372 lines
44 KiB
XML
372 lines
44 KiB
XML
<document id="9989BD7A04A5586F9A582703FD5ED49A" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="99789763659385B1D36C2384FC4F9D5D" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiocamax dorotheae Thuy 2013, sp. nov." docType="treatment" docUuid="412FC94A-B8E6-48B2-915D-E719B0AD10CC" docUuidSource="ZooBank" docVersion="10" lastPageNumber="203" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="199" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F11902E79B81F33C44F10D7DAC18E3D5">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:namePart id="30E8D072C0A9AC36E9CF63185F647066">Thuy, Ben</mods:namePart>
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<mods:title id="CA870DA9C994B01E3446AF99E38981AA">European Journal of Taxonomy</mods:title>
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<mods:part id="6C88854C23FBE030FC9511C6F8D17A6F">
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<mods:date id="41BC647BF2D8E6790552D2DEAD4213D9">2013</mods:date>
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<mods:number id="84AE7DD4AFC245FE11FF5B11E883941B">2013-07-02</mods:number>
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<mods:number id="EA11685855199DFEE8A56F846666F38E">48</mods:number>
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<treatment id="99789763659385B1D36C2384FC4F9D5D" ID-DOI="http://doi.org/10.5281/zenodo.3844141" ID-GBIF-Taxon="164044797" ID-Zenodo-Dep="3844141" LSID="urn:lsid:zoobank.org:act:412FC94A-B8E6-48B2-915D-E719B0AD10CC" httpUri="http://treatment.plazi.org/id/99789763659385B1D36C2384FC4F9D5D" lastPageId="202" lastPageNumber="203" pageId="198" pageNumber="199">
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<subSubSection id="59CB75FE659385BDD36C2384FC2099B0" box="[604,984,1709,1736]" pageId="198" pageNumber="199" type="nomenclature">
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<paragraph id="116E2675659385BDD36C2384FC2099B0" blockId="198.[361,1227,1709,1813]" box="[604,984,1709,1736]" pageId="198" pageNumber="199">
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<heading id="4A269119659385BDD36C2384FC2099B0" box="[604,984,1709,1736]" centered="true" fontSize="11" level="2" pageId="198" pageNumber="199" reason="2">
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<taxonomicName id="D6D15DF6659385BDD36C2384FC8F99B0" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" box="[604,887,1709,1736]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="198" pageNumber="199" phylum="Echinodermata" rank="species" species="dorotheae" status="sp. nov.">
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<emphasis id="23A5FA67659385BDD36C2384FC8F99B0" bold="true" box="[604,887,1709,1736]" italics="true" pageId="198" pageNumber="199">Ophiocamax dorotheae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3896471C659385BDD24E2387FC2099B0" box="[894,984,1710,1736]" pageId="198" pageNumber="199" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE659385BDD05923FAFC9D986C" pageId="198" pageNumber="199" type="description">
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<paragraph id="116E2675659385BDD05923FAFB339996" blockId="198.[361,1227,1709,1813]" box="[361,1227,1747,1774]" pageId="198" pageNumber="199">
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<uri id="65402A77659385BDD05923FAFB339996" box="[361,1227,1747,1774]" pageId="198" pageNumber="199">
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urn:lsid:zoobank.org:act:
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<uuid id="65771CA0659385BDD3B723FAFB339996" box="[647,1227,1747,1774]" pageId="198" pageNumber="199">412FC94A-B8E6-48B2-915D-E719B0AD10CC</uuid>
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</uri>
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</paragraph>
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<paragraph id="116E2675659385BDD3FE23D3FC9D986C" blockId="198.[361,1227,1709,1813]" box="[718,869,1786,1813]" pageId="198" pageNumber="199">
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<figureCitation id="89EA3AF0659385BDD3FE23D3FCDD986D" box="[718,805,1786,1813]" captionStart="Fig" captionStartId="200.[189,231,1754,1780]" captionTargetBox="[258,1330,265,1715]" captionTargetId="figure@200.[258,1330,265,1715]" captionTargetPageId="200" captionText="Fig. 35. Skeletal plates and arm fragments of fossil and Recent ophiacanthid brittle stars; lateral arm plates (LAPs) in external (a) and internal (b) views. 1-3. Ophiocamax vitrea Lyman, 1878, Recent. 1. Proximal LAP. 2. Median LAP. 3. Dorsal arm plate. 4-5. Ophiocamax hystrix Lyman, 1878, Recent. 4. Proximal LAP. 5. Median LAP. 6. Ophiocamax austera Verrill, 1899, Recent; proximal LAP. 7-11. Ophiocamax dorotheae sp. nov. from the late Oxfordian (Late Jurassic) of Savigna, France. 7. GZG.INV.78763 (holotype), proximal LAP. 8. GZG.INV.78764 (paratype), median LAP. 9. GZG.INV.78765 (paratype), distal LAP. 10. GZG.INV.78766 (paratype), proximal arm fragment in ventral view. 11. GZG.INV.78767 (paratype), median arm fragment in dorsal view. One common scale bar per species except for 10 and 11." figureDoi="http://doi.org/10.5281/zenodo.3822911" httpUri="https://zenodo.org/record/3822911/files/figure.png" pageId="198" pageNumber="199">Fig. 35</figureCitation>
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: 7-11
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</paragraph>
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</subSubSection>
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<subSubSection id="59CB75FE659385BDD18D2216FD4C9893" pageId="198" pageNumber="199" type="reference_group">
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<paragraph id="116E2675659385BDD18D2216FCB39821" blockId="198.[189,843,1855,1882]" box="[189,843,1855,1882]" pageId="198" pageNumber="199">
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<treatmentCitationGroup id="31C1015B659385BDD18D2216FCB39821" box="[189,843,1855,1882]" pageId="198" pageNumber="199">
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<taxonomicName id="D6D15DF6659385BDD18D2216FCBE9821" authorityName="Muller & Troschel" authorityYear="1842" box="[189,838,1855,1882]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="198" pageNumber="199" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67659385BDD18D2216FEAE9821" box="[189,342,1855,1881]" italics="true" pageId="198" pageNumber="199">Ophiacantha</emphasis>
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?
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<emphasis id="23A5FA67659385BDD05A2269FDE09821" box="[362,536,1856,1881]" italics="true" pageId="198" pageNumber="199">suprajurassica</emphasis>
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<treatmentCitation id="90700064659385BDD3102216FD049822" author="Hess H." box="[544,764,1855,1882]" page="1065" pageId="198" pageNumber="199" year="1965">
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<bibRefCitation id="75405B84659385BDD3102216FD049822" author="Hess H." box="[544,764,1855,1882]" pageId="198" pageNumber="199" pagination="1059 - 1082" refId="ref122348" refString="Hess H. 1965 a. Mikropalaontologische Untersuchungen an Ophiuren IV: Die Ophiuren aus dem Renggeri-Ton (Unter-Oxford) von Chapois (Jura) und Longecombe (Ain). Eclogae geologicae Helvetiae 58: 1059 - 1082." type="journal article" year="1965">Hess, 1965a: 1065</bibRefCitation>
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</treatmentCitation>
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, 1077
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="116E2675659385BDD18D22ADFC6698E6" blockId="198.[189,1399,1924,2027]" box="[189,926,1924,1950]" pageId="198" pageNumber="199">
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<treatmentCitationGroup id="31C1015B659385BDD18D22ADFC6698E6" box="[189,926,1924,1950]" pageId="198" pageNumber="199">
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<taxonomicName id="D6D15DF6659385BDD18D22ADFDCC98E6" authorityName="Muller & Troschel" authorityYear="1842" box="[189,564,1924,1950]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="198" pageNumber="199" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67659385BDD18D22ADFEAE98E6" box="[189,342,1924,1950]" italics="true" pageId="198" pageNumber="199">Ophiacantha</emphasis>
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? sp. or
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<emphasis id="23A5FA67659385BDD08322ADFDCC98E6" box="[435,564,1924,1950]" italics="true" pageId="198" pageNumber="199">Ophiothrix</emphasis>
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</taxonomicName>
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? sp. –
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<treatmentCitation id="90700064659385BDD3B722ADFCB898E6" author="Hess H." box="[647,832,1924,1950]" page="419" pageId="198" pageNumber="199" year="1960">
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||
<bibRefCitation id="75405B84659385BDD3B722ADFCB898E6" author="Hess H." box="[647,832,1924,1950]" pageId="198" pageNumber="199" pagination="385 - 421" refId="ref122234" refString="Hess H. 1960. Ophiurenreste aus dem Malm des Schweizer Juras und des Departements Haut-Rhin. Eclogae geologicae Helvetiae 53: 385 - 421." type="journal article" year="1960">Hess 1960: 419</bibRefCitation>
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</treatmentCitation>
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, fig. 39.
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</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="116E2675659385BDD18D2283FD4C9893" blockId="198.[189,1399,1924,2027]" pageId="198" pageNumber="199">
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||
<treatmentCitationGroup id="31C1015B659385BDD18D2283FD4C9893" pageId="198" pageNumber="199">
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||
<taxonomicName id="D6D15DF6659385BDD18D2283FDE498BC" authorityName="Muller & Troschel" authorityYear="1842" box="[189,540,1962,1988]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="198" pageNumber="199" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67659385BDD18D2283FEAE98BC" box="[189,342,1962,1988]" italics="true" pageId="198" pageNumber="199">Ophiacantha</emphasis>
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||
?
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<emphasis id="23A5FA67659385BDD05E2282FDE498BC" box="[366,540,1963,1988]" italics="true" pageId="198" pageNumber="199">suprajurassica</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="90700064659385BDD3712282FCE898BC" author="Hess H." box="[577,784,1962,1989]" page="1030" pageId="198" pageNumber="199" year="1966">
|
||
<bibRefCitation id="75405B84659385BDD3712282FCE898BC" author="Hess H." box="[577,784,1962,1989]" pageId="198" pageNumber="199" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess 1966: 1030</bibRefCitation>
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||
</treatmentCitation>
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||
, 1054, figs 76-77 (material incorrectly assigned to
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<taxonomicName id="D6D15DF6659385BDD18D22F8FD5F9893" authorityName="Muller & Troschel" authorityYear="1842" box="[189,679,2000,2027]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="198" pageNumber="199" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67659385BDD18D22F8FEAE9893" box="[189,342,2001,2027]" italics="true" pageId="198" pageNumber="199">Ophiacantha</emphasis>
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?
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<emphasis id="23A5FA67659385BDD05A22FBFDE09893" box="[362,536,2002,2027]" italics="true" pageId="198" pageNumber="199">suprajurassica</emphasis>
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Hess, 1965
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</taxonomicName>
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||
).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59CB75FE659285BCD18D2422FB559E9D" pageId="199" pageNumber="200" type="diagnosis">
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||
<paragraph id="116E2675659285BCD18D2422FECE9E5D" blockId="199.[189,310,267,293]" box="[189,310,267,293]" pageId="199" pageNumber="200">
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||
<heading id="4A269119659285BCD18D2422FECE9E5D" bold="true" box="[189,310,267,293]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
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||
<emphasis id="23A5FA67659285BCD18D2422FECE9E5D" bold="true" box="[189,310,267,293]" pageId="199" pageNumber="200">Diagnosis</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="116E2675659285BCD18D2413FB559E9D" blockId="199.[189,1399,314,486]" pageId="199" pageNumber="200">
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Species of
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<taxonomicName id="D6D15DF6659285BCD0742413FE229E2C" authorityName="Lyman" authorityYear="1878" box="[324,474,314,340]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="199" pageNumber="200" phylum="Echinodermata" rank="genus">
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<emphasis id="23A5FA67659285BCD0742413FE229E2C" box="[324,474,314,340]" italics="true" pageId="199" pageNumber="200">Ophiocamax</emphasis>
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</taxonomicName>
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||
with large LAPs displaying large, well-defined, strongly protruding spur on the outer proximal edge paralleled by a large, non-prominent, rounded area of densely meshed stereom on the inner distal edge; up to five spine articulations, two dorsalmost of which always largest; ridge on inner side of LAP relatively short, with strongly widened dorsal tip; dorsal arm plates irregularly pentagonal to blazon-shaped, with small granules or thorns scattered on outer surface.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59CB75FE659285BCD18D2727FDD99D03" pageId="199" pageNumber="200" type="etymology">
|
||
<paragraph id="116E2675659285BCD18D2727FEBC9D50" blockId="199.[189,324,526,552]" box="[189,324,526,552]" pageId="199" pageNumber="200">
|
||
<heading id="4A269119659285BCD18D2727FEBC9D50" bold="true" box="[189,324,526,552]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
|
||
<emphasis id="23A5FA67659285BCD18D2727FEBC9D50" bold="true" box="[189,324,526,552]" pageId="199" pageNumber="200">Etymology</emphasis>
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||
</heading>
|
||
</paragraph>
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<paragraph id="116E2675659285BCD18D2714FDD99D03" blockId="199.[189,1399,573,635]" pageId="199" pageNumber="200">Species named in honour of Dorothea Hause-Reitner for her friendship and generous assistance in scanning electron microscopy.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="59CB75FE659285BCD18D278DFBD69C00" pageId="199" pageNumber="200" type="materials_examined">
|
||
<paragraph id="116E2675659285BCD18D278DFE949DC6" blockId="199.[189,364,676,702]" box="[189,364,676,702]" pageId="199" pageNumber="200">
|
||
<heading id="4A269119659285BCD18D278DFE949DC6" bold="true" box="[189,364,676,702]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
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||
<emphasis id="23A5FA67659285BCD18D278DFE949DC6" bold="true" box="[189,364,676,702]" pageId="199" pageNumber="200">Type material</emphasis>
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||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E2675659285BCD1DA27FDFE719C69" blockId="199.[189,393,724,785]" pageId="199" pageNumber="200">
|
||
<materialsCitation id="A1B92C28659285BCD1DA27FDFE719C69" ID-GBIF-Occurrence="2612041331" collectionCode="GZG" pageId="199" pageNumber="200" specimenCode="GZG.INV.78763" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="23A5FA67659285BCD1DA27FDFEA49D96" bold="true" box="[234,348,724,750]" pageId="199" pageNumber="200">
|
||
<typeStatus id="CE6A98D7659285BCD1DA27FDFEA49D96" box="[234,348,724,750]" pageId="199" pageNumber="200" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<specimenCode id="41778E0E659285BCD18D27DEFE7E9C69" box="[189,390,759,785]" pageId="199" pageNumber="200">GZG.INV.78763</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="116E2675659285BCD1DA2613FE9F9C2C" blockId="199.[189,1070,826,889]" box="[234,359,826,852]" pageId="199" pageNumber="200">
|
||
<emphasis id="23A5FA67659285BCD1DA2613FE9F9C2C" bold="true" box="[234,359,826,852]" pageId="199" pageNumber="200">
|
||
<typeStatus id="CE6A98D7659285BCD1DA2613FE9F9C2C" box="[234,359,826,852]" pageId="199" pageNumber="200" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="116E2675659285BCD18D2677FBD69C00" blockId="199.[189,1070,826,889]" box="[189,1070,862,889]" pageId="199" pageNumber="200">GZG.INV.78764, GZG.INV.78765, GZG.INV.78766 and GZG.INV.78767.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE659285BCD18D2688FAD99C92" pageId="199" pageNumber="200" type="distribution">
|
||
<paragraph id="116E2675659285BCD18D2688FE039CC2" blockId="199.[189,507,928,955]" box="[189,507,928,955]" pageId="199" pageNumber="200">
|
||
<heading id="4A269119659285BCD18D2688FE039CC2" bold="true" box="[189,507,928,955]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
|
||
<emphasis id="23A5FA67659285BCD18D2688FE039CC2" bold="true" box="[189,507,928,955]" pageId="199" pageNumber="200">Type locality and horizon</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E2675659285BCD18D26F9FAD99C92" blockId="199.[189,1313,976,1003]" box="[189,1313,976,1003]" pageId="199" pageNumber="200">
|
||
Savigna,
|
||
<collectingCountry id="69C666E5659285BCD01A26F8FE869C93" box="[298,382,977,1003]" name="France" pageId="199" pageNumber="200">France</collectingCountry>
|
||
; sample S2b of
|
||
<bibRefCitation id="75405B84659285BCD30926F9FD3F9C92" author="Gale A. S." box="[569,711,976,1002]" pageId="199" pageNumber="200" pagination="69 - 89" refId="ref121688" refString="Gale A. S. 2011. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Departement du Jura, France. Swiss Journal of Palaeontology 130 (1): 69 - 89. http: // dx. doi. org / 10.1007 / s 13358 - 010 - 0008 - x" type="journal article" year="2011">Gale (2011)</bibRefCitation>
|
||
,
|
||
<emphasis id="23A5FA67659285BCD3E326F8FC8E9C92" box="[723,886,977,1002]" italics="true" pageId="199" pageNumber="200">Bimammatum</emphasis>
|
||
Zone, late Oxfordian, Late Jurassic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE659285BCD18D213BFE1F9A4E" pageId="199" pageNumber="200" type="materials_examined">
|
||
<paragraph id="116E2675659285BCD18D213BFE4C9B54" blockId="199.[189,436,1042,1068]" box="[189,436,1042,1068]" pageId="199" pageNumber="200">
|
||
<heading id="4A269119659285BCD18D213BFE4C9B54" bold="true" box="[189,436,1042,1068]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
|
||
<emphasis id="23A5FA67659285BCD18D213BFE4C9B54" bold="true" box="[189,436,1042,1068]" pageId="199" pageNumber="200">Additional material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E2675659285BCD18D216BFE1F9A4E" blockId="199.[189,1399,1090,1334]" pageId="199" pageNumber="200">
|
||
GZG.INV.78768 (29 dissociated LAPs) from sample S2b of
|
||
<bibRefCitation id="75405B84659285BCD245216BFBFA9B24" author="Gale A. S." box="[885,1026,1090,1116]" pageId="199" pageNumber="200" pagination="69 - 89" refId="ref121688" refString="Gale A. S. 2011. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Departement du Jura, France. Swiss Journal of Palaeontology 130 (1): 69 - 89. http: // dx. doi. org / 10.1007 / s 13358 - 010 - 0008 - x" type="journal article" year="2011">Gale (2011)</bibRefCitation>
|
||
, GZG.INV.78769 (6 dissociated LAPs) from sample S1 of
|
||
<bibRefCitation id="75405B84659285BCD0C0214FFD879BF9" author="Gale A. S." box="[496,639,1126,1153]" pageId="199" pageNumber="200" pagination="69 - 89" refId="ref121688" refString="Gale A. S. 2011. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Departement du Jura, France. Swiss Journal of Palaeontology 130 (1): 69 - 89. http: // dx. doi. org / 10.1007 / s 13358 - 010 - 0008 - x" type="journal article" year="2011">Gale (2011)</bibRefCitation>
|
||
, GZG.INV.78770 (8 dissociated LAPs) from sample S2a of
|
||
<bibRefCitation id="75405B84659285BCD40F214FFEF59BDD" author="Gale A. S." pageId="199" pageNumber="200" pagination="69 - 89" refId="ref121688" refString="Gale A. S. 2011. Asteroidea (Echinodermata) from the Oxfordian (Late Jurassic) of Savigna, Departement du Jura, France. Swiss Journal of Palaeontology 130 (1): 69 - 89. http: // dx. doi. org / 10.1007 / s 13358 - 010 - 0008 - x" type="journal article" year="2011">Gale (2011)</bibRefCitation>
|
||
, all from the
|
||
<emphasis id="23A5FA67659285BCD08921A2FDCA9BDD" box="[441,562,1163,1189]" italics="true" pageId="199" pageNumber="200">Bifurcatus</emphasis>
|
||
Zone, late Oxfordian of Savigna,
|
||
<collectingCountry id="69C666E5659285BCD2EB21A2FBD79BDD" box="[987,1071,1163,1189]" name="France" pageId="199" pageNumber="200">France</collectingCountry>
|
||
; dissociated LAP from the
|
||
<emphasis id="23A5FA67659285BCD18D2186FECE9BB1" box="[189,310,1199,1225]" italics="true" pageId="199" pageNumber="200">Bifurcatus</emphasis>
|
||
Zone, late Oxfordian of Guldental,
|
||
<collectingCountry id="69C666E5659285BCD3C22186FC799BB1" box="[754,897,1199,1225]" name="Switzerland" pageId="199" pageNumber="200">Switzerland</collectingCountry>
|
||
, the original material of
|
||
<bibRefCitation id="75405B84659285BCD5842186FAB29BB2" author="Hess H." box="[1204,1354,1199,1226]" pageId="199" pageNumber="200" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess (1966)</bibRefCitation>
|
||
; 26 dissociated LAPs from the
|
||
<emphasis id="23A5FA67659285BCD0CA21FDFD9E9B95" box="[506,614,1236,1261]" italics="true" pageId="199" pageNumber="200">Renggeri</emphasis>
|
||
Member, early Oxfordian of Chapois,
|
||
<collectingCountry id="69C666E5659285BCD51C21FDFB879B96" box="[1068,1151,1236,1262]" name="France" pageId="199" pageNumber="200">France</collectingCountry>
|
||
, the original material of
|
||
<bibRefCitation id="75405B84659285BCD1EE21D1FE869A6A" author="Hess H." box="[222,382,1271,1298]" pageId="199" pageNumber="200" pagination="1059 - 1082" refId="ref122348" refString="Hess H. 1965 a. Mikropalaontologische Untersuchungen an Ophiuren IV: Die Ophiuren aus dem Renggeri-Ton (Unter-Oxford) von Chapois (Jura) und Longecombe (Ain). Eclogae geologicae Helvetiae 58: 1059 - 1082." type="journal article" year="1965">Hess (1965a)</bibRefCitation>
|
||
; 2 dissociated LAPs from the
|
||
<emphasis id="23A5FA67659285BCD3D621D1FCAA9A69" box="[742,850,1272,1297]" italics="true" pageId="199" pageNumber="200">Renggeri</emphasis>
|
||
Member of Longecombe,
|
||
<collectingCountry id="69C666E5659285BCD5BF21D1FB1A9A6A" box="[1167,1250,1272,1298]" name="France" pageId="199" pageNumber="200">France</collectingCountry>
|
||
, the original material of
|
||
<bibRefCitation id="75405B84659285BCD0742035FE1B9A4E" author="Hess H." box="[324,483,1307,1334]" pageId="199" pageNumber="200" pagination="1059 - 1082" refId="ref122348" refString="Hess H. 1965 a. Mikropalaontologische Untersuchungen an Ophiuren IV: Die Ophiuren aus dem Renggeri-Ton (Unter-Oxford) von Chapois (Jura) und Longecombe (Ain). Eclogae geologicae Helvetiae 58: 1059 - 1082." type="journal article" year="1965">Hess (1965a)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE659285B2D18D2077FB339AD6" lastPageId="201" lastPageNumber="202" pageId="199" pageNumber="200" type="description">
|
||
<paragraph id="116E2675659285BCD18D2077FEB69A00" blockId="199.[189,334,1374,1400]" box="[189,334,1374,1400]" pageId="199" pageNumber="200">
|
||
<heading id="4A269119659285BCD18D2077FEB69A00" bold="true" box="[189,334,1374,1400]" fontSize="11" level="3" pageId="199" pageNumber="200" reason="3">
|
||
<emphasis id="23A5FA67659285BCD18D2077FEB69A00" bold="true" box="[189,334,1374,1400]" pageId="199" pageNumber="200">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E2675659285BCD1DA20A7FEA49AD0" blockId="199.[189,1399,1422,2027]" box="[234,348,1422,1448]" pageId="199" pageNumber="200">
|
||
<emphasis id="23A5FA67659285BCD1DA20A7FEA49AD0" bold="true" box="[234,348,1422,1448]" pageId="199" pageNumber="200">
|
||
<typeStatus id="CE6A98D7659285BCD1DA20A7FEA49AD0" box="[234,348,1422,1448]" pageId="199" pageNumber="200" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="116E2675659285BCD18D2098FC8E9806" blockId="199.[189,1399,1422,2027]" pageId="199" pageNumber="200">GZG.INV.78763 is a dissociated, large, proximal LAP; nearly as high as wide; dorsal edge strongly concave as a result of a well-developed constriction; distal edge strongly convex; proximal edge irregularly undulose, with a large, sharply defined, prominent and strongly protruding, oval, horizontally elongate spur composed of densely meshed stereom; weakly defined, shallow depression in ventral half of proximal edge, composed of more finely meshed stereom; outer surface with finely meshed stereom, fining towards proximal edge. Five large ear-shaped spine articulations freestanding on strongly elevated distal half of LAP; strong dorsalward increase in size of spine articulations and of gaps separating them; ventral and dorsal lobes merged into continuous volute; dorsal lobe with conspicuous central notch; gap between spine articulations and distal edge of LAP as wide as one spine articulation, strongly increasing in width dorsalwards, parallel to dorsalward increase in size of spine articulations; dorsalmost spine articulation below dorsal edge of LAP, pointing dorsalwards; spine articulations proximally sharply bordered by undulose edge of elevated distal half of LAP. Ventral edge of LAP nearly straight, with very small, weakly concave, almost indiscernible tentacle notch.</paragraph>
|
||
<paragraph id="116E2675659285B2D18D22A0FE659E31" blockId="199.[189,1399,1422,2027]" lastBlockId="201.[189,1398,267,329]" lastPageId="201" lastPageNumber="202" pageId="199" pageNumber="200">Inner side of LAP with very large, broad, sharply defined, prominent ridge with evenly bent, ventroproximally pointing ventral part, and strongly widened, near-triangular dorsal part; inner side of distal edge of LAP with large, well-defined, rounded, non-prominent spur composed of densely meshed stereom; inner side of tentacle notch very small, semi-circular, well defined laterally. No perforations or furrow discernible.</paragraph>
|
||
<caption id="45AE76FD659D85B3D18D23F3FA8E9893" ID-DOI="http://doi.org/10.5281/zenodo.3822911" ID-Zenodo-Dep="3822911" httpUri="https://zenodo.org/record/3822911/files/figure.png" pageId="200" pageNumber="201" startId="200.[189,231,1754,1780]" targetBox="[258,1330,265,1715]" targetPageId="200">
|
||
<paragraph id="116E2675659D85B3D18D23F3FA8E9893" blockId="200.[189,1399,1754,2027]" pageId="200" pageNumber="201">
|
||
<emphasis id="23A5FA67659D85B3D18D23F3FEEE998C" bold="true" box="[189,278,1754,1780]" pageId="200" pageNumber="201">Fig. 35.</emphasis>
|
||
Skeletal plates and arm fragments of fossil and Recent ophiacanthid brittle stars; lateral arm plates (LAPs) in external (a) and internal (b) views.
|
||
<emphasis id="23A5FA67659D85B3D3FF23D4FD04986F" bold="true" box="[719,764,1789,1815]" pageId="200" pageNumber="201">1-3.</emphasis>
|
||
<taxonomicName id="D6D15DF6659D85B3D23423D4FB7D9860" authority="Lyman, 1878" authorityName="Lyman" authorityYear="1878" box="[772,1157,1789,1816]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="200" pageNumber="201" phylum="Echinodermata" rank="species" species="vitrea">
|
||
<emphasis id="23A5FA67659D85B3D23423D4FC19986F" box="[772,993,1789,1815]" italics="true" pageId="200" pageNumber="201">Ophiocamax vitrea</emphasis>
|
||
Lyman, 1878
|
||
</taxonomicName>
|
||
, Recent.
|
||
<emphasis id="23A5FA67659D85B3D5DF23D4FAFD986F" bold="true" box="[1263,1285,1789,1815]" pageId="200" pageNumber="201">1.</emphasis>
|
||
Proximal LAP.
|
||
<emphasis id="23A5FA67659D85B3D0312208FEEF9843" bold="true" box="[257,279,1825,1851]" pageId="200" pageNumber="201">2.</emphasis>
|
||
Median LAP.
|
||
<emphasis id="23A5FA67659D85B3D0F42208FE229843" bold="true" box="[452,474,1825,1851]" pageId="200" pageNumber="201">3.</emphasis>
|
||
Dorsal arm plate.
|
||
<emphasis id="23A5FA67659D85B3D3872208FD1D9843" bold="true" box="[695,741,1825,1851]" pageId="200" pageNumber="201">4-5.</emphasis>
|
||
<taxonomicName id="D6D15DF6659D85B3D3DE2208FB789843" authority="Lyman, 1878" authorityName="Lyman" authorityYear="1878" box="[750,1152,1825,1851]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="200" pageNumber="201" phylum="Echinodermata" rank="species" species="hystrix">
|
||
<emphasis id="23A5FA67659D85B3D3DE2208FC229843" box="[750,986,1825,1851]" italics="true" pageId="200" pageNumber="201">Ophiocamax hystrix</emphasis>
|
||
Lyman, 1878
|
||
</taxonomicName>
|
||
, Recent.
|
||
<emphasis id="23A5FA67659D85B3D5DD2208FAFB9843" bold="true" box="[1261,1283,1825,1851]" pageId="200" pageNumber="201">4.</emphasis>
|
||
Proximal LAP.
|
||
<emphasis id="23A5FA67659D85B3D031226DFEEF9826" bold="true" box="[257,279,1860,1886]" pageId="200" pageNumber="201">5.</emphasis>
|
||
Median LAP.
|
||
<emphasis id="23A5FA67659D85B3D0F6226DFE239826" bold="true" box="[454,475,1860,1886]" pageId="200" pageNumber="201">6.</emphasis>
|
||
<taxonomicName id="D6D15DF6659D85B3D0D5226DFC819826" authority="Verrill, 1899" authorityName="Verrill" authorityYear="1899" box="[485,889,1860,1886]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="200" pageNumber="201" phylum="Echinodermata" rank="species" species="austera">
|
||
<emphasis id="23A5FA67659D85B3D0D5226DFD219826" box="[485,729,1860,1886]" italics="true" pageId="200" pageNumber="201">Ophiocamax austera</emphasis>
|
||
Verrill, 1899
|
||
</taxonomicName>
|
||
, Recent; proximal LAP.
|
||
<emphasis id="23A5FA67659D85B3D5AE226DFB219826" bold="true" box="[1182,1241,1860,1886]" pageId="200" pageNumber="201">7-11.</emphasis>
|
||
<taxonomicName id="D6D15DF6659D85B3D5D2226DFECA98F9" authority="Thuy, 2013" authorityName="Thuy" authorityYear="2013" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="200" pageNumber="201" phylum="Echinodermata" rank="species" species="dorotheae" status="sp. nov.">
|
||
<emphasis id="23A5FA67659D85B3D5D2226DFECA98F9" italics="true" pageId="200" pageNumber="201">Ophiocamax dorotheae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3896471C659D85B3D00E2241FE6698FA" box="[318,414,1896,1922]" pageId="200" pageNumber="201" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the late Oxfordian (Late Jurassic) of Savigna, France.
|
||
<emphasis id="23A5FA67659D85B3D5A1224EFB5F98F9" bold="true" box="[1169,1191,1895,1921]" pageId="200" pageNumber="201">7.</emphasis>
|
||
GZG.INV.78763 (holotype), proximal LAP.
|
||
<emphasis id="23A5FA67659D85B3D0CB22A3FDE998DC" bold="true" box="[507,529,1930,1956]" pageId="200" pageNumber="201">8.</emphasis>
|
||
GZG.INV.78764 (paratype), median LAP.
|
||
<emphasis id="23A5FA67659D85B3D53E22A3FBDC98DC" bold="true" box="[1038,1060,1930,1956]" pageId="200" pageNumber="201">9.</emphasis>
|
||
GZG.INV.78765 (paratype), distal LAP.
|
||
<emphasis id="23A5FA67659D85B3D0742284FE9198BF" bold="true" box="[324,361,1965,1991]" pageId="200" pageNumber="201">10.</emphasis>
|
||
GZG.INV.78766 (paratype), proximal arm fragment in ventral view.
|
||
<emphasis id="23A5FA67659D85B3D5BA2284FB5398BF" bold="true" box="[1162,1195,1965,1991]" pageId="200" pageNumber="201">11.</emphasis>
|
||
GZG.INV.78767 (paratype), median arm fragment in dorsal view. One common scale bar per species except for 10 and 11.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="116E2675659C85B2D1DA2458FD489EF3" blockId="201.[189,1399,369,647]" box="[234,688,369,395]" pageId="201" pageNumber="202">
|
||
<emphasis id="23A5FA67659C85B2D1DA2458FD489EF3" bold="true" box="[234,688,369,395]" pageId="201" pageNumber="202">
|
||
<typeStatus id="CE6A98D7659C85B2D1DA2458FEA49EF3" box="[234,348,369,395]" pageId="201" pageNumber="202" type="paratype">Paratype</typeStatus>
|
||
supplements and variation
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="116E2675659C85B2D18D24BCFE919D46" blockId="201.[189,1399,369,647]" pageId="201" pageNumber="202">
|
||
GZG.INV.78764 is a dissociated median LAP; slightly wider than high; very well in agreement with
|
||
<typeStatus id="CE6A98D7659C85B2D18D2490FEDF9EAB" box="[189,295,441,467]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
; dorsal edge less strongly concave; spur on proximal edge slightly smaller than in
|
||
<typeStatus id="CE6A98D7659C85B2D4382490FA8B9EAB" box="[1288,1395,441,467]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
. Five spine articulations similar to those of
|
||
<typeStatus id="CE6A98D7659C85B2D3F524F4FCD79E8F" box="[709,815,477,503]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
; gap between spine articulations and distal edge of LAP slightly narrower. Ventral edge of LAP weakly concave, with very small, almost indiscernible tentacle notch.
|
||
</paragraph>
|
||
<paragraph id="116E2675659C85B2D18D2760FBC69DFF" blockId="201.[189,1399,369,647]" pageId="201" pageNumber="202">Inner side of LAP with very large, well-defined, short ridge with strongly widened ventral and dorsal edges; inner side of tentacle notch very small, sharply defined, semi circular.</paragraph>
|
||
<paragraph id="116E2675659C85B2D18D2787FB3E9C2F" blockId="201.[189,1399,686,963]" pageId="201" pageNumber="202">
|
||
GZG.INV.78765 is a dissociated distal LAP; almost twice wider than high; very well in agreement with
|
||
<typeStatus id="CE6A98D7659C85B2D1CB27FBFE9D9D94" box="[251,357,722,748]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
; depression in ventral half of outer surface smaller and even less well defined than in
|
||
<typeStatus id="CE6A98D7659C85B2D18D27DFFEDF9C68" box="[189,295,758,784]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
, almost indiscernible. Four spine articulations similar to those of
|
||
<typeStatus id="CE6A98D7659C85B2D50727DFFB599C68" box="[1079,1185,758,784]" pageId="201" pageNumber="202" type="holotype">holotype</typeStatus>
|
||
; dorsalmost spine articulation not pointing dorsalwards; gap between spine articulations and distal edge of LAP much narrower. Ventral edge of LAP gently concave; tentacle notch invisible in external view.
|
||
</paragraph>
|
||
<paragraph id="116E2675659C85B2D18D264BFBBE9CBB" blockId="201.[189,1399,686,963]" pageId="201" pageNumber="202">Inner side of LAP with large, poorly defined, broad, weakly prominent ridge displaying strongly widened ventral part and dorso-proximalwards pointing dorsal tip; spur on inner side of distal edge slightly less well defined; inner side of tentacle notch very small, more than semi circular.</paragraph>
|
||
<paragraph id="116E2675659C85B2D18D26C2FD8A9A5B" blockId="201.[189,1399,1003,1316]" pageId="201" pageNumber="202">GZG.INV.78766 is an articulated arm fragment preserving two median segments; LAPs well in agreement with dissociated ones described above; dorsal arm plates large, widely separated by LAPs, slightly longer than wide, widest distally, irregularly pentagonal to blazon shaped, with gently convex distal edge, slightly proximalwards converging lateral edges and nearly right proximal angle; outer surface of dorsal arm plates with small, scattered granules or thorns; ventral arm plates widely separated by LAPs; slighty wider than long, widest distally, with gently convex to slightly angular distal edge, short, straight lateral edges, small half-circular tentacle notches and obtuse proximal angle; few very short arm fragments preserved, originally cylindrical, with moderately coarsely meshed outer surface stereom, no further details preserved.</paragraph>
|
||
<paragraph id="116E2675659C85B2D18D2062FB339AD6" blockId="201.[189,1399,1355,1454]" pageId="201" pageNumber="202">GZG.INV.78767 is an articulated arm fragment preserving three median to distal segments; one second ventralmost arm spine nearly completely preserved, straight, cylindrical, almost as long as one arm segment, with moderately coarsely meshed and slightly longitudinally elongate stereom.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE659C85B1D18D20FFFD0C9ECC" lastPageId="202" lastPageNumber="203" pageId="201" pageNumber="202" type="discussion">
|
||
<paragraph id="116E2675659C85B2D18D20FFFED79A88" blockId="201.[189,303,1494,1520]" box="[189,303,1494,1520]" pageId="201" pageNumber="202">
|
||
<heading id="4A269119659C85B2D18D20FFFED79A88" bold="true" box="[189,303,1494,1520]" fontSize="11" level="3" pageId="201" pageNumber="202" reason="3">
|
||
<emphasis id="23A5FA67659C85B2D18D20FFFED79A88" bold="true" box="[189,303,1494,1520]" pageId="201" pageNumber="202">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E2675659C85B2D18D232DFE539846" blockId="201.[189,1399,1540,1854]" pageId="201" pageNumber="202">
|
||
The highly distinctive shape and position of the spine articulations, combined with the lack of a conspicuous outer surface ornament, the distinctively shaped ridge on the inner side and the very small, well-defined, semi-circular tentacle notch unambiguously place these LAPs in the genus
|
||
<taxonomicName id="D6D15DF6659C85B2D5F22365FAA0991E" authorityName="Lyman" authorityYear="1878" box="[1218,1368,1612,1638]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="201" pageNumber="202" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA67659C85B2D5F22365FAA0991E" box="[1218,1368,1612,1638]" italics="true" pageId="201" pageNumber="202">Ophiocamax</emphasis>
|
||
</taxonomicName>
|
||
or at least in a very closely related form (more closely related than to
|
||
<taxonomicName id="D6D15DF6659C85B2D2FC2359FBAD99F2" authorityName="Lyman" authorityYear="1869" box="[972,1109,1648,1674]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiomitra" kingdom="Animalia" order="Ophiurida" pageId="201" pageNumber="202" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA67659C85B2D2FC2359FBAD99F2" box="[972,1109,1648,1674]" italics="true" pageId="201" pageNumber="202">Ophiomitra</emphasis>
|
||
</taxonomicName>
|
||
, on account of the much closer similarities in LAP morphology). The low number of spine articulations and the very weakly ventralwards protruding row of spine articulations precludes assignment to the closely related Late Jurassic
|
||
<taxonomicName id="D6D15DF6659C85B2D01123F2FCFD998E" authority="(Quenstedt, 1876)" baseAuthorityName="Quenstedt" baseAuthorityYear="1876" box="[289,773,1755,1782]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiosternle" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="201" pageNumber="202" phylum="Echinodermata" rank="species" species="crinitum">
|
||
<emphasis id="23A5FA67659C85B2D01123F2FDDD998D" box="[289,549,1755,1781]" italics="true" pageId="201" pageNumber="202">Ophiosternle crinitum</emphasis>
|
||
(Quenstedt, 1876)
|
||
</taxonomicName>
|
||
. The present LAPs are by far the oldest record that is assignable to
|
||
<taxonomicName id="D6D15DF6659C85B2D06B23D6FE099861" authorityName="Lyman" authorityYear="1878" box="[347,497,1791,1817]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="201" pageNumber="202" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA67659C85B2D06B23D6FE099861" box="[347,497,1791,1817]" italics="true" pageId="201" pageNumber="202">Ophiocamax</emphasis>
|
||
</taxonomicName>
|
||
, predating the Miocene records from
|
||
<collectingCountry id="69C666E5659C85B2D2992229FC149862" box="[937,1004,1792,1818]" name="Japan" pageId="201" pageNumber="202">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="75405B84659C85B2D2CA23D6FB7E9861" author="Ishida Y." box="[1018,1158,1791,1817]" pageId="201" pageNumber="202" pagination="53 - 59" refId="ref123173" refString="Ishida Y. 2001. Cenozoic ophiuroids from Japan; particularly those conspecific with extant species. In: J. - P. Feral & B. David (eds) Echinoderm Research 2001: 53 - 59. Swets & Zeitlinger, Lisse." type="journal article" year="2001">Ishida 2001</bibRefCitation>
|
||
) and
|
||
<collectingCountry id="69C666E5659C85B2D5F623D6FA8F9861" box="[1222,1399,1791,1817]" name="Saint Vincent and the Grenadines" pageId="201" pageNumber="202">the Grenadines</collectingCountry>
|
||
(Jagt
|
||
<emphasis id="23A5FA67659C85B2D1CD220DFEC09845" box="[253,312,1827,1853]" italics="true" pageId="201" pageNumber="202">et al.</emphasis>
|
||
in press).
|
||
</paragraph>
|
||
<paragraph id="116E2675659C85B1D18D224CFD0C9ECC" blockId="201.[189,1398,1893,2027]" lastBlockId="202.[189,1399,267,437]" lastPageId="202" lastPageNumber="203" pageId="201" pageNumber="202">
|
||
The present LAPs differ in that the two dorsalmost spine articulations invariably are the largest, rather than much smaller than the remaining ones as in the LAPs of most extant species of
|
||
<taxonomicName id="D6D15DF6659C85B2D5E922A0FA9798DB" authorityName="Lyman" authorityYear="1878" box="[1241,1391,1929,1955]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="201" pageNumber="202" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="23A5FA67659C85B2D5E922A0FA9798DB" box="[1241,1391,1929,1955]" italics="true" pageId="201" pageNumber="202">Ophiocamax</emphasis>
|
||
</taxonomicName>
|
||
. Greatest similarities are shared with the LAPs of extant
|
||
<taxonomicName id="D6D15DF6659C85B2D2752284FC5398BF" authorityName="Lyman" authorityYear="1878" box="[837,939,1965,1991]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="201" pageNumber="202" phylum="Echinodermata" rank="species" species="vitrea">
|
||
<emphasis id="23A5FA67659C85B2D2752284FC5398BF" box="[837,939,1965,1991]" italics="true" pageId="201" pageNumber="202">O. vitrea</emphasis>
|
||
</taxonomicName>
|
||
, especially on account of the shape and position of the spine articulations, the presence of a well-developed spur on the outer proximal and inner distal edge, and the development of the tentacle notch. In addition, the shape and outer surface ornament of the dorsal arm plates is markedly similar in
|
||
<taxonomicName id="D6D15DF6659F85B1D3C02419FCA19E31" authorityName="Lyman" authorityYear="1878" box="[752,857,304,330]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="202" pageNumber="203" phylum="Echinodermata" rank="species" species="vitrea">
|
||
<emphasis id="23A5FA67659F85B1D3C02419FCA19E31" box="[752,857,304,330]" italics="true" pageId="202" pageNumber="203">O. vitrea</emphasis>
|
||
</taxonomicName>
|
||
and these fossil specimens. The shape of the ridge on the inner side, in contrast, is more reminiscent of that observed in the LAPs of
|
||
<taxonomicName id="D6D15DF6659F85B1D5E2247AFAB69E15" authorityName="Verrill" authorityYear="1899" box="[1234,1358,339,365]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiocamax" kingdom="Animalia" order="Ophiurida" pageId="202" pageNumber="203" phylum="Echinodermata" rank="species" species="austera">
|
||
<emphasis id="23A5FA67659F85B1D5E2247AFAB69E15" box="[1234,1358,339,365]" italics="true" pageId="202" pageNumber="203">O. austera</emphasis>
|
||
</taxonomicName>
|
||
. In the light of these minor, yet significant, morphological differences and the huge stratigraphic gap, the fossil LAPs are here described as a new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="59CB75FE659F85B1D18D24F5FC4F9D5D" box="[189,951,476,550]" pageId="202" pageNumber="203" type="distribution">
|
||
<paragraph id="116E2675659F85B1D18D24F5FEB79E8E" blockId="202.[189,335,476,502]" box="[189,335,476,502]" pageId="202" pageNumber="203">
|
||
<heading id="4A269119659F85B1D18D24F5FEB79E8E" bold="true" box="[189,335,476,502]" fontSize="11" level="3" pageId="202" pageNumber="203" reason="3">
|
||
<emphasis id="23A5FA67659F85B1D18D24F5FEB79E8E" bold="true" box="[189,335,476,502]" pageId="202" pageNumber="203">Occurrence</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="116E2675659F85B1D18D2722FC4F9D5D" blockId="202.[189,951,523,550]" box="[189,951,523,550]" pageId="202" pageNumber="203">
|
||
Early to late Oxfordian of
|
||
<collectingCountry id="69C666E5659F85B1D0C32725FDBE9D5E" box="[499,582,524,550]" name="France" pageId="202" pageNumber="203">France</collectingCountry>
|
||
, late Oxfordian of
|
||
<collectingCountry id="69C666E5659F85B1D2132722FC4B9D5D" box="[803,947,523,549]" name="Switzerland" pageId="202" pageNumber="203">Switzerland</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |