361 lines
42 KiB
XML
361 lines
42 KiB
XML
<document id="FBA6633B23A11F2618BE117EC1A44DC2" ID-CLB-Dataset="8243" ID-DOI="10.11646/zootaxa.4923.1.1" ID-GBIF-Dataset="0a027439-c601-4533-bd03-f0e5d4442801" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496757" ID-ZooBank="41D057A7-F855-47C2-A598-8000CFD83A5B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612332986149" checkinUser="plazi" docAuthor="Gunter, Nicole L. & Weir, Thomas A." docDate="2021" docId="1A3236786230223C09D9FA1FFC286E8F" docLanguage="en" docName="zootaxa.4923.1.1.pdf" docOrigin="Zootaxa 4923 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lepanus cardwellensis Gunter & Weir 2021, new species" docType="treatment" docVersion="11" lastPageNumber="33" masterDocId="E60B4E00622E221C094EFFBDFFC26960" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species" masterLastPageNumber="66" masterPageNumber="1" pageNumber="31" updateTime="1698926692322" updateUser="ExternalLinkService">
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<mods:title id="42AEA07E90BE6305C945F146360D6CB1">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species</mods:title>
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<mods:namePart id="5256E22FAF8ADF94EC02E9271AF830BA">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="33F55B733C75A5F88A4C9403BBC0FEC9">Weir, Thomas A.</mods:namePart>
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<mods:nameIdentifier id="4BA090499A5A962C6D7DC62F4FD25F95" type="email">ngunter@cmnh.org</mods:nameIdentifier>
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<mods:title id="5AD5CA3000BDAFCE75F87E69F23F20F6">Zootaxa</mods:title>
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<mods:part id="A7D76AA8E808337F366906847DC67303">
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<mods:date id="AD6857FA0DEACE5410F9901BF3CA6C75">2021</mods:date>
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<mods:number id="16EA4B091CBD10B1E5DC78AB58FB0BB4">2021-02-03</mods:number>
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<mods:number id="DDD581D039B3631EFB051FA3F427218A">4923</mods:number>
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<treatment id="1A3236786230223C09D9FA1FFC286E8F" ID-DOI="http://doi.org/10.5281/zenodo.4496740" ID-GBIF-Taxon="177324106" ID-Zenodo-Dep="4496740" LSID="urn:lsid:plazi:treatment:1A3236786230223C09D9FA1FFC286E8F" httpUri="http://treatment.plazi.org/id/1A3236786230223C09D9FA1FFC286E8F" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31">
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<emphasis id="A0EF5B7C6230220209D9FA1FFCCE6CDB" bold="true" box="[151,780,1441,1468]" pageId="30" pageNumber="31">
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<taxonomicName id="559BFCED6230220209D9FA1FFDAD6CDB" ID-CoL="7WWVQ" authority="Gunter & Weir" authorityName="Gunter & Weir" authorityYear="2021" box="[151,623,1441,1468]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="cardwellensis" status="sp. nov.">
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<emphasis id="A0EF5B7C6230220209D9FA1FFE686CDC" bold="true" box="[151,426,1441,1468]" italics="true" pageId="30" pageNumber="31">Lepanus cardwellensis</emphasis>
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Gunter & Weir
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</taxonomicName>
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,
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<taxonomicNameLabel id="BBDCE607623022020B32FA1FFCCE6CDB" box="[636,780,1441,1468]" pageId="30" pageNumber="31" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="DA81D4E56230220209D9FA78FE246C80" box="[151,486,1477,1504]" pageId="30" pageNumber="31" type="description">
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<paragraph id="9224876E6230220209D9FA78FE246C80" blockId="30.[151,780,1441,1504]" box="[151,486,1477,1504]" pageId="30" pageNumber="31">
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(
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<figureCitation id="0AA09BEB6230220209D1FA78FEE26C80" box="[159,288,1477,1504]" captionStart="FIGURE 2" captionStartId="28.[151,250,1798,1822]" captionTargetBox="[255,1320,187,1768]" captionTargetId="figure-26@28.[255,1332,181,1774]" captionTargetPageId="28" captionText="FIGURE 2. Some diagnostic characters of the Lepanus ustulatus species group. A, Dorsal view of male protibia of Lepanus ustulatus; B, dorsal view of female protibia of Lepanus ustulatus; C, dorsal view of male protibia of Lepanus globulus; D, male metatibia of Lepanus ustulatus; E, female metatibia of Lepanus ustulatus; F, male metatibia of Lepanus lemannae; G, lateral view of male protibia of Lepanus cameroni; H, lateral view of male protibia of Lepanus globulus; I, lateral view of male protibia of Lepanus cardwellensis; J, head of Lepanus cameroni showing eye canthus not dividing the eye; K, posterolateral view of pronotum of Lepanus globulus showing corners distinct; L, head of Lepanus globulus showing eye canthus almost dividing the eye; M, posterolateral view of pronotum of Lepanus cardwellensis showing corners rounded." figureDoi="http://doi.org/10.5281/zenodo.4496761" httpUri="https://zenodo.org/record/4496761/files/figure.png" pageId="30" pageNumber="31">Figs. 2I, M</figureCitation>
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;
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<figureCitation id="0AA09BEB623022020862FA7BFE8D6C80" box="[300,335,1478,1504]" captionStart="FIGURE 3" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[185,1384,196,1902]" captionTargetId="figure-19@29.[183,1404,181,1919]" captionTargetPageId="29" captionText="FIGURE 3. Dorsal habitus of species within the Lepanus ustulatus species group. A, Lepanus cameroni; B, Lepanus cardwellensis; C, Lepanus globulus; D, Lepanus lemannae; E, Lepanus ustulatus. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4496763" httpUri="https://zenodo.org/record/4496763/files/figure.png" pageId="30" pageNumber="31">3B</figureCitation>
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;
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<figureCitation id="0AA09BEB623022020815FA7BFEBD6C80" box="[347,383,1478,1504]" captionStart="FIGURE 4" captionStartId="30.[151,250,1263,1287]" captionTargetBox="[157,1422,197,1228]" captionTargetId="figure-26@30.[151,1436,181,1239]" captionTargetPageId="30" captionText="FIGURE 4. Pygidial configuration of species within the Lepanus ustulatus species group. A, Lepanus cameroni; B, Lepanus cardwellensis; C, Lepanus globulus; D, Lepanus lemannae; E, Lepanus ustulatus. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496765" httpUri="https://zenodo.org/record/4496765/files/figure.png" pageId="30" pageNumber="31">4B</figureCitation>
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;
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<figureCitation id="0AA09BEB6230220208C5FA78FE6C6C80" box="[395,430,1477,1504]" captionStart="FIGURE 5" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[217,1358,189,1913]" captionTargetId="figure-19@31.[213,1373,181,1919]" captionTargetPageId="31" captionText="FIGURE 5. Left, dorsal and right view of aedeagi of species within the Lepanus ustulatus species group. A, Lepanus cameroni; B, Lepanus cardwellensis; C, Lepanus globulus; D, Lepanus lemannae; E, Lepanus ustulatus. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496767" httpUri="https://zenodo.org/record/4496767/files/figure.png" pageId="30" pageNumber="31">5B</figureCitation>
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;
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<figureCitation id="0AA09BEB6230220208F4FA7BFE1C6C80" box="[442,478,1478,1504]" captionStart="FIGURE 7" captionStartId="33.[151,250,1247,1271]" captionTargetBox="[154,1425,186,1216]" captionTargetId="figure-19@33.[151,1436,181,1223]" captionTargetPageId="33" captionText="FIGURE 7. A, Map of known records of Lepanus cardwellensis, Lepanus globulus, and Lepanus lemannae with the zones of insect endemism of the Wet Tropics sensu Yeates & Montieth (2008) numbered and indicated by dashed lines. Zone numbers are as follows: 1 Mount Finnigan; 2 Thornton Peak; 3 Windsor Tableland; 4 Carbine Tableland; 5 Hann Tableland; 6 Black Mountain; 7 Lamb Range; 8 Walsh/Hugh Nelson Range; 9 Atherton Tableland; 10 Bellenden Ker; 11 Malbon Thompson Range; 12 Walter Hill Range; 13 Kirrama/Cardwell Ranges; 14 Seaview Range; 15 Hinchinbrook Island; 16 Paluma/Bluewater Ranges; 17 Mount Elliot, B, predicted distribution of Lepanus globulus; C, predicted distribution of Lepanus lemannae." figureDoi="http://doi.org/10.5281/zenodo.4496771" httpUri="https://zenodo.org/record/4496771/files/figure.png" pageId="30" pageNumber="31">7A</figureCitation>
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)
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<emphasis id="A0EF5B7C6230220209D9F9B3FEE46F47" bold="true" box="[151,294,1549,1576]" pageId="30" pageNumber="31">Type series.</emphasis>
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<typeStatus id="4D2039CC62302202087FF9B3FE606F48" box="[305,418,1550,1576]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
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: ♂
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</emphasis>
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“
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QLD,
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N Ingham
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,
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<date id="E625A1AE623022020AA9F9B3FBB66F48" box="[999,1140,1550,1576]" pageId="30" pageNumber="31" value="1982-08-09">
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,
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<collectorName id="3F6EE2B8623022020DCCF9B3FAD56F47" box="[1154,1303,1549,1576]" pageId="30" pageNumber="31">S. & J. Peck</collectorName>
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SBP102” / “rainforest, carrion bait” / “
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<collectionCode id="F48A1FAB623022020881F98FFDD46F2C" box="[463,534,1586,1612]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="30" pageNumber="31">ANIC</collectionCode>
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database 25 058170” / “
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25, det
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<collectorName id="3F6EE2B8623022020D58F98FFB476F2B" box="[1046,1157,1585,1612]" pageId="30" pageNumber="31">T.A. Weir</collectorName>
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2011” / “
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Image”, [
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,
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], (in
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)
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.
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<emphasis id="A0EF5B7C623022020B24F9EBFC1C6F0F" bold="true" box="[618,990,1621,1650]" pageId="30" pageNumber="31">
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<typeStatus id="4D2039CC623022020B24F9EBFD206F10" box="[618,738,1622,1648]" pageId="30" pageNumber="31" type="paratype">Paratypes</typeStatus>
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(4x
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<specimenCount id="849D4CE7623022020A59F9EAFCEF6F12" box="[791,813,1623,1650]" pageId="30" pageNumber="31" type="female">♀</specimenCount>
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):
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:
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</emphasis>
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,
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N Ingham
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,
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|
||
<elevation id="19B6605D623022020C2CF9EBFA5E6F10" box="[1378,1436,1622,1648]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="30" pageNumber="31" unit="m" value="80.0">80 m</elevation>
|
||
</quantity>
|
||
[
|
||
<geoCoordinate id="F7AFE1A96230220209D1F9C7FEE56FF4" box="[159,295,1658,1684]" degrees="18" direction="south" minutes="32" orientation="latitude" pageId="30" pageNumber="31" precision="15" seconds="00" value="-18.533333">18°32’00”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F7AFE1A962302202087AF9C4FE096FF4" box="[308,459,1657,1684]" degrees="146" direction="east" minutes="11" orientation="longitude" pageId="30" pageNumber="31" precision="15" seconds="00" value="146.18333">146°11’00”E</geoCoordinate>
|
||
],
|
||
<date id="E625A1AE623022020891F9C4FD996FF3" box="[479,603,1657,1683]" pageId="30" pageNumber="31" value="1982-08-09">
|
||
<collectingDate id="F6615846623022020891F9C4FD996FF3" box="[479,603,1657,1683]" pageId="30" pageNumber="31" value="1982-08-09">9.viii.1982</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="3F6EE2B8623022020B29F9C7FD7E6FF3" box="[615,700,1657,1684]" pageId="30" pageNumber="31">S. Peck</collectorName>
|
||
&
|
||
<collectorName id="3F6EE2B8623022020BACF9C7FCF06FF3" box="[738,818,1657,1684]" pageId="30" pageNumber="31">J. Peck</collectorName>
|
||
(25-65534
|
||
<specimenCount id="849D4CE7623022020AF8F9C7FB9C6FF4" box="[950,1118,1657,1684]" pageId="30" pageNumber="31" type="female">2 specimens ♀</specimenCount>
|
||
in
|
||
<collectionCode id="F48A1FAB623022020DCCF9C7FB096FF4" box="[1154,1227,1658,1684]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="30" pageNumber="31">ANIC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="22F38D33623022020D91F9C4FED66FBC" ID-GBIF-Occurrence="3034608588" collectingDate="2012-03-15" collectingDateMax="2012-03-16" collectingDateMin="2012-03-15" collectionCode="ANIC" collectorName="G. B. Monteith" elevation="130" latitude="-18.528889" location="Cardwell Range" longLatPrecision="21" longitude="146.18611" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
|
||
<location id="9744D1B5623022020D91F9C4FA5A6FF4" LSID="urn:lsid:plazi:treatment:1A3236786230223C09D9FA1FFC286E8F:9744D1B5623022020D91F9C4FA5A6FF4" box="[1247,1432,1657,1684]" latitude="-18.528889" longLatPrecision="21" longitude="146.18611" name="Cardwell Range" pageId="30" pageNumber="31" stateProvince="Queensland">Cardwell Range</location>
|
||
,
|
||
<geoCoordinate id="F7AFE1A96230220209D9F923FEDC6FD8" box="[151,286,1694,1720]" degrees="18" direction="south" minutes="31" orientation="latitude" pageId="30" pageNumber="31" precision="15" seconds="44" value="-18.528889">18°31’44”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F7AFE1A9623022020861F920FE066FD7" box="[303,452,1693,1719]" degrees="146" direction="east" minutes="11" orientation="longitude" pageId="30" pageNumber="31" precision="15" seconds="10" value="146.18611">146°11’10”E</geoCoordinate>
|
||
,
|
||
<quantity id="55632A8B62302202089AF923FDE06FD8" box="[468,546,1694,1720]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="30" pageNumber="31" unit="m" value="130.0">
|
||
<elevation id="19B6605D62302202089AF923FDE06FD8" box="[468,546,1694,1720]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="30" pageNumber="31" unit="m" value="130.0">130 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="E625A1AE623022020B7DF920FD1B6FD8" box="[563,729,1693,1720]" pageId="30" pageNumber="31" value="2012-03-15" valueMax="2012-03-16" valueMin="2012-03-15">
|
||
<collectingDate id="F6615846623022020B7DF920FD1B6FD8" box="[563,729,1693,1720]" pageId="30" pageNumber="31" value="2012-03-15" valueMax="2012-03-16" valueMin="2012-03-15">15–16.iii.2012</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="3F6EE2B8623022020BA7F923FC536FD7" box="[745,913,1693,1720]" pageId="30" pageNumber="31">G.B. Monteith</collectorName>
|
||
(25-053916
|
||
<specimenCount id="849D4CE7623022020D65F923FB126FD8" box="[1067,1232,1693,1720]" pageId="30" pageNumber="31" type="female">1 specimen ♀</specimenCount>
|
||
in
|
||
<collectionCode id="F48A1FAB623022020DB2F923FA826FD8" box="[1276,1344,1694,1720]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="30" pageNumber="31">ANIC</collectionCode>
|
||
also as COL2408)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="22F38D3362302202086EF97CFC8C6E60" ID-GBIF-Occurrence="3034608757" collectingDate="1986-12-29" collectionCode="CMNC" collectorName="H. Howden & A. Howden" elevation="700" latitude="-18.166666" location="Kirrama Range" longLatPrecision="21" longitude="145.78334" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="paratype">
|
||
<location id="9744D1B562302202086EF97CFE0D6FBC" LSID="urn:lsid:plazi:treatment:1A3236786230223C09D9FA1FFC286E8F:9744D1B562302202086EF97CFE0D6FBC" box="[288,463,1729,1756]" latitude="-18.166666" longLatPrecision="21" longitude="145.78334" name="Kirrama Range" pageId="30" pageNumber="31" stateProvince="Queensland">Kirrama Range</location>
|
||
,
|
||
<locationDeviation id="83969E4C623022020895F97FFD276FBB" box="[475,741,1729,1756]" location="Kennedy" pageId="30" pageNumber="31">
|
||
<quantity id="55632A8B623022020895F97FFDE16FBB" box="[475,547,1729,1756]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="30" pageNumber="31" unit="km" value="18.0">18 km</quantity>
|
||
NW of Kennedy
|
||
</locationDeviation>
|
||
[
|
||
<geoCoordinate id="F7AFE1A9623022020BBAF97FFCBF6FBC" box="[756,893,1730,1756]" degrees="18" direction="south" minutes="10" orientation="latitude" pageId="30" pageNumber="31" precision="15" seconds="00" value="-18.166666">18°10’00”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F7AFE1A9623022020AC7F97CFBE06FBC" box="[905,1058,1729,1756]" degrees="145" direction="east" minutes="47" orientation="longitude" pageId="30" pageNumber="31" precision="15" seconds="00" value="145.78334">145°47’00”E</geoCoordinate>
|
||
],
|
||
<quantity id="55632A8B623022020D78F97FFBBD6FBC" box="[1078,1151,1730,1756]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="31" unit="m" value="700.0">
|
||
<elevation id="19B6605D623022020D78F97FFBBD6FBC" box="[1078,1151,1730,1756]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="30" pageNumber="31" unit="m" value="700.0">700 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="E625A1AE623022020DC2F97CFACC6FBB" box="[1164,1294,1729,1755]" pageId="30" pageNumber="31" value="1986-12-29">
|
||
<collectingDate id="F6615846623022020DC2F97CFACC6FBB" box="[1164,1294,1729,1755]" pageId="30" pageNumber="31" value="1986-12-29">29.xii.1986</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="3F6EE2B8623022020C54F97FFA5E6FBB" box="[1306,1436,1729,1756]" pageId="30" pageNumber="31">H. Howden</collectorName>
|
||
&
|
||
<collectorName id="3F6EE2B86230220209FCF95BFEF56F9F" box="[178,311,1765,1792]" pageId="30" pageNumber="31">A. Howden</collectorName>
|
||
, (CMNEN-00030067
|
||
<specimenCount id="849D4CE7623022020B7EF95BFD096E60" box="[560,715,1765,1792]" pageId="30" pageNumber="31" type="female">1 specimen ♀</specimenCount>
|
||
in
|
||
<collectionCode id="F48A1FAB623022020BA1F95BFC856E60" box="[751,839,1766,1792]" pageId="30" pageNumber="31">CMNC</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA81D4E56230223C0989F8B4FDBD6FAF" lastPageId="32" lastPageNumber="33" pageId="30" pageNumber="31" type="description">
|
||
<paragraph id="9224876E623022020989F8B4FE1A6E28" blockId="30.[151,1436,1549,2008]" pageId="30" pageNumber="31">
|
||
<emphasis id="A0EF5B7C623022020989F8B4FE9B6E43" bold="true" box="[199,345,1801,1827]" pageId="30" pageNumber="31">Description.</emphasis>
|
||
Uniform in colour, dark brown to black. Antennal clubs grey. Dorsal surfaces nitid with extremely fine setae (often rubbed off).
|
||
</paragraph>
|
||
<paragraph id="9224876E623022020989F8ECFB036E0C" blockId="30.[151,1436,1549,2008]" box="[199,1217,1873,1900]" pageId="30" pageNumber="31">
|
||
Total length: 4.0 mm.
|
||
<typeStatus id="4D2039CC62302202088EF8ECFDEA6E0B" box="[448,552,1873,1899]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
|
||
measurements: Total length 4.0 mm; elytral width
|
||
<quantity id="55632A8B623022020D2AF8EFFB7F6E0C" box="[1124,1213,1874,1900]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="30" pageNumber="31" unit="mm" value="2.9">2.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9224876E623022020989F8C8FEB56EB8" blockId="30.[151,1436,1549,2008]" pageId="30" pageNumber="31">
|
||
<emphasis id="A0EF5B7C623022020989F8C8FECA6EEF" box="[199,264,1909,1935]" italics="true" pageId="30" pageNumber="31">Male.</emphasis>
|
||
Head: Width to length ratio 35: 24. Surface smooth and nitid, with very fine punctation. Dorsal part of eyes wide, separated by an interocular space approximately 11 times eye width (54: 5); eye canthus dividing or nearly dividing eye.
|
||
</paragraph>
|
||
<caption id="C6E4D7E66231220309D9F82AFE4D6E97" ID-DOI="http://doi.org/10.5281/zenodo.4496767" ID-Zenodo-Dep="4496767" httpUri="https://zenodo.org/record/4496767/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1943,1967]" targetBox="[217,1358,189,1913]" targetPageId="31">
|
||
<paragraph id="9224876E6231220309D9F82AFE4D6E97" blockId="31.[151,1436,1943,2040]" pageId="31" pageNumber="32">
|
||
<emphasis id="A0EF5B7C6231220309D9F82AFED06ED0" bold="true" box="[151,274,1943,1968]" pageId="31" pageNumber="32">FIGURE 5.</emphasis>
|
||
Left, dorsal and right view of aedeagi of species within the
|
||
<taxonomicName id="559BFCED623122030A24F825FBE06ED0" baseAuthorityName="Lansberge" baseAuthorityYear="1874" box="[874,1058,1944,1968]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="ustulatus">
|
||
<emphasis id="A0EF5B7C623122030A24F825FBE06ED0" box="[874,1058,1944,1968]" italics="true" pageId="31" pageNumber="32">Lepanus ustulatus</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="A0EF5B7C623122030DF3F82AFB0D6ECF" bold="true" box="[1213,1231,1943,1967]" pageId="31" pageNumber="32">A</emphasis>
|
||
,
|
||
<taxonomicName id="559BFCED623122030D95F825FA576ECF" authorityName="Gunter & Weir" authorityYear="2021" box="[1243,1429,1944,1967]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="cameroni">
|
||
<emphasis id="A0EF5B7C623122030D95F825FA576ECF" box="[1243,1429,1944,1967]" italics="true" pageId="31" pageNumber="32">Lepanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="A0EF5B7C6231220309D9F801FF6A6EB4" bold="true" box="[151,168,1980,2004]" pageId="31" pageNumber="32">B</emphasis>
|
||
,
|
||
<taxonomicName id="559BFCED6231220309F9F801FE5C6EB4" authorityName="Gunter & Weir" authorityYear="2021" box="[183,414,1980,2004]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="cardwellensis">
|
||
<emphasis id="A0EF5B7C6231220309F9F801FE5C6EB4" box="[183,414,1980,2004]" italics="true" pageId="31" pageNumber="32">Lepanus cardwellensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="A0EF5B7C6231220308E0F806FE026EB3" bold="true" box="[430,448,1979,2003]" pageId="31" pageNumber="32">C</emphasis>
|
||
,
|
||
<taxonomicName id="559BFCED623122030881F801FD466EB4" authorityName="DET" authorityYear="1971" baseAuthorityName="Macleay" baseAuthorityYear="1887" box="[463,644,1980,2004]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="globulus">
|
||
<emphasis id="A0EF5B7C623122030881F801FD466EB4" box="[463,644,1980,2004]" italics="true" pageId="31" pageNumber="32">Lepanus globulus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="A0EF5B7C623122030BDDF801FD676EB4" bold="true" box="[659,677,1980,2004]" pageId="31" pageNumber="32">D</emphasis>
|
||
,
|
||
<taxonomicName id="559BFCED623122030BFAF801FCB66EB4" authorityName="Gunter & Weir" authorityYear="2021" box="[692,884,1980,2004]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="lemannae">
|
||
<emphasis id="A0EF5B7C623122030BFAF801FCB66EB4" box="[692,884,1980,2004]" italics="true" pageId="31" pageNumber="32">Lepanus lemannae</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="A0EF5B7C623122030ACDF801FC566EB4" bold="true" box="[899,916,1980,2004]" pageId="31" pageNumber="32">E</emphasis>
|
||
,
|
||
<taxonomicName id="559BFCED623122030AEDF801FB9E6EB4" baseAuthorityName="Lansberge" baseAuthorityYear="1874" box="[931,1116,1980,2004]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="ustulatus">
|
||
<emphasis id="A0EF5B7C623122030AEDF801FB9E6EB4" box="[931,1116,1980,2004]" italics="true" pageId="31" pageNumber="32">Lepanus ustulatus</emphasis>
|
||
</taxonomicName>
|
||
. All specimens in comparative scale. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C6E4D7E6620E223C09D9FC5AFE066D42" ID-DOI="http://doi.org/10.5281/zenodo.4496769" ID-Zenodo-Dep="4496769" httpUri="https://zenodo.org/record/4496769/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,999,1023]" targetBox="[154,1436,187,969]" targetPageId="32">
|
||
<paragraph id="9224876E620E223C09D9FC5AFE066D42" blockId="32.[151,1436,999,1059]" pageId="32" pageNumber="33">
|
||
<emphasis id="A0EF5B7C620E223C09D9FC5AFEE96A9F" bold="true" box="[151,299,999,1023]" pageId="32" pageNumber="33">FIGURE 6. A</emphasis>
|
||
, Map of known records of
|
||
<taxonomicName id="559BFCED620E223C0B0DFC55FD396A9F" baseAuthorityName="Lansberge" baseAuthorityYear="1874" box="[579,763,999,1023]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ustulatus">
|
||
<emphasis id="A0EF5B7C620E223C0B0DFC55FD396A9F" box="[579,763,999,1023]" italics="true" pageId="32" pageNumber="33">Lepanus ustulatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="A0EF5B7C620E223C0A47FC5AFCD86A9F" bold="true" box="[777,794,999,1023]" pageId="32" pageNumber="33">B</emphasis>
|
||
, predicted distribution of
|
||
<taxonomicName id="559BFCED620E223C0D6DFC55FB1E6A9F" baseAuthorityName="Lansberge" baseAuthorityYear="1874" box="[1059,1244,999,1023]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ustulatus">
|
||
<emphasis id="A0EF5B7C620E223C0D6DFC55FB1E6A9F" box="[1059,1244,999,1023]" italics="true" pageId="32" pageNumber="33">Lepanus ustulatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="A0EF5B7C620E223C0DA7FC5AFB396A9F" bold="true" box="[1257,1275,999,1023]" pageId="32" pageNumber="33">C</emphasis>
|
||
, map of known records of
|
||
<taxonomicName id="559BFCED620E223C084AFBB1FE7D6D43" authorityName="Gunter & Weir" authorityYear="2021" box="[260,447,1036,1059]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cameroni">
|
||
<emphasis id="A0EF5B7C620E223C084AFBB1FE7D6D43" box="[260,447,1036,1059]" italics="true" pageId="32" pageNumber="33">Lepanus cameroni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9224876E620E223C0989FBECFDDD6DD4" blockId="32.[151,1437,1105,2031]" pageId="32" pageNumber="33">Prothorax: Pronotum with posterolateral angles rounded and obtuse, lateral margins bordered. Pronotal surface smooth, nitid, finely and densely punctate. Hypomeral surface very finely reticulate, hypomeral stria absent. Pronotum width to length ratio 57: 35.</paragraph>
|
||
<paragraph id="9224876E620E223C0989FB00FD636C7F" blockId="32.[151,1437,1105,2031]" pageId="32" pageNumber="33">Elytra: Surface smooth, nitid. Striae superficial, impunctate. Stria 6 not extending to elytral base and stria 7 extending to the elytral base. 8th stria impunctate where it curves inwards posteriorly. Ratio of length of the elytra along suture to maximum elytral width 65: 72.</paragraph>
|
||
<paragraph id="9224876E620E223C0989FA97FED66CCF" blockId="32.[151,1437,1105,2031]" pageId="32" pageNumber="33">Legs: Ventral ridge of protibiae without crenulations but with small flange. Protibia with 2 teeth on the outer edge, front edge truncate, apical digit narrow and sharp, apical spur absent. Mesotibia with a brush of setae apically on inner side. Metatibia with inner edge nearly straight and not crenulate. Basal metatarsomere with slight lobe on inner edge.</paragraph>
|
||
<paragraph id="9224876E620E223C0989FA04FF3C6F7C" blockId="32.[151,1437,1105,2031]" pageId="32" pageNumber="33">Abdomen: Pygidium with a transverse sinuate fold across base that is curved and not impressed medially. Pygidial disc with a large flat, shiny, finely reticulate central area that occupies about half the disc. Ventrite 6 impunctate.</paragraph>
|
||
<paragraph id="9224876E620E223C0989F998FDB96FE7" blockId="32.[151,1437,1105,2031]" pageId="32" pageNumber="33">Pterothorax: Mesometaventral suture slightly arched. Medial lobe of metaventrite smooth, nitid, virtually impunctate, broadly margined beside mesocoxae. Mesoventrite smooth, nitid, and notched anteriorly. Mesepimeron smooth. Metanepisternum finely reticulate.</paragraph>
|
||
<paragraph id="9224876E620E223C0989F92CFDBD6FAF" blockId="32.[151,1437,1105,2031]" pageId="32" pageNumber="33">
|
||
<emphasis id="A0EF5B7C620E223C0989F92CFEE16FCB" box="[199,291,1681,1707]" italics="true" pageId="32" pageNumber="33">Female.</emphasis>
|
||
Protibia with front edge truncate and apical spur arising directly from front edge. Apical digit absent. Mesotibia without a brush of setae apically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA81D4E5620E223C0989F964FA406F93" box="[199,1410,1753,1780]" pageId="32" pageNumber="33" type="etymology">
|
||
<paragraph id="9224876E620E223C0989F964FA406F93" blockId="32.[151,1437,1105,2031]" box="[199,1410,1753,1780]" pageId="32" pageNumber="33">
|
||
<emphasis id="A0EF5B7C620E223C0989F964FE8C6F93" bold="true" box="[199,334,1753,1779]" pageId="32" pageNumber="33">Etymology.</emphasis>
|
||
Named after the Cardwell Range of the Australian Wet Tropics, where most records are known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA81D4E5620E223C0989F940FAE46E5B" pageId="32" pageNumber="33" type="distribution">
|
||
<paragraph id="9224876E620E223C0989F940FAE46E5B" blockId="32.[151,1437,1105,2031]" pageId="32" pageNumber="33">
|
||
<emphasis id="A0EF5B7C620E223C0989F940FEA36E77" bold="true" box="[199,353,1789,1815]" pageId="32" pageNumber="33">Distribution.</emphasis>
|
||
Known from rainforests below
|
||
<quantity id="55632A8B620E223C0B8DF940FCCB6E78" box="[707,777,1789,1816]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="32" pageNumber="33" unit="m" value="700.0">700 m</quantity>
|
||
in the contiguous Cardwell and Kirrama Ranges in the Wet Tropics (WET) IBRA bioregion [Zone 13] (
|
||
<figureCitation id="0AA09BEB620E223C0BCCF89CFD1E6E5B" box="[642,732,1825,1851]" captionStart="FIGURE 7" captionStartId="33.[151,250,1247,1271]" captionTargetBox="[154,1425,186,1216]" captionTargetId="figure-19@33.[151,1436,181,1223]" captionTargetPageId="33" captionText="FIGURE 7. A, Map of known records of Lepanus cardwellensis, Lepanus globulus, and Lepanus lemannae with the zones of insect endemism of the Wet Tropics sensu Yeates & Montieth (2008) numbered and indicated by dashed lines. Zone numbers are as follows: 1 Mount Finnigan; 2 Thornton Peak; 3 Windsor Tableland; 4 Carbine Tableland; 5 Hann Tableland; 6 Black Mountain; 7 Lamb Range; 8 Walsh/Hugh Nelson Range; 9 Atherton Tableland; 10 Bellenden Ker; 11 Malbon Thompson Range; 12 Walter Hill Range; 13 Kirrama/Cardwell Ranges; 14 Seaview Range; 15 Hinchinbrook Island; 16 Paluma/Bluewater Ranges; 17 Mount Elliot, B, predicted distribution of Lepanus globulus; C, predicted distribution of Lepanus lemannae." figureDoi="http://doi.org/10.5281/zenodo.4496771" httpUri="https://zenodo.org/record/4496771/files/figure.png" pageId="32" pageNumber="33">Fig. 7A</figureCitation>
|
||
). This species may represent a short-range endemic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA81D4E5620E223C0989F8F8FC286E8F" pageId="32" pageNumber="33" type="discussion">
|
||
<paragraph id="9224876E620E223C0989F8F8FC286E8F" blockId="32.[151,1437,1105,2031]" pageId="32" pageNumber="33">
|
||
<emphasis id="A0EF5B7C620E223C0989F8F8FE8D6E3F" bold="true" box="[199,335,1861,1887]" pageId="32" pageNumber="33">Comments.</emphasis>
|
||
Only known from a total of five specimens collected from either dung or carrion traps. This species has been referred to under the informal name of “
|
||
<taxonomicName id="559BFCED620E223C0BBDF8D7FC446EE4" authority="NQ" authorityName="NQ" box="[755,902,1897,1924]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0EF5B7C620E223C0BBDF8D7FC906EE3" box="[755,850,1898,1923]" italics="true" pageId="32" pageNumber="33">Lepanus</emphasis>
|
||
NQ
|
||
</taxonomicName>
|
||
25”. The
|
||
<taxonomicName id="559BFCED620E223C0AA7F8D7FBBB6EE4" authority="NQ" authorityName="NQ" box="[1001,1145,1897,1924]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A0EF5B7C620E223C0AA7F8D7FB8A6EE3" box="[1001,1096,1898,1923]" italics="true" pageId="32" pageNumber="33">Lepanus</emphasis>
|
||
NQ
|
||
</taxonomicName>
|
||
<specimenCount id="849D4CE7620E223C0D37F8D4FAC66EE3" box="[1145,1284,1897,1924]" pageId="32" pageNumber="33" type="generic">25 specimen</specimenCount>
|
||
sequenced in
|
||
<bibRefCitation id="F60AFA9F620E223C09D9F830FEB66EC7" author="Gunter, N. L. & Weir, T. A." box="[151,372,1933,1959]" pageId="32" pageNumber="33" pagination="41 - 80" refId="ref76546" refString="Gunter, N. L. & Weir, T. A. (2019 a) Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group. Zootaxa, 4564 (1), 41 - 80. https: // doi. org / 10.11646 / zootaxa. 4564.1.2" type="journal article" year="2019">
|
||
Gunter
|
||
<emphasis id="A0EF5B7C620E223C09A5F833FEE16EC7" box="[235,291,1933,1959]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
was collected from Cardwell Range, is designated as a
|
||
<typeStatus id="4D2039CC620E223C0AAEF833FB826EC8" box="[992,1088,1934,1960]" pageId="32" pageNumber="33" type="paratype">paratype</typeStatus>
|
||
here (ANIC 25-053831 also as COL2408) and was recovered within the clade that contains all species in the
|
||
<taxonomicName id="559BFCED620E223C0D4BF80FFB126EAB" baseAuthorityName="Lansberge" baseAuthorityYear="1874" box="[1029,1232,1969,1995]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="ustulatus">
|
||
<emphasis id="A0EF5B7C620E223C0D4BF80FFB126EAB" box="[1029,1232,1969,1995]" italics="true" pageId="32" pageNumber="33">Lepanus ustulatus</emphasis>
|
||
</taxonomicName>
|
||
species group. Its relationship with
|
||
<taxonomicName id="559BFCED620E223C0815F86BFE1B6E8F" authorityName="DET" authorityYear="1971" baseAuthorityName="Macleay" baseAuthorityYear="1887" box="[347,473,2005,2031]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="globulus">
|
||
<emphasis id="A0EF5B7C620E223C0815F86BFE1B6E8F" box="[347,473,2005,2031]" italics="true" pageId="32" pageNumber="33">L. globulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="559BFCED620E223C0B5EF86BFD546E8F" authorityName="Gunter & Weir" authorityYear="2021" box="[528,662,2006,2031]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="cameroni">
|
||
<emphasis id="A0EF5B7C620E223C0B5EF86BFD546E8F" box="[528,662,2006,2031]" italics="true" pageId="32" pageNumber="33">L. cameroni</emphasis>
|
||
</taxonomicName>
|
||
is only moderately supported.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |