo (2004), this study. 20, Tanaka et al. (2015). 21, Sánchez-Bayo &amp; Goka (2006a; 2006b). 22, Van der Meeren et al. (2010). 23, Martens (1991). 24, Zhai &amp; Zhao (2014). 25, This study. 26, Okubo &amp; Terauchi (1992). 27, This study. 28, Okubo (1990). 29, Nakao &amp; Tsukagoshi (2002; 2008). 30, This study. 31, Okubo (1974)." httpUri="https://zenodo.org/record/3363551/files/figure.png" pageId="28">Fig. 9</figureCitation>
, see
<tableCitation id="6C2A030CBA59FF8812DCD1AB4B72FA6F" box="[192,274,1485,1511]" captionStart="TABLE 1" captionStartId="1.[151,244,1160,1187]" captionTargetBox="[163,1412,1310,1957]" captionText="TABLE 1. Adult Ilyocypris specimens examined for this study. Initial discriminations of species were based on lateral views of carapaces, and confirmed by dissections of male specimens. Figures in brackets indicate the number of specimens with tubercles on the carapace. Note that tubercles vary from weakly to well-developed and were no- ticeable on whole specimens or valves in dorsal view using a stereo optical microscope." httpUri="http://table.plazi.org/id/75D7663FBA44FF95128BD0EE4E0FFA87" pageId="28" tableUuid="75D7663FBA44FF95128BD0EE4E0FFA87">Table 1</tableCitation>
for more details). Both known occurrences of this species fall into the temperate, without dry season, hot summer Köppen climatic zone (Cfa) (
<bibRefCitation id="45394B46BA59FF881022D1974892F983" author="Peel, M. C. &amp; Finlayson, B. L. &amp; McMahon, T. A." box="[574,754,1521,1547]" pageId="28" pagination="1633 - 1644" refId="ref22573" refString="Peel, M. C., Finlayson, B. L. &amp; McMahon, T. A. (2007) Updated world map of the Koppen-Geiger climate classification. Hydrology and Earth System Sciences Discussions, European Geosciences Union, 11, 1633 - 1644. https: // doi. org / 10.5194 / hess- 11 - 1633 - 2007" type="journal article" year="2007">
Peel
<emphasis id="13DCEAA5BA59FF88106AD19448C7F983" box="[630,679,1521,1547]" italics="true" pageId="28">et al</emphasis>
. 2007
</bibRefCitation>
).
<bibRefCitation id="45394B46BA59FF881118D1974986F983" author="Kim, W. &amp; Min, G. S." box="[772,998,1521,1547]" pageId="28" pagination="307 - 322" refId="ref21073" refString="Kim, W. &amp; Min, G. S. (1991 a) Redescription of four Recent freshwater ostracod (Crustacea: Ostracoda) species from Korea. Korean Journal of Zoology, 34, 307 - 322." type="journal article" year="1991">Kim &amp; Min (1991a)</bibRefCitation>
reported “
<taxonomicName id="E6A84D34BA59FF881641D1944EB9F983" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[1117,1241,1521,1547]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA59FF881641D1944EB9F983" box="[1117,1241,1521,1547]" italics="true" pageId="28">I. dentifera</emphasis>
</taxonomicName>
”, which possibly represents this species, from various places throughout
<collectingCountry id="59BF7627BA59FF881116D27349FCF9A7" box="[778,924,1557,1583]" name="South Korea" pageId="28">South Korea</collectingCountry>
. However, considering the close similarity of this species with the true
<taxonomicName id="E6A84D34BA59FF8813AFD25C4850F9DB" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[435,560,1593,1619]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA59FF8813AFD25C4850F9DB" box="[435,560,1593,1619]" italics="true" pageId="28">I. dentifera</emphasis>
</taxonomicName>
, it is unknown how many of these records potentially refer to
<taxonomicName id="E6A84D34BA59FF8816F6D25F4FFCF9DB" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[1258,1436,1593,1619]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF8816F6D25F4FFCF9DB" box="[1258,1436,1593,1619]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF88128BD23B4A8EF9FF" box="[151,238,1629,1655]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="211736B7BA59FF8812DBD2E748D4F96B" blockId="28.[151,1437,153,1763]" pageId="28">
<emphasis id="13DCEAA5BA59FF8812DBD2E74B4CF913" bold="true" box="[199,300,1665,1691]" pageId="28">Ecology.</emphasis>
<taxonomicName id="E6A84D34BA59FF88132ED2E74B84F913" authority="Smith &amp; Zhai &amp; Chang, 2019" authorityName="Smith &amp; Zhai &amp; Chang" authorityYear="2019" box="[306,484,1665,1691]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="incus" status="sp. nov.">
<emphasis id="13DCEAA5BA59FF88132ED2E74B84F913" box="[306,484,1665,1691]" italics="true" pageId="28">Ilyocypris incus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="08EF57DEBA59FF8813F6D2E74820F913" box="[490,576,1665,1691]" pageId="28" rank="species">sp. nov.</taxonomicNameLabel>
was found in a rice field with
<taxonomicName id="E6A84D34BA59FF88118ED2E44E6EF913" authorityName="G. O. Sars-Chen" authorityYear="1903" box="[914,1038,1665,1691]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="dentifera">
<emphasis id="13DCEAA5BA59FF88118ED2E44E6EF913" box="[914,1038,1665,1691]" italics="true" pageId="28">I. dentifera</emphasis>
</taxonomicName>
and
<taxonomicName id="E6A84D34BA59FF881658D2E44EDDF913" authorityName="Okubo" authorityYear="1990" box="[1092,1213,1666,1691]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="13DCEAA5BA59FF881658D2E44EDDF913" box="[1092,1213,1666,1691]" italics="true" pageId="28">I. japonica</emphasis>
</taxonomicName>
during early July in
<collectingCountry id="59BF7627BA59FF88128BD2C34ABFF937" box="[151,223,1701,1727]" name="South Korea" pageId="28">Korea</collectingCountry>
.
<bibRefCitation id="45394B46BA59FF8812F5D2C34B1FF937" author="Okubo, I." box="[233,383,1701,1727]" pageId="28" pagination="46 - 50" refId="ref22322" refString="Okubo, I. (1974) Freshwater Ostracoda from Japan, XI. Ilyocypris angulata Sars, 1902. The Research Bulletin of Okayama Shujitsu Junior College, 4, 46 - 50." type="journal article" year="1974">Okubo (1974</bibRefCitation>
, as
<taxonomicName id="E6A84D34BA59FF8813BBD2C04841F937" authorityName="Sars" authorityYear="1903" box="[423,545,1701,1727]" class="Ostracoda" family="Ilyocyprididae" genus="Ilyocypris" kingdom="Animalia" order="Podocopida" pageId="28" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="13DCEAA5BA59FF8813BBD2C04841F937" box="[423,545,1701,1727]" italics="true" pageId="28">I. angulata</emphasis>
</taxonomicName>
) reported that it is found from spring to summer, sometimes autumn, “creeping” in mud, although the habitat was not mentioned.
</paragraph>
</subSubSection>
</treatment>
</document>