treatments-xml/data/99/78/97/99789763658585A9D0962134FD10996A.xml
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<document id="F93FEA9B90F781AA90BAAE7F994BEDE4" ID-CLB-Dataset="22249" ID-DOI="10.5852/ejt.2013.48" ID-GBIF-Dataset="02866e62-ab61-4cf4-9d98-0248e251084e" ID-ISSN="2118-9773" ID-Zenodo-Dep="3822836" ID-ZooBank="E7080722-E348-448D-96E5-D537F4865BB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1589219740188" checkinUser="carolina" docAuthor="Thuy, Ben" docDate="2013" docId="99789763658585A9D0962134FD10996A" docLanguage="en" docName="ejt-2013-48.pdf.imf" docOrigin="European Journal of Taxonomy 48" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiochondrus punctatus Thuy 2013, comb. nov." docType="treatment" docVersion="9" lastPageNumber="211" masterDocId="6541EF1B6555857BD1302529FFF89F78" masterDocTitle="Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)" masterLastPageNumber="242" masterPageNumber="1" pageNumber="209" updateTime="1698839159859" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B8B5AAD48C1F91694B4C1FCBCF2AA8D7">Temporary expansion to shelf depths rather than an onshore-offshore trend: the shallow-water rise and demise of the modern deep-sea brittle star family Ophiacanthidae (Echinodermata: Ophiuroidea)</mods:title>
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<mods:namePart id="2909D88864429AE6D14E28477B94F524">Thuy, Ben</mods:namePart>
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<mods:title id="B04562F86ECC0856D639AEED829CCD1E">European Journal of Taxonomy</mods:title>
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<mods:date id="5D0793D9B3A6B71036FF12767B7CAA9B">2013</mods:date>
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<treatment id="99789763658585A9D0962134FD10996A" ID-DOI="http://doi.org/10.5281/zenodo.3844097" ID-GBIF-Taxon="164044795" ID-Zenodo-Dep="3844097" LSID="urn:lsid:plazi:treatment:99789763658585A9D0962134FD10996A" httpUri="http://treatment.plazi.org/id/99789763658585A9D0962134FD10996A" lastPageId="210" lastPageNumber="211" pageId="208" pageNumber="209">
<subSubSection id="59CB75FE658585ABD0962134FCA69B24" pageId="208" pageNumber="209" type="nomenclature">
<paragraph id="116E2675658585ABD0962134FCA69B24" blockId="208.[422,1166,1053,1116]" pageId="208" pageNumber="209">
<taxonomicName id="D6D15DF6658585ABD0962134FBFD9B40" authority="(Kutscher &amp; Jagt, 2000)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Kutscher &amp; Jagt" baseAuthorityYear="2000" box="[422,1029,1053,1080]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="208" pageNumber="209" phylum="Echinodermata" rank="species" species="punctatus" status="comb. nov.">
<emphasis id="23A5FA67658585ABD0962134FD1A9B40" bold="true" box="[422,738,1053,1080]" italics="true" pageId="208" pageNumber="209">Ophiochondrus punctatus</emphasis>
(
<bibRefCitation id="75405B84658585ABD3C12134FC049B40" author="Kutscher M. &amp; Jagt J. W. M." box="[753,1020,1053,1080]" pageId="208" pageNumber="209" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="3896471C658585ABD53C2134FB769B40" box="[1036,1166,1053,1080]" pageId="208" pageNumber="209" rank="species">comb. nov.</taxonomicNameLabel>
<figureCitation id="89EA3AF0658585ABD3E52168FCD49B24" box="[725,812,1089,1116]" captionStart="Fig" captionStartId="209.[189,232,1648,1674]" captionTargetBox="[297,1290,265,1608]" captionTargetId="figure@209.[297,1290,265,1608]" captionTargetPageId="209" captionText="Fig. 37. Lateral arm plates (LAPs) of fossil and Recent ophiacanthid brittle stars in external (a) and internal (b) views. 1-2. Ophiochondrus convolutus Lyman, 1869, Recent. 1. Proximal LAP. 2. Median LAP. 3. Ophiochondrus stelliger Lyman, 1879, Recent; proximal LAP. 4-5. Ophiochondrus punctatus (Kutscher &amp; Jagt, 2000) comb. nov. from the early Maastrichtian (Early Cretaceous) of Rügen, Germany. 4. GZG.INV.78775, proximal LAP. 5. GZG.INV.78776, distal LAP. 6-7. Ophiochondrus? semirotundus (Kutscher &amp; Jagt, 2000) comb. nov. from the early Maastrichtian of Rügen, Germany. 6. GZG.INV.78800, proximal LAP. 7. GZG.INV.78801, median LAP. 8-9. Ophiacanthidae gen. et sp. nov. innom. from the late Oxfordian (Late Jurassic) of the Plettenberg, Germany. 8. GZG.INV.78802, proximal LAP. 9. GZG.INV.78803, median or distal LAP. 10-11. Ophiojagtus argoviensis (Hess, 1966) comb. nov. from the late Oxfordian (Late Jurassic) of Savigna, France. 10. NHMB M1124, proximal LAP. 11. NHMB M11225, proximal to median LAP. One common scale bar per species." figureDoi="http://doi.org/10.5281/zenodo.3822915" httpUri="https://zenodo.org/record/3822915/files/figure.png" pageId="208" pageNumber="209">Fig. 37</figureCitation>
: 4-5
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658585ABD18D21ADFC159BE7" box="[189,1005,1156,1183]" pageId="208" pageNumber="209" type="reference_group">
<paragraph id="116E2675658585ABD18D21ADFC159BE7" blockId="208.[189,1005,1156,1183]" box="[189,1005,1156,1183]" pageId="208" pageNumber="209">
<treatmentCitationGroup id="31C1015B658585ABD18D21ADFC159BE7" box="[189,1005,1156,1183]" pageId="208" pageNumber="209">
<taxonomicName id="D6D15DF6658585ABD18D21ADFCCF9BE6" authority="Kutscher &amp; Jagt, 2000: 62 - 63" authorityName="Muller &amp; Troschel" authorityPageNumber="62 - 63" authorityYear="1842" box="[189,823,1156,1182]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="208" pageNumber="209" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658585ABD18D21ADFEAE9BE6" box="[189,342,1156,1182]" italics="true" pageId="208" pageNumber="209">Ophiacantha</emphasis>
?
<treatmentCitation id="90700064658585ABD05A21ACFCCF9BE6" author="Kutscher M. &amp; Jagt J. W. M." box="[362,823,1156,1182]" page="63" pageId="208" pageNumber="209" year="2000">
<emphasis id="23A5FA67658585ABD05A21ACFE299BE6" box="[362,465,1157,1182]" italics="true" pageId="208" pageNumber="209">punctata</emphasis>
<bibRefCitation id="75405B84658585ABD0E821ADFCCF9BE6" author="Kutscher M. &amp; Jagt J. W. M." box="[472,823,1156,1182]" pageId="208" pageNumber="209" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000: 62-63</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, pl. 25 figs 3-5.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658585ABD18D21EFFCE19A9F" pageId="208" pageNumber="209" type="diagnosis">
<paragraph id="116E2675658585ABD18D21EFFECE9B98" blockId="208.[189,310,1222,1248]" box="[189,310,1222,1248]" pageId="208" pageNumber="209">
<heading id="4A269119658585ABD18D21EFFECE9B98" bold="true" box="[189,310,1222,1248]" fontSize="11" level="3" pageId="208" pageNumber="209" reason="3">
<emphasis id="23A5FA67658585ABD18D21EFFECE9B98" bold="true" box="[189,310,1222,1248]" pageId="208" pageNumber="209">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658585ABD18D21DCFCE19A9F" blockId="208.[189,1399,1269,1512]" pageId="208" pageNumber="209">
Species of
<taxonomicName id="D6D15DF6658585ABD00E21DCFE0D9A77" authorityName="Lyman" authorityYear="1869" box="[318,501,1269,1295]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="208" pageNumber="209" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658585ABD00E21DCFE0D9A77" box="[318,501,1269,1295]" italics="true" pageId="208" pageNumber="209">Ophiochondrus</emphasis>
</taxonomicName>
with small LAPs displaying a coarsely meshed stereom on the outer surface with trabecular intersections strongly thickened into slightly vertically elongate granules or knobs; small, very poorly defined, weakly prominent and protruding spur on the outer proximal edge, paralleled on the inner distal edge by a moderately well-defined, rounded area of densely meshed stereom; up to eight small, slightly oval, spine articulations, proximally bordered by thick, slightly knobby ridge; gap between spine articulations and distal edge of LAP with trabecular intersections thickened into small granules; ridge on inner side relatively small, short.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658585ABD18D2326FC569905" pageId="208" pageNumber="209" type="materials_examined">
<paragraph id="116E2675658585ABD18D2326FE529951" blockId="208.[189,426,1551,1577]" box="[189,426,1551,1577]" pageId="208" pageNumber="209">
<heading id="4A269119658585ABD18D2326FE529951" bold="true" box="[189,426,1551,1577]" fontSize="11" level="3" pageId="208" pageNumber="209" reason="3">
<emphasis id="23A5FA67658585ABD18D2326FE529951" bold="true" box="[189,426,1551,1577]" pageId="208" pageNumber="209">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658585ABD18D2316FC569905" blockId="208.[189,1399,1599,1662]" pageId="208" pageNumber="209">
GZG.INV.78775, GZG.INV.78776 and 260 dissociated LAPs in the Manfred Kutscher Collection in Sassnitz,
<collectingCountry id="69C666E5658585ABD01C234AFE629905" box="[300,410,1635,1661]" name="Germany" pageId="208" pageNumber="209">Germany</collectingCountry>
, the
<typeStatus id="CE6A98D7658585ABD0E1234DFDFC9906" box="[465,516,1636,1662]" pageId="208" pageNumber="209">type</typeStatus>
material of
<bibRefCitation id="75405B84658585ABD3A2234AFC529905" author="Kutscher M. &amp; Jagt J. W. M." box="[658,938,1635,1662]" pageId="208" pageNumber="209" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt (2000)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658585A9D18D238CFB3B9C7F" lastPageId="210" lastPageNumber="211" pageId="208" pageNumber="209" type="description">
<paragraph id="116E2675658585ABD18D238CFEB699C7" blockId="208.[189,334,1701,1727]" box="[189,334,1701,1727]" pageId="208" pageNumber="209">
<heading id="4A269119658585ABD18D238CFEB699C7" bold="true" box="[189,334,1701,1727]" fontSize="11" level="3" pageId="208" pageNumber="209" reason="3">
<emphasis id="23A5FA67658585ABD18D238CFEB699C7" bold="true" box="[189,334,1701,1727]" pageId="208" pageNumber="209">Description</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658585A9D18D23FCFDCD9D50" blockId="208.[189,1399,1748,2027]" lastBlockId="210.[189,1399,267,775]" lastPageId="210" lastPageNumber="211" pageId="208" pageNumber="209">Small, dissociated LAPs, proximal ones slightly higher than wide, distal ones approximately 1.5 times wider than high; proximal to median LAPs of massive, thick, rounded aspect; dorsal edge straight (proximal LAPs) to concave (median and distal LAPs) as a result of a well-developed constriction; distal edge convex; proximal edge nearly straight, with small, very poorly defined, weakly prominent and very slightly protruding spur ventrally bordered by small, poorly defined, non-depressed area of slightly more finely meshed stereom; outer surface with coarsely meshed stereom slightly fining towards proximal edge; trabecular intersections of coarsely meshed stereom thickened into large, very weakly vertically elongate granules or knobs. Eight (proximal LAPs) to six (distal LAPs) small, ear-shaped, slightly oval spine articulations freestanding on weakly to fairly elevated distal portion of LAP; spine articulations proximally bordered by thick, well-defined, knobby ridge composed of vertically merged thickened trabeculae of coarsely meshed outer surface stereom; ventral and dorsal lobes of spine articulations separated proximally by very small notch; spine articulations nearly of equal size or with very weak dorsalward decrease in size; gap between spine articulations and distal edge of LAP at most as wide as one spine articulation, with dorsalward increase in width, with trabecular intersections of moderately finely meshed stereom thickened into small granules. Ventral edge of LAP weakly concave; tentacle notch invisible in external view.</paragraph>
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<paragraph id="116E2675658485AAD18D2359FB319893" blockId="209.[189,1399,1648,2027]" pageId="209" pageNumber="210">
<emphasis id="23A5FA67658485AAD18D2359FEE399F3" bold="true" box="[189,283,1648,1675]" pageId="209" pageNumber="210">Fig. 37.</emphasis>
Lateral arm plates (LAPs) of fossil and Recent ophiacanthid brittle stars in external (a) and internal (b) views.
<emphasis id="23A5FA67658485AAD0AE23BDFE3499D6" bold="true" box="[414,460,1684,1710]" pageId="209" pageNumber="210">1-2.</emphasis>
<taxonomicName id="D6D15DF6658485AAD0E523BDFC4199D6" authority="Lyman, 1869" authorityName="Lyman" authorityYear="1869" box="[469,953,1684,1710]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="209" pageNumber="210" phylum="Echinodermata" rank="species" species="convolutus">
<emphasis id="23A5FA67658485AAD0E523BDFCEB99D6" box="[469,787,1684,1710]" italics="true" pageId="209" pageNumber="210">Ophiochondrus convolutus</emphasis>
Lyman, 1869
</taxonomicName>
, Recent.
<emphasis id="23A5FA67658485AAD51823BDFBC699D6" bold="true" box="[1064,1086,1684,1710]" pageId="209" pageNumber="210">1.</emphasis>
Proximal LAP.
<emphasis id="23A5FA67658485AAD5CE23BDFAEC99D6" bold="true" box="[1278,1300,1684,1710]" pageId="209" pageNumber="210">2.</emphasis>
Median LAP.
<emphasis id="23A5FA67658485AAD031239EFEEF99A9" bold="true" box="[257,279,1719,1745]" pageId="209" pageNumber="210">3.</emphasis>
<taxonomicName id="D6D15DF6658485AAD010239EFD1F99A9" authority="Lyman, 1879" authorityName="Lyman" authorityYear="1879" box="[288,743,1719,1746]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="209" pageNumber="210" phylum="Echinodermata" rank="species" species="stelliger">
<emphasis id="23A5FA67658485AAD010239EFDC799A9" box="[288,575,1719,1745]" italics="true" pageId="209" pageNumber="210">Ophiochondrus stelliger</emphasis>
Lyman, 1879
</taxonomicName>
, Recent; proximal LAP.
<emphasis id="23A5FA67658485AAD53E239EFBC499A9" bold="true" box="[1038,1084,1719,1745]" pageId="209" pageNumber="210">4-5.</emphasis>
<taxonomicName id="D6D15DF6658485AAD575239EFE17998C" authority="(Kutscher &amp; Jagt, 2000)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Kutscher &amp; Jagt" baseAuthorityYear="2000" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="209" pageNumber="210" phylum="Echinodermata" rank="species" species="punctatus" status="comb. nov.">
<emphasis id="23A5FA67658485AAD575239EFA8E99A9" box="[1093,1398,1719,1745]" italics="true" pageId="209" pageNumber="210">Ophiochondrus punctatus</emphasis>
(
<bibRefCitation id="75405B84658485AAD1F523F3FE1E998C" author="Kutscher M. &amp; Jagt J. W. M." box="[197,486,1754,1781]" pageId="209" pageNumber="210" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="3896471C658485AAD0CD23F3FD7E998D" box="[509,646,1754,1781]" pageId="209" pageNumber="210" rank="species">comb. nov.</taxonomicNameLabel>
from the early Maastrichtian (Early Cretaceous) of Rügen, Germany.
<emphasis id="23A5FA67658485AAD00B23D4FEA9986F" bold="true" box="[315,337,1789,1815]" pageId="209" pageNumber="210">4.</emphasis>
GZG.INV.78775, proximal LAP.
<emphasis id="23A5FA67658485AAD3DF23D7FCFD9860" bold="true" box="[751,773,1790,1816]" pageId="209" pageNumber="210">5.</emphasis>
GZG.INV.78776, distal LAP.
<emphasis id="23A5FA67658485AAD54923D4FB50986F" bold="true" box="[1145,1192,1789,1815]" pageId="209" pageNumber="210">6-7.</emphasis>
<taxonomicName id="D6D15DF6658485AAD58323D4FA92986F" authorityName="Lyman" authorityYear="1869" box="[1203,1386,1789,1815]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="209" pageNumber="210" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658485AAD58323D4FA92986F" box="[1203,1386,1789,1815]" italics="true" pageId="209" pageNumber="210">Ophiochondrus</emphasis>
</taxonomicName>
?
<emphasis id="23A5FA67658485AAD18D2208FEA29843" box="[189,346,1825,1851]" italics="true" pageId="209" pageNumber="210">semirotundus</emphasis>
(
<bibRefCitation id="75405B84658485AAD05C2208FD789843" author="Kutscher M. &amp; Jagt J. W. M." box="[364,640,1825,1851]" pageId="209" pageNumber="210" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000</bibRefCitation>
) comb. nov. from the early Maastrichtian of Rügen, Germany.
<emphasis id="23A5FA67658485AAD18D226DFF2B9826" bold="true" box="[189,211,1860,1886]" pageId="209" pageNumber="210">6.</emphasis>
GZG.INV.78800, proximal LAP.
<emphasis id="23A5FA67658485AAD35A226DFD789826" bold="true" box="[618,640,1860,1886]" pageId="209" pageNumber="210">7.</emphasis>
GZG.INV.78801, median LAP.
<emphasis id="23A5FA67658485AAD532226DFBC99826" bold="true" box="[1026,1073,1860,1886]" pageId="209" pageNumber="210">8-9.</emphasis>
<taxonomicName id="D6D15DF6658485AAD50A226DFB0A9826" baseAuthorityName="Martynov" baseAuthorityYear="2010" box="[1082,1266,1860,1886]" class="Ophiuroidea" family="Ophiacanthidae" kingdom="Animalia" order="Ophiurida" pageId="209" pageNumber="210" phylum="Echinodermata" rank="family">Ophiacanthidae</taxonomicName>
gen. et sp. nov. innom. from the late Oxfordian (Late Jurassic) of the Plettenberg, Germany.
<emphasis id="23A5FA67658485AAD5BC224EFB5998F9" bold="true" box="[1164,1185,1895,1921]" pageId="209" pageNumber="210">8.</emphasis>
GZG.INV.78802, proximal LAP.
<emphasis id="23A5FA67658485AAD04122A3FE7F98DC" bold="true" box="[369,391,1930,1956]" pageId="209" pageNumber="210">9.</emphasis>
GZG.INV.78803, median or distal LAP.
<emphasis id="23A5FA67658485AAD25922A3FC4B98DC" bold="true" box="[873,947,1930,1956]" pageId="209" pageNumber="210">10-11.</emphasis>
<taxonomicName id="D6D15DF6658485AAD28B22A3FA8E98DD" authority="(Hess, 1966)" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Hess" baseAuthorityYear="1966" box="[955,1398,1930,1957]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiojagtus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="209" pageNumber="210" phylum="Echinodermata" rank="species" species="argoviensis" status="comb. nov.">
<emphasis id="23A5FA67658485AAD28B22A3FB2198DC" box="[955,1241,1930,1956]" italics="true" pageId="209" pageNumber="210">Ophiojagtus argoviensis</emphasis>
(
<bibRefCitation id="75405B84658485AAD5D922A3FA9698DD" author="Hess H." box="[1257,1390,1930,1957]" pageId="209" pageNumber="210" pagination="1025 - 1063" refId="ref122420" refString="Hess H. 1966. Mikropalaontologische Untersuchungen an Ophiuren V: Die Ophiuren aus dem Argovien (unteres Ober-Oxford) vom Guldental (Kt. Solothurn) und von Savigna (Dept. Jura). Eclogae geologicae Helvetiae 59: 1025 - 1063." type="journal article" year="1966">Hess, 1966</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="3896471C658485AAD18D2284FEB998B0" box="[189,321,1965,1992]" pageId="209" pageNumber="210" rank="species">comb. nov.</taxonomicNameLabel>
from the late Oxfordian (Late Jurassic) of Savigna, France.
<emphasis id="23A5FA67658485AAD5292284FBC598BF" bold="true" box="[1049,1085,1965,1991]" pageId="209" pageNumber="210">10.</emphasis>
NHMB M1124, proximal LAP.
<emphasis id="23A5FA67658485AAD03022F8FEDB9893" bold="true" box="[256,291,2001,2027]" pageId="209" pageNumber="210">11.</emphasis>
NHMB M11225, proximal to median LAP. One common scale bar per species.
</paragraph>
</caption>
<paragraph id="116E2675658785A9D18D271DFB3B9C7F" blockId="210.[189,1399,267,775]" pageId="210" pageNumber="211">Inner side of LAPs with relatively small, short, well-defined, prominent ridge with ventro-proximalwards bent ventral part not merged with ventral portion of LAP; dorsal tip of ridge slightly widened, with short ventro-proximalwards projecting extension; small, round knob distally bordering ridge in distal LAPs; inner side of distal edge of proximal to median LAPs with relatively small, moderately well-defined, round, very weakly protruding area of more densely meshed stereom; inner side of tentacle notch very small, shallow, moderately well defined laterally. No perforations or furrow discernible.</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658785A9D18D2619FD389AE7" pageId="210" pageNumber="211" type="discussion">
<paragraph id="116E2675658785A9D18D2619FED79C32" blockId="210.[189,303,816,842]" box="[189,303,816,842]" pageId="210" pageNumber="211">
<heading id="4A269119658785A9D18D2619FED79C32" bold="true" box="[189,303,816,842]" fontSize="11" level="3" pageId="210" pageNumber="211" reason="3">
<emphasis id="23A5FA67658785A9D18D2619FED79C32" bold="true" box="[189,303,816,842]" pageId="210" pageNumber="211">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658785A9D18D2649FA8F9BDB" blockId="210.[189,1399,864,1187]" pageId="210" pageNumber="211">
<bibRefCitation id="75405B84658785A9D18D2649FE379C02" author="Kutscher M. &amp; Jagt J. W. M." box="[189,463,864,891]" pageId="210" pageNumber="211" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt (2000)</bibRefCitation>
described this LAP
<typeStatus id="CE6A98D7658785A9D38A2648FD159C03" box="[698,749,865,891]" pageId="210" pageNumber="211">type</typeStatus>
from the early Maastrichtian of
<collectingCountry id="69C666E5658785A9D5552649FB2A9C02" box="[1125,1234,864,890]" name="Germany" pageId="210" pageNumber="211">Germany</collectingCountry>
and
<collectingCountry id="69C666E5658785A9D4392649FA8E9C02" box="[1289,1398,864,890]" name="Denmark" pageId="210" pageNumber="211">Denmark</collectingCountry>
in detail and tentatively assigned it to
<taxonomicName id="D6D15DF6658785A9D3B126ACFCE29CE7" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[641,794,901,927]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="210" pageNumber="211" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658785A9D3B126ACFCE29CE7" box="[641,794,901,927]" italics="true" pageId="210" pageNumber="211">Ophiacantha</emphasis>
</taxonomicName>
using open nomenclature. They stressed, however, that this assignment was highly doubtful when the fossil LAPs were compared with those of extant species of
<taxonomicName id="D6D15DF6658785A9D07326E6FE249C91" authorityName="Muller &amp; Troschel" authorityYear="1842" box="[323,476,975,1001]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiacantha" kingdom="Animalia" order="Ophiurida" pageId="210" pageNumber="211" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658785A9D07326E6FE249C91" box="[323,476,975,1001]" italics="true" pageId="210" pageNumber="211">Ophiacantha</emphasis>
</taxonomicName>
. Thanks to a systematic SEM-supported assessment of LAP morphologies in Recent ophiacanthids performed in the course of the present study, it is now possible to constrain the generic placement of these LAPs even further. Strongest similarities are shared with the LAPs of
<taxonomicName id="D6D15DF6658785A9D18D2117FE8C9B20" authorityName="Lyman" authorityYear="1869" box="[189,372,1086,1112]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="210" pageNumber="211" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658785A9D18D2117FE8C9B20" box="[189,372,1086,1112]" italics="true" pageId="210" pageNumber="211">Ophiochondrus</emphasis>
</taxonomicName>
, mainly on account of the massive, thick, rounded general aspect of the LAPs, the shape, size and position of the spine articulations, the outer surface ornament, and the shape of the ridge on the inner side. It is the first fossil record of
<taxonomicName id="D6D15DF6658785A9D3B021A1FCCF9BDA" authorityName="Lyman" authorityYear="1869" box="[640,823,1160,1186]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="210" pageNumber="211" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658785A9D3B021A1FCCF9BDA" box="[640,823,1160,1186]" italics="true" pageId="210" pageNumber="211">Ophiochondrus</emphasis>
</taxonomicName>
or at least of a very closely related, extinct genus.
</paragraph>
<paragraph id="116E2675658785A9D18D21E5FD389AE7" blockId="210.[189,1399,1228,1439]" pageId="210" pageNumber="211">
The LAPs of
<taxonomicName id="D6D15DF6658785A9D06721E5FD7D9B9E" authorityName="Thuy" authorityYear="2013" baseAuthorityName="Kutscher &amp; Jagt" baseAuthorityYear="2000" box="[343,645,1228,1254]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="210" pageNumber="211" phylum="Echinodermata" rank="species" species="punctatus" status="comb. nov.">
<emphasis id="23A5FA67658785A9D06721E5FD7D9B9E" box="[343,645,1228,1254]" italics="true" pageId="210" pageNumber="211">Ophiochondrus punctatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3896471C658785A9D3BA21E5FCF29B9E" box="[650,778,1228,1254]" pageId="210" pageNumber="211" rank="species">comb. nov.</taxonomicNameLabel>
cannot be confused with any other fossil
<typeStatus id="CE6A98D7658785A9D5D521E5FAE09B9E" box="[1253,1304,1228,1254]" pageId="210" pageNumber="211">type</typeStatus>
of LAP. Those of co-occurring
<taxonomicName id="D6D15DF6658785A9D0FD21D8FD7C9A73" authorityName="Lyman" authorityYear="1869" box="[461,644,1265,1291]" class="Ophiuroidea" family="Ophiacanthidae" genus="Ophiochondrus" kingdom="Animalia" order="Ophiurida" pageId="210" pageNumber="211" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658785A9D0FD21D8FD7C9A73" box="[461,644,1265,1291]" italics="true" pageId="210" pageNumber="211">Ophiochondrus</emphasis>
</taxonomicName>
?
<emphasis id="23A5FA67658785A9D3AA21D8FCCF9A73" box="[666,823,1265,1291]" italics="true" pageId="210" pageNumber="211">semirotundus</emphasis>
(
<bibRefCitation id="75405B84658785A9D27721D8FBAF9A73" author="Kutscher M. &amp; Jagt J. W. M." box="[839,1111,1265,1291]" pageId="210" pageNumber="211" pagination="45 - 107" refId="ref124720" refString="Kutscher M. &amp; Jagt J. W. M. 2000. Early Maastrichtian ophiuroids from Rugen (northeast Germany) and Mon (Denmark). In: Jagt J. W. M. 2000. Late Cretaceous-Early Palaeogene echinoderms and the K / T boundary in the southeast Netherlands and the northeast Belgium - Part 3: Ophiuroids. Scripta Geologica 121: 45 - 107." type="journal article" year="2000">Kutscher &amp; Jagt, 2000</bibRefCitation>
) comb. nov. have fewer spine articulations not bordered proximally by a ridge, and large contact surfaces with the opposite LAP on the inner side. LAPs assigned to
<taxonomicName id="D6D15DF6658785A9D3422012FCCC9A2D" authorityName="Thuy" authorityYear="2011" box="[626,820,1339,1365]" class="Ophiuroidea" family="Ophiacanthidae" genus="Inexpectacantha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="210" pageNumber="211" phylum="Echinodermata" rank="genus">
<emphasis id="23A5FA67658785A9D3422012FCCC9A2D" box="[626,820,1339,1365]" italics="true" pageId="210" pageNumber="211">Inexpectacantha</emphasis>
</taxonomicName>
display strongly tilted spine articulations with a ventral lobe commonly merged with the outer surface stereom, and a ridge on the inner side lacking the ventro-proximalwards projecting extension.
</paragraph>
</subSubSection>
<subSubSection id="59CB75FE658785A9D18D20E1FD10996A" box="[189,744,1480,1555]" pageId="210" pageNumber="211" type="distribution">
<paragraph id="116E2675658785A9D18D20E1FEB79A9A" blockId="210.[189,335,1480,1506]" box="[189,335,1480,1506]" pageId="210" pageNumber="211">
<heading id="4A269119658785A9D18D20E1FEB79A9A" bold="true" box="[189,335,1480,1506]" fontSize="11" level="3" pageId="210" pageNumber="211" reason="3">
<emphasis id="23A5FA67658785A9D18D20E1FEB79A9A" bold="true" box="[189,335,1480,1506]" pageId="210" pageNumber="211">Occurrence</emphasis>
</heading>
</paragraph>
<paragraph id="116E2675658785A9D18D20D1FD10996A" blockId="210.[189,744,1528,1555]" box="[189,744,1528,1555]" pageId="210" pageNumber="211">
Early Maastrichtian of
<collectingCountry id="69C666E5658785A9D0FC20D0FDC1996B" box="[460,569,1529,1555]" name="Germany" pageId="210" pageNumber="211">Germany</collectingCountry>
and
<collectingCountry id="69C666E5658785A9D34220D1FD1B996A" box="[626,739,1528,1554]" name="Denmark" pageId="210" pageNumber="211">Denmark</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>