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<document ID-DOI="10.11865/zs.2021404" ID-ISSN="2095-6827" ID-Zenodo-Dep="7176018" ID-ZooBank="511CFFE2-ED49-4C53-9E66-C431C944ED88" approvalRequired="7" approvalRequired_for_textStreams="7" checkinTime="1665295219367" checkinUser="felipe" docAuthor="Xu, Yuan, Xiong, Yun &amp; Liang, Hongbin" docDate="2021" docId="857C87D1171CFF8634A21C642D80FDCE" docLanguage="en" docName="ZoolSyst.46.4.300316.pdf" docOrigin="Zoological Systematics 46 (4)" docSource="http://zoobank.org/511cffe2-ed49-4c53-9e66-c431c944ed88" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Lilioceris fouana Pic 1932" docType="treatment" docVersion="3" lastPageNumber="307" masterDocId="7945FFA91719FF81342A1E292C07FF91" masterDocTitle="A review of the neptis species group of the genus Lilioceris (Coleoptera: Chrysomelidae: Criocerinae)" masterLastPageNumber="316" masterPageNumber="300" pageNumber="305" updateTime="1665314060196" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A review of the neptis species group of the genus Lilioceris (Coleoptera: Chrysomelidae: Criocerinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Yuan</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; College of Life Science, University of Chinese Academy of Sciences, Beijing 100049, China Corresponding author, E-mail: lianghb @ ioz. ac. cn</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xiong, Yun</mods:namePart>
<mods:affiliation>Gongshan Protection Sub-Bureau, Gaoligongshan National Nature Reserve, Gongshan 673500, Yunnan, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:title>Zoological Systematics</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="volume">
<mods:number>46</mods:number>
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<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:identifier type="DOI">10.11865/zs.2021404</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7172322" ID-Zenodo-Dep="7172322" LSID="urn:lsid:plazi:treatment:857C87D1171CFF8634A21C642D80FDCE" httpUri="http://treatment.plazi.org/id/857C87D1171CFF8634A21C642D80FDCE" lastPageId="7" lastPageNumber="307" pageId="5" pageNumber="305">
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<paragraph blockId="5.[136,691,589,678]" box="[136,691,589,614]" pageId="5" pageNumber="305">
<heading box="[136,691,589,614]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="305" reason="2">
<taxonomicName authority="Pic, 1932" authorityName="Pic" authorityYear="1932" box="[136,435,589,614]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="305" phylum="Arthropoda" rank="species" species="fouana">
<emphasis bold="true" box="[136,435,589,614]" pageId="5" pageNumber="305">
<emphasis bold="true" box="[136,326,589,613]" italics="true" pageId="5" pageNumber="305">Lilioceris fouana</emphasis>
<bibRefCitation author="Pic" box="[332,435,590,614]" pageId="5" pageNumber="305" pagination="1036" refId="ref11391" refString="Pic, M. 1932. Nouveautes diverses. Melanges Exotico-Entomologiques, 59: 1036." type="journal article" year="1932">Pic, 1932</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation box="[468,564,590,614]" captionStart="Figures 710" captionStartId="6.[155,227,2006,2028]" captionTargetBox="[220,1371,482,1970]" captionTargetId="figure-143@6.[189,1399,479,1976]" captionTargetPageId="6" captionText="Figures 710. Habitus of Lilioceris spp. 78. L. fouana, ♀, syntype, China (Yunnan), photoed by Christophe Rivier. 910. L. neptis, ♂, China (Zhejiang: Tianmu Shan). 7, 9. Dorsal view. 8, 10. Ventral view. Scale bars=5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7176024" httpUri="https://zenodo.org/record/7176024/files/figure.png" pageId="5" pageNumber="305">Figs 78</figureCitation>
,
<figureCitation box="[575,604,590,614]" captionStart="Figures 1519" captionStartId="10.[155,227,1993,2015]" captionTargetBox="[189,1399,260,1960]" captionTargetId="figure-68@10.[189,1399,260,1960]" captionTargetPageId="10" captionText="Figures 1519. Pronotum, mesoventral process and metaventrite of Lilioceris spp. 15. L. vietnamica, ♂, China (Yunnan: Menghai). 16. L. cantonensis, ♂, China (Zhejiang: Tianmu Shan). 17. L. fouana, ♂, China (Sichuan: Luding). 18. L. neptis, ♀, China (Hong Kong: Honghualing). 19. L. subpolita, ♂, Japan (Osaka: Minomo). A. Pronotum. B. Mesoventral process. C. Metaventral disc." figureDoi="http://doi.org/10.5281/zenodo.7176030" httpUri="https://zenodo.org/record/7176030/files/figure.png" pageId="5" pageNumber="305">17</figureCitation>
,
<figureCitation box="[615,683,590,614]" captionStart-0="Figures 2024" captionStart-1="Figures 2529" captionStartId-0="11.[155,227,1722,1744]" captionStartId-1="12.[155,227,1989,2011]" captionTargetBox-0="[208,1384,248,1673]" captionTargetBox-1="[211,1385,890,1940]" captionTargetId-0="figure-173@11.[189,1399,245,1683]" captionTargetId-1="figure-315@12.[189,1398,888,1947]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="Figures 2024. Male genitalia of Lilioceris spp. 20. L. cantonensis, China (Zhejiang: Tianmu Shan). 21. L. neptis, China (Fujian: Dehua). 22. L. vietnamica, China (Yunnan: Menghai). 23. L. subpolita, Japan (Osaka: Minomo). 24. L. fouana, China (Sichuan: Luding).A. Aedeagus, lateral view. B. Aedeagus, dorsal view. C. Dorsal sclerite, dorsal view. D. Sclerites in internal sac, lateral view. Scale bars: AB=0.5 mm; CD=0.2mm." captionText-1="Figures 2529. Female reproductive organs of Lilioceris spp. 25. L. fouana, China (Sichuan: Luding). 26. L. vietnamica, China (Yunnan: Jinghong). 27. L. cantonensis, China (Zhejiang: Tianmu Shan). 28. Lilioceris neptis, China (Guangxi: Guilin). 29. L. subpolita, Japan (Mont Takao). A. Dorsal view. B. Ventral view. C. Spermatheca. Scale bars: AB=0.5mm; C=0.1mm." figureDoi-0="http://doi.org/10.5281/zenodo.7176032" figureDoi-1="http://doi.org/10.5281/zenodo.7176034" httpUri-0="https://zenodo.org/record/7176032/files/figure.png" httpUri-1="https://zenodo.org/record/7176034/files/figure.png" pageId="5" pageNumber="305">2425</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="305" type="reference_group">
<paragraph blockId="5.[136,691,589,678]" box="[136,613,624,646]" pageId="5" pageNumber="305">
<treatmentCitationGroup box="[136,613,624,646]" pageId="5" pageNumber="305">
<treatmentCitation author="Pic" box="[136,441,624,646]" page="11" pageId="5" pageNumber="305" year="1932">
<taxonomicName authority="Pic, 1932: 11" authorityName="Pic" authorityPageNumber="11" authorityYear="1932" box="[136,441,624,646]" class="Insecta" family="Chrysomelidae" genus="Crioceris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="305" phylum="Arthropoda" rank="species" species="fouanus">
<emphasis box="[136,309,624,646]" italics="true" pageId="5" pageNumber="305">Crioceris fouanus</emphasis>
<bibRefCitation author="Pic" box="[315,441,624,646]" pageId="5" pageNumber="305" pagination="1036" refId="ref11391" refString="Pic, M. 1932. Nouveautes diverses. Melanges Exotico-Entomologiques, 59: 1036." type="journal article" year="1932">Pic, 1932: 11</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<collectingCountry box="[454,516,624,646]" name="China" pageId="5" pageNumber="305">China</collectingCountry>
:
<collectingRegion box="[524,602,624,645]" country="China" name="Yunnan" pageId="5" pageNumber="305">Yunnan</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="5.[136,691,589,678]" box="[136,596,656,678]" pageId="5" pageNumber="305">
<treatmentCitationGroup box="[136,596,656,678]" pageId="5" pageNumber="305">
<treatmentCitation author="Gressitt &amp; Kimoto &amp; The Chrysomelidae &amp; Coleopt &amp; China and Korea &amp; Pacific Insects Monographs" box="[136,592,656,678]" page="48" pageId="5" pageNumber="305" year="1961">
<taxonomicName authority=": Gressitt &amp; Kimoto, 1961: 48" authorityName=": Gressitt &amp; Kimoto" authorityPageNumber="48" authorityYear="1961" box="[136,592,656,678]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="305" phylum="Arthropoda" rank="species" species="fouana">
<emphasis box="[136,301,656,677]" italics="true" pageId="5" pageNumber="305">Lilioceris fouana</emphasis>
:
<bibRefCitation author="Gressitt &amp; Kimoto &amp; The Chrysomelidae &amp; Coleopt &amp; China and Korea &amp; Pacific Insects Monographs" box="[313,592,656,678]" pageId="5" pageNumber="305" pagination="1 - 299" refId="ref10792" refString="Gressitt, J. L., Kimoto, S. 1961. The Chrysomelidae (Coleopt.) of China and Korea, part 1. Pacific Insects Monographs, 1 A: 1 - 299." type="journal article" year="1961">Gressitt &amp; Kimoto, 1961: 48</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="307" pageId="5" pageNumber="305" type="materials_examined">
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">
Material examined. Total
<specimenCount box="[471,621,709,734]" pageId="5" pageNumber="305" type="generic">18 specimens</specimenCount>
.
<collectingCountry box="[633,701,709,733]" name="China" pageId="5" pageNumber="305">China</collectingCountry>
:
<collectingRegion box="[713,785,710,734]" country="China" name="Gansu" pageId="5" pageNumber="305">Gansu</collectingRegion>
:
<specimenCount box="[798,832,710,734]" pageId="5" pageNumber="305" type="male">1♂</specimenCount>
, Kangxian, Qinghe Forest Farm /
<date box="[1210,1343,710,734]" pageId="5" pageNumber="305" value="1998-07-14">1998.VII.14</date>
, Decheng Yuan coll.;
<specimenCount box="[271,304,744,768]" pageId="5" pageNumber="305" type="female">1♀</specimenCount>
, Wenxian /
<date box="[451,584,744,768]" pageId="5" pageNumber="305" value="1988-07-20">1988.VII.20</date>
, Hongjian Wang coll.;
<collectingRegion box="[855,945,744,768]" country="China" name="Shaanxi" pageId="5" pageNumber="305">Shaanxi</collectingRegion>
:
<specimenCount box="[961,995,744,768]" pageId="5" pageNumber="305" type="male">1♂</specimenCount>
(
<collectionCode box="[1015,1092,744,768]" country="USA" httpUri="http://grbio.org/cool/3dng-x47a" name="Carthage College" pageId="5" pageNumber="305">CCCC</collectionCode>
), Huxian, Huashuping,
<quantity box="[1363,1447,744,768]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="305" unit="m" value="1800.0">1800 m</quantity>
,
<date box="[136,258,779,803]" pageId="5" pageNumber="305" value="2020-06-19">2020.VI.19</date>
J.T. Zhao coll;
<specimenCount box="[422,455,779,803]" pageId="5" pageNumber="305" type="female">1♀</specimenCount>
, Hanzhong, Liping township /
<date box="[785,896,779,803]" pageId="5" pageNumber="305" value="1958-05-16">1958.V.16</date>
, Shimei Song coll.;
<collectingRegion box="[1109,1178,779,803]" country="China" name="Hubei" pageId="5" pageNumber="305">Hubei</collectingRegion>
:
<specimenCount box="[1188,1223,779,803]" pageId="5" pageNumber="305" type="male">1♂</specimenCount>
, Xingshan, Longmen He /
<date box="[193,317,814,838]" pageId="5" pageNumber="305" value="1993-06-23">1993.VI.23</date>
, Wenzhu Li coll.;
<specimenCount box="[523,556,814,838]" pageId="5" pageNumber="305" type="female">1♀</specimenCount>
, Xingshan, Longmen He /
<date box="[860,971,814,838]" pageId="5" pageNumber="305" value="1994-05-08">1994.V.08</date>
, Youwei Zhang coll.;
<collectingRegion box="[1216,1306,813,837]" country="China" name="Sichuan" pageId="5" pageNumber="305">Sichuan</collectingRegion>
:
<specimenCount box="[1318,1353,814,838]" pageId="5" pageNumber="305" type="male">1♂</specimenCount>
, Luding, Xinxing township /
<quantity box="[351,431,848,872]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="305" unit="m" value="1800.0">1800 m</quantity>
,
<date box="[442,566,848,872]" pageId="5" pageNumber="305" value="1983-06-13">1983.VI.13</date>
, Shuyong Wang coll.;
<specimenCount box="[810,845,848,872]" pageId="5" pageNumber="305" type="male">1♂</specimenCount>
, Emei Shan /
<date box="[996,1120,848,872]" pageId="5" pageNumber="305" value="1955-06-26">1955.VI.26</date>
, Xingchi Yang coll.;
<specimenCount box="[1348,1383,848,872]" pageId="5" pageNumber="305" type="male">1♂</specimenCount>
, Emei Shan,
<quantity box="[207,355,883,907]" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="2.1" metricValueMin="1.1" pageId="5" pageNumber="305" unit="m" value="1600.0" valueMax="2100.0" valueMin="1100.0">11002100 m</quantity>
,
<date box="[370,493,883,907]" pageId="5" pageNumber="305" value="1955-06-20">1955.VI.20</date>
, Yunzhen Zi coll.;
<specimenCount box="[711,744,883,907]" pageId="5" pageNumber="305" type="female">1♀</specimenCount>
, Luding, Xinxing township /
<date box="[1081,1204,883,907]" pageId="5" pageNumber="305" value="1983-06-13">1983.VI.13</date>
, Shuyong Wang coll.;
<collectingRegion box="[136,222,918,941]" country="China" name="Yunnan" pageId="5" pageNumber="305">Yunnan</collectingRegion>
:
<specimenCount box="[233,268,918,942]" pageId="5" pageNumber="305" type="male">1♂</specimenCount>
, Gongshan, Bingzhongluo village /
<date box="[656,777,917,941]" pageId="5" pageNumber="305" value="2002-04-26">2002.IV.26</date>
, Hongbin Liang &amp; Weidong Ba coll.;
<specimenCount box="[1188,1221,918,942]" pageId="5" pageNumber="305" type="female">1♀</specimenCount>
, Lijiang, Yulongshan Nature Reserve /
<date box="[333,457,952,976]" pageId="5" pageNumber="305" value="2005-06-27">2005.VI.27</date>
, Wenhua Lu coll.,
<specimenCount box="[671,704,952,976]" pageId="5" pageNumber="305" type="male">1♂</specimenCount>
(
<collectionCode box="[722,789,952,976]" pageId="5" pageNumber="305">MHU</collectionCode>
), Lushui, Pianma township /
<date box="[1119,1257,952,976]" pageId="5" pageNumber="305" value="2004-05-09" valueMax="2004-05-11" valueMin="2004-05-09">2004.V.911</date>
, Xiujuan Yang &amp; Yushuang Liu coll.;
<specimenCount box="[352,388,987,1011]" pageId="5" pageNumber="305" type="male">2♂</specimenCount>
, Wuding, Chadian, Changji Road,
<geoCoordinate box="[759,881,987,1011]" degrees="25.74144" direction="north" orientation="latitude" pageId="5" pageNumber="305" precision="1" value="25.74144">N25.74144</geoCoordinate>
,
<geoCoordinate box="[892,1021,987,1011]" degrees="102.30336" direction="east" orientation="longitude" pageId="5" pageNumber="305" precision="1" value="102.30336">E102.30336</geoCoordinate>
/
<quantity box="[1041,1121,987,1011]" metricMagnitude="3" metricUnit="m" metricValue="2.296" pageId="5" pageNumber="305" unit="m" value="2296.0">2296 m</quantity>
,
<date box="[1132,1262,987,1011]" pageId="5" pageNumber="305" value="2020-07-11">2020.VII.11</date>
D1, Yuan Xu and Neng Zhang coll;
<specimenCount box="[332,365,1022,1046]" pageId="5" pageNumber="305" type="female">1♀</specimenCount>
(
<collectionCode box="[380,447,1022,1046]" pageId="5" pageNumber="305">MHU</collectionCode>
), Zhenyuan, Jiujia township /
<date box="[773,928,1022,1046]" pageId="5" pageNumber="305" value="2009-08-01" valueMax="2009-08-03" valueMin="2009-08-01">2009.VIII.13</date>
, Jishan Xu &amp; Jianxiong Zhang coll;
<specimenCount box="[1324,1357,1022,1046]" pageId="5" pageNumber="305" type="female">1♀</specimenCount>
(
<collectionCode box="[1372,1439,1022,1046]" pageId="5" pageNumber="305">MHU</collectionCode>
), Lushui, Pianma,
<date box="[320,458,1056,1080]" pageId="5" pageNumber="305" value="2004-05-09" valueMax="2004-05-11" valueMin="2004-05-09">2004.V.911</date>
, Xiujuan Yang, Yushuang Liu coll.;
<specimenCount box="[852,886,1056,1080]" pageId="5" pageNumber="305" type="female">1♀</specimenCount>
(
<collectionCode box="[901,988,1056,1080]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="305">MNHN</collectionCode>
),
<collectingRegion box="[1007,1088,1057,1080]" country="China" name="Yunnan" pageId="5" pageNumber="305">Yunnan</collectingRegion>
fou / Type /
<taxonomicName authorityName=": Gressitt &amp; Kimoto" authorityYear="1961" box="[1232,1306,1056,1080]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="305" phylum="Arthropoda" rank="species" species="fouana">
<emphasis box="[1232,1306,1056,1080]" italics="true" pageId="5" pageNumber="305">fouana</emphasis>
</taxonomicName>
n sp / Type /
<typeStatus box="[136,256,1091,1115]" pageId="5" pageNumber="305">SYNTYPE</typeStatus>
/
<typeStatus box="[277,364,1091,1115]" pageId="5" pageNumber="305">Syntype</typeStatus>
<taxonomicName authority="(Pic, 1932)" baseAuthorityName="Pic" baseAuthorityYear="1932" box="[371,690,1091,1115]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="305" phylum="Arthropoda" rank="species" species="fouanus">
<emphasis box="[371,565,1091,1115]" italics="true" pageId="5" pageNumber="305">Lilioceris fouanus</emphasis>
(
<bibRefCitation author="Pic" box="[579,682,1091,1115]" pageId="5" pageNumber="305" pagination="1036" refId="ref11391" refString="Pic, M. 1932. Nouveautes diverses. Melanges Exotico-Entomologiques, 59: 1036." type="journal article" year="1932">Pic, 1932</bibRefCitation>
)
</taxonomicName>
/ MNHN Paris EC11741.
</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">Diagnosis. Femora and ventral surface of body black; antennomeres 510 twice as long as wide; pronotal disc a short longitudinal row of 35 small punctures; elytral punctures sparse throughout, large on basal half, slightly diminishing posteriorly, but not absent near apex; apex of mesoventral process strongly widened.</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">
Redescription. BL
<quantity box="[399,528,1229,1253]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="5" pageNumber="305" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.56.5 mm</quantity>
, BW 2.53.0 mm. Head, antennae, legs and ventral side of body black, pronotum, scutellum and elytra brownish red.
</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">
Head (
<figureCitation box="[259,355,1299,1323]" captionStart="Figures 710" captionStartId="6.[155,227,2006,2028]" captionTargetBox="[220,1371,482,1970]" captionTargetId="figure-143@6.[189,1399,479,1976]" captionTargetPageId="6" captionText="Figures 710. Habitus of Lilioceris spp. 78. L. fouana, ♀, syntype, China (Yunnan), photoed by Christophe Rivier. 910. L. neptis, ♂, China (Zhejiang: Tianmu Shan). 7, 9. Dorsal view. 8, 10. Ventral view. Scale bars=5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7176024" httpUri="https://zenodo.org/record/7176024/files/figure.png" pageId="5" pageNumber="305">Figs 78</figureCitation>
). HL/HW 1.31.6; vertex with a deep groove in middle, punctate laterally; occipit with a shallow furrow medially, sparsely punctate; gena punctate and setose; frontal tubercle glabrous, flat; frontoclypeal area triangular, disc with punctures and setae; labrum transverse, with long setae; antennae filiform, shorter than half of body length, antennomeres 14 nearly globular, 2 shortest, 510 cylindrical, twice as long as wide; antennomeres 1 and 2 sparsely pubescent and punctate, 311 densely pubescent and punctate.
</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">
Pronotum (
<figureCitation box="[310,380,1472,1496]" captionStart="Figures 710" captionStartId="6.[155,227,2006,2028]" captionTargetBox="[220,1371,482,1970]" captionTargetId="figure-143@6.[189,1399,479,1976]" captionTargetPageId="6" captionText="Figures 710. Habitus of Lilioceris spp. 78. L. fouana, ♀, syntype, China (Yunnan), photoed by Christophe Rivier. 910. L. neptis, ♂, China (Zhejiang: Tianmu Shan). 7, 9. Dorsal view. 8, 10. Ventral view. Scale bars=5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7176024" httpUri="https://zenodo.org/record/7176024/files/figure.png" pageId="5" pageNumber="305">Figs 7</figureCitation>
,
<figureCitation box="[393,442,1472,1496]" captionStart="Figures 1519" captionStartId="10.[155,227,1993,2015]" captionTargetBox="[189,1399,260,1960]" captionTargetId="figure-68@10.[189,1399,260,1960]" captionTargetPageId="10" captionText="Figures 1519. Pronotum, mesoventral process and metaventrite of Lilioceris spp. 15. L. vietnamica, ♂, China (Yunnan: Menghai). 16. L. cantonensis, ♂, China (Zhejiang: Tianmu Shan). 17. L. fouana, ♂, China (Sichuan: Luding). 18. L. neptis, ♀, China (Hong Kong: Honghualing). 19. L. subpolita, ♂, Japan (Osaka: Minomo). A. Pronotum. B. Mesoventral process. C. Metaventral disc." figureDoi="http://doi.org/10.5281/zenodo.7176030" httpUri="https://zenodo.org/record/7176030/files/figure.png" pageId="5" pageNumber="305">17A</figureCitation>
). PW/HW 1.72.0, PL/PW 0.80.9; anterior angle protruding, posterior angle not protruding; sides constricted in middle; middle of disc with a longitudinal row of 35 small punctures; posterior transverse impression indistinct, anterior transverse impression and basal transverse groove absent. Scutellum triangular and pubescent.
</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">
Elytra (
<figureCitation box="[270,336,1576,1600]" captionStart="Figures 710" captionStartId="6.[155,227,2006,2028]" captionTargetBox="[220,1371,482,1970]" captionTargetId="figure-143@6.[189,1399,479,1976]" captionTargetPageId="6" captionText="Figures 710. Habitus of Lilioceris spp. 78. L. fouana, ♀, syntype, China (Yunnan), photoed by Christophe Rivier. 910. L. neptis, ♂, China (Zhejiang: Tianmu Shan). 7, 9. Dorsal view. 8, 10. Ventral view. Scale bars=5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7176024" httpUri="https://zenodo.org/record/7176024/files/figure.png" pageId="5" pageNumber="305">Fig. 7</figureCitation>
). EL/EW 2.02.3; sutural angle rounded; humeri protruding, humeral groove distinct, basal impression indistinct; intervals smooth; strial punctures sparse and large on basal half, diminishing posteriorly, but not absent, interspace between two neighboring punctures in a stria larger than that in
<taxonomicName baseAuthorityName="Weise" baseAuthorityYear="1922" box="[834,927,1646,1669]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="305" phylum="Arthropoda" rank="species" species="neptis">
<emphasis box="[834,927,1646,1669]" italics="true" pageId="5" pageNumber="305">L. neptis</emphasis>
</taxonomicName>
; scutellary striole with 46 punctures; epipleura with upper margin slightly raised, with a row of fine punctures laterally.
</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">
Mesoventrite pubescent, apical portion of mesoventral process strongly widened, convex, tuberculate, horizontally connected with metaventrite (
<figureCitation box="[460,560,1749,1773]" captionStart="Figures 1519" captionStartId="10.[155,227,1993,2015]" captionTargetBox="[189,1399,260,1960]" captionTargetId="figure-68@10.[189,1399,260,1960]" captionTargetPageId="10" captionText="Figures 1519. Pronotum, mesoventral process and metaventrite of Lilioceris spp. 15. L. vietnamica, ♂, China (Yunnan: Menghai). 16. L. cantonensis, ♂, China (Zhejiang: Tianmu Shan). 17. L. fouana, ♂, China (Sichuan: Luding). 18. L. neptis, ♀, China (Hong Kong: Honghualing). 19. L. subpolita, ♂, Japan (Osaka: Minomo). A. Pronotum. B. Mesoventral process. C. Metaventral disc." figureDoi="http://doi.org/10.5281/zenodo.7176030" httpUri="https://zenodo.org/record/7176030/files/figure.png" pageId="5" pageNumber="305">Fig. 17B</figureCitation>
). metaventral disc with a triangular setose patch near anterior margin (
<figureCitation box="[1338,1438,1749,1773]" captionStart="Figures 1519" captionStartId="10.[155,227,1993,2015]" captionTargetBox="[189,1399,260,1960]" captionTargetId="figure-68@10.[189,1399,260,1960]" captionTargetPageId="10" captionText="Figures 1519. Pronotum, mesoventral process and metaventrite of Lilioceris spp. 15. L. vietnamica, ♂, China (Yunnan: Menghai). 16. L. cantonensis, ♂, China (Zhejiang: Tianmu Shan). 17. L. fouana, ♂, China (Sichuan: Luding). 18. L. neptis, ♀, China (Hong Kong: Honghualing). 19. L. subpolita, ♂, Japan (Osaka: Minomo). A. Pronotum. B. Mesoventral process. C. Metaventral disc." figureDoi="http://doi.org/10.5281/zenodo.7176030" httpUri="https://zenodo.org/record/7176030/files/figure.png" pageId="5" pageNumber="305">Fig. 17C</figureCitation>
); metepisternum densely pubescent.
</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">Abdominal sternite with sparse setae and punctures, transverse impressions distinct in sternites 25, with dense punctures and pubescence.</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" pageId="5" pageNumber="305">Legs slender; tibiae with dense punctures, pubescence and two spurs; femora with dense pubescence on dorsal surface, with sparse setae on ventral surface, with middle area widened.</paragraph>
<paragraph blockId="5.[136,1452,709,2051]" lastBlockId="6.[136,1452,236,434]" lastPageId="6" lastPageNumber="306" pageId="5" pageNumber="305">
Male genitalia (
<figureCitation box="[359,497,1957,1981]" captionStart="Figures 2024" captionStartId="11.[155,227,1722,1744]" captionTargetBox="[208,1384,248,1673]" captionTargetId="figure-173@11.[189,1399,245,1683]" captionTargetPageId="11" captionText="Figures 2024. Male genitalia of Lilioceris spp. 20. L. cantonensis, China (Zhejiang: Tianmu Shan). 21. L. neptis, China (Fujian: Dehua). 22. L. vietnamica, China (Yunnan: Menghai). 23. L. subpolita, Japan (Osaka: Minomo). 24. L. fouana, China (Sichuan: Luding).A. Aedeagus, lateral view. B. Aedeagus, dorsal view. C. Dorsal sclerite, dorsal view. D. Sclerites in internal sac, lateral view. Scale bars: AB=0.5 mm; CD=0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7176032" httpUri="https://zenodo.org/record/7176032/files/figure.png" pageId="5" pageNumber="305">Figs 24AD</figureCitation>
). Median foramen occupying 1/5 length of median lobe (
<figureCitation box="[1112,1212,1957,1981]" captionStart="Figures 2024" captionStartId="11.[155,227,1722,1744]" captionTargetBox="[208,1384,248,1673]" captionTargetId="figure-173@11.[189,1399,245,1683]" captionTargetPageId="11" captionText="Figures 2024. Male genitalia of Lilioceris spp. 20. L. cantonensis, China (Zhejiang: Tianmu Shan). 21. L. neptis, China (Fujian: Dehua). 22. L. vietnamica, China (Yunnan: Menghai). 23. L. subpolita, Japan (Osaka: Minomo). 24. L. fouana, China (Sichuan: Luding).A. Aedeagus, lateral view. B. Aedeagus, dorsal view. C. Dorsal sclerite, dorsal view. D. Sclerites in internal sac, lateral view. Scale bars: AB=0.5 mm; CD=0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7176032" httpUri="https://zenodo.org/record/7176032/files/figure.png" pageId="5" pageNumber="305">Fig. 24A</figureCitation>
); apex of median lobe round (
<figureCitation box="[215,314,1992,2016]" captionStart="Figures 2024" captionStartId="11.[155,227,1722,1744]" captionTargetBox="[208,1384,248,1673]" captionTargetId="figure-173@11.[189,1399,245,1683]" captionTargetPageId="11" captionText="Figures 2024. Male genitalia of Lilioceris spp. 20. L. cantonensis, China (Zhejiang: Tianmu Shan). 21. L. neptis, China (Fujian: Dehua). 22. L. vietnamica, China (Yunnan: Menghai). 23. L. subpolita, Japan (Osaka: Minomo). 24. L. fouana, China (Sichuan: Luding).A. Aedeagus, lateral view. B. Aedeagus, dorsal view. C. Dorsal sclerite, dorsal view. D. Sclerites in internal sac, lateral view. Scale bars: AB=0.5 mm; CD=0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7176032" httpUri="https://zenodo.org/record/7176032/files/figure.png" pageId="5" pageNumber="305">Fig. 24B</figureCitation>
); tegmen Y shaped and weak, basal piece of tegmen triangular and relatively broad, lateral lobes weakly scleritized, combined with second connecting membrane; internal sac membranous, with dorsal, median, and ventral sclerites weakly scleritized (
<figureCitation box="[344,479,236,261]" captionStart="Figures 2024" captionStartId="11.[155,227,1722,1744]" captionTargetBox="[208,1384,248,1673]" captionTargetId="figure-173@11.[189,1399,245,1683]" captionTargetPageId="11" captionText="Figures 2024. Male genitalia of Lilioceris spp. 20. L. cantonensis, China (Zhejiang: Tianmu Shan). 21. L. neptis, China (Fujian: Dehua). 22. L. vietnamica, China (Yunnan: Menghai). 23. L. subpolita, Japan (Osaka: Minomo). 24. L. fouana, China (Sichuan: Luding).A. Aedeagus, lateral view. B. Aedeagus, dorsal view. C. Dorsal sclerite, dorsal view. D. Sclerites in internal sac, lateral view. Scale bars: AB=0.5 mm; CD=0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7176032" httpUri="https://zenodo.org/record/7176032/files/figure.png" pageId="6" pageNumber="306">Figs 24CD</figureCitation>
), dorsal sclerite with straight extension (
<figureCitation box="[910,1008,236,261]" captionStart="Figures 2024" captionStartId="11.[155,227,1722,1744]" captionTargetBox="[208,1384,248,1673]" captionTargetId="figure-173@11.[189,1399,245,1683]" captionTargetPageId="11" captionText="Figures 2024. Male genitalia of Lilioceris spp. 20. L. cantonensis, China (Zhejiang: Tianmu Shan). 21. L. neptis, China (Fujian: Dehua). 22. L. vietnamica, China (Yunnan: Menghai). 23. L. subpolita, Japan (Osaka: Minomo). 24. L. fouana, China (Sichuan: Luding).A. Aedeagus, lateral view. B. Aedeagus, dorsal view. C. Dorsal sclerite, dorsal view. D. Sclerites in internal sac, lateral view. Scale bars: AB=0.5 mm; CD=0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7176032" httpUri="https://zenodo.org/record/7176032/files/figure.png" pageId="6" pageNumber="306">Fig. 24C</figureCitation>
).
</paragraph>
<paragraph blockId="6.[136,1452,236,434]" pageId="6" pageNumber="306">
Female reproductive organs (
<figureCitation box="[503,639,271,295]" captionStart="Figures 2529" captionStartId="12.[155,227,1989,2011]" captionTargetBox="[211,1385,890,1940]" captionTargetId="figure-315@12.[189,1398,888,1947]" captionTargetPageId="12" captionText="Figures 2529. Female reproductive organs of Lilioceris spp. 25. L. fouana, China (Sichuan: Luding). 26. L. vietnamica, China (Yunnan: Jinghong). 27. L. cantonensis, China (Zhejiang: Tianmu Shan). 28. Lilioceris neptis, China (Guangxi: Guilin). 29. L. subpolita, Japan (Mont Takao). A. Dorsal view. B. Ventral view. C. Spermatheca. Scale bars: AB=0.5mm; C=0.1mm." figureDoi="http://doi.org/10.5281/zenodo.7176034" httpUri="https://zenodo.org/record/7176034/files/figure.png" pageId="6" pageNumber="306">Figs 25AC</figureCitation>
). Tergites 8 and 9, sternites 8 and 9 of female sclerotized, posterior areas of tergite 8 and sternite 8 with setae, without apodemes, spiculum gastrale short, X shaped, strongly widened in distal part, apical margin truncate; vaginal palpi with dense setae, very short and cylindrical; spermatheca complicatedly folded.
</paragraph>
<paragraph blockId="6.[136,1452,236,434]" box="[189,966,375,399]" pageId="6" pageNumber="306">
Distribution (
<figureCitation box="[333,413,375,399]" captionStart="Figure 30" captionStartId="13.[155,218,1402,1424]" captionTargetBox="[188,1397,536,1367]" captionTargetId="graphics-520@13.[188,1397,536,1368]" captionTargetPageId="13" captionText="Figure 30. Distribution map of Lilioceris spp. in East Asia." figureDoi="http://doi.org/10.5281/zenodo.7176036" httpUri="https://zenodo.org/record/7176036/files/figure.png" pageId="6" pageNumber="306">Fig. 30</figureCitation>
).
<collectingCountry box="[432,496,375,399]" name="China" pageId="6" pageNumber="306">China</collectingCountry>
(
<collectingRegion box="[510,582,375,399]" country="China" name="Gansu" pageId="6" pageNumber="306">Gansu</collectingRegion>
,
<collectingRegion box="[593,682,375,399]" country="China" name="Shaanxi" pageId="6" pageNumber="306">Shaanxi</collectingRegion>
,
<collectingRegion box="[692,760,375,399]" country="China" name="Hubei" pageId="6" pageNumber="306">Hubei</collectingRegion>
,
<collectingRegion box="[770,859,375,399]" country="China" name="Sichuan" pageId="6" pageNumber="306">Sichuan</collectingRegion>
,
<collectingRegion box="[868,955,375,399]" country="China" name="Yunnan" pageId="6" pageNumber="306">Yunnan</collectingRegion>
).
</paragraph>
<paragraph blockId="6.[136,1452,236,434]" lastBlockId="7.[136,1452,236,607]" lastPageId="7" lastPageNumber="307" pageId="6" pageNumber="306">
Host plant and habitat. This species feeds on
<taxonomicName box="[683,832,410,433]" class="Magnoliopsida" family="Hernandiaceae" genus="Dioscorea" kingdom="Plantae" order="Laurales" pageId="6" pageNumber="306" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[683,793,410,433]" italics="true" pageId="6" pageNumber="306">Dioscorea</emphasis>
sp.
</taxonomicName>
(
<taxonomicName authorityName="R.Brown" authorityYear="1810" box="[847,1006,410,434]" class="Liliopsida" family="Dioscoreaceae" kingdom="Plantae" order="Dioscoreales" pageId="6" pageNumber="306" phylum="Tracheophyta" rank="family">Dioscoreaceae</taxonomicName>
) according to the observation of the first author (
<collectionCode box="[218,259,237,261]" pageId="7" pageNumber="307">XY</collectionCode>
) in
<collectingRegion box="[298,380,237,260]" country="China" name="Yunnan" pageId="7" pageNumber="307">Yunnan</collectingRegion>
(
<figureCitation box="[393,472,236,261]" captionStart="Figure 33" captionStartId="14.[155,218,2049,2071]" captionTargetBox="[497,1091,1566,2012]" captionTargetId="figure-340@14.[497,1091,1566,2012]" captionTargetPageId="14" captionText="Figure 33. Host plant of Lilioceris fouana: Dioscorea sp., Wuding, 2020.VII.11, photoed by Yuan Xu." figureDoi="http://doi.org/10.5281/zenodo.7176040" httpUri="https://zenodo.org/record/7176040/files/figure.png" pageId="7" pageNumber="307">Fig. 33</figureCitation>
). It is confined to west central
<collectingCountry box="[795,861,236,260]" name="China" pageId="7" pageNumber="307">China</collectingCountry>
, at higher elevation (
<quantity box="[1081,1228,236,261]" metricMagnitude="3" metricUnit="m" metricValue="2.048" metricValueMax="2.296" metricValueMin="1.8" pageId="7" pageNumber="307" unit="m" value="2048.0" valueMax="2296.0" valueMin="1800.0">18002296 m</quantity>
) than other members of the species group. One collecting locality of
<taxonomicName authorityName=": Gressitt &amp; Kimoto" authorityYear="1961" box="[650,753,271,295]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="307" phylum="Arthropoda" rank="species" species="fouana">
<emphasis box="[650,753,271,295]" italics="true" pageId="7" pageNumber="307">L. fouana</emphasis>
</taxonomicName>
in Wuding County of
<collectingRegion box="[997,1180,271,295]" country="China" name="Yunnan" pageId="7" pageNumber="307">Yunnan Province</collectingRegion>
is situated in subtropical area. The vegetation is subtropical evergreen forest. The climate is characterized by distinct rainy summer and dry winter, annual temperature generally ranges 6 to 22°C. The forests are composed of tall trees, woody vines and epiphytes. The host plant
<taxonomicName box="[195,341,376,399]" class="Magnoliopsida" family="Hernandiaceae" genus="Dioscorea" kingdom="Plantae" order="Laurales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[195,305,376,399]" italics="true" pageId="7" pageNumber="307">Dioscorea</emphasis>
sp.
</taxonomicName>
shares habitat with other plants such as
<taxonomicName box="[758,956,376,399]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis box="[758,956,376,399]" italics="true" pageId="7" pageNumber="307">Pinus yunnanensis</emphasis>
</taxonomicName>
(
<taxonomicName box="[968,1069,375,399]" class="Pinopsida" family="Pinaceae" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="family">Pinaceae</taxonomicName>
),
<taxonomicName box="[1085,1181,375,399]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[1085,1146,375,399]" italics="true" pageId="7" pageNumber="307">Alnus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName authorityName="S.F.Gray" authorityYear="1822" box="[1194,1317,375,399]" class="Magnoliopsida" family="Betulaceae" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="family">Betulaceae</taxonomicName>
),
<taxonomicName class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[1333,1452,375,399]" italics="true" pageId="7" pageNumber="307">Eucalyptus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName box="[188,306,410,434]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
),
<taxonomicName box="[333,482,410,434]" class="Polypodiopsida" family="Pteridaceae" genus="Adiantum" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[333,437,410,434]" italics="true" pageId="7" pageNumber="307">Adiantum</emphasis>
sp.
</taxonomicName>
(
<taxonomicName box="[503,632,410,434]" class="Polypodiopsida" family="Pteridaceae" kingdom="Plantae" order="Polypodiales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="family">Pteridaceae</taxonomicName>
),
<taxonomicName box="[658,772,410,434]" class="Magnoliopsida" family="Caprifoliaceae" genus="Abelia" kingdom="Plantae" order="Dipsacales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[658,727,410,434]" italics="true" pageId="7" pageNumber="307">Abelia</emphasis>
sp.
</taxonomicName>
(
<taxonomicName box="[794,956,410,434]" class="Magnoliopsida" family="Caprifoliaceae" kingdom="Plantae" order="Dipsacales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="family">Caprifoliaceae</taxonomicName>
),
<taxonomicName box="[981,1133,410,433]" class="Magnoliopsida" family="Asteraceae" genus="Ageratina" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[981,1087,410,433]" italics="true" pageId="7" pageNumber="307">Ageratina</emphasis>
sp.
</taxonomicName>
(
<taxonomicName box="[1154,1277,410,434]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
),
<taxonomicName box="[1303,1445,410,433]" class="Magnoliopsida" family="Asteraceae" genus="Artemisia" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[1303,1407,410,433]" italics="true" pageId="7" pageNumber="307">Artemisia</emphasis>
sp.
</taxonomicName>
(
<taxonomicName box="[143,263,444,468]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) and
<taxonomicName box="[323,419,445,468]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[323,382,445,468]" italics="true" pageId="7" pageNumber="307">Ficus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName box="[433,545,444,468]" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" order="Rosales" pageId="7" pageNumber="307" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7176024" ID-Zenodo-Dep="7176024" httpUri="https://zenodo.org/record/7176024/files/figure.png" pageId="6" pageNumber="306" startId="6.[155,227,2006,2028]" targetBox="[220,1371,482,1970]" targetPageId="6">
<paragraph blockId="6.[155,1432,2006,2060]" pageId="6" pageNumber="306">
Figures 710. Habitus of
<taxonomicName box="[403,560,2006,2028]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="species" species="undefined-7">
<emphasis box="[403,495,2006,2027]" italics="true" pageId="6" pageNumber="306">Lilioceris</emphasis>
spp. 7
</taxonomicName>
8.
<taxonomicName authorityName=": Gressitt &amp; Kimoto" authorityYear="1961" box="[596,688,2006,2028]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="species" species="fouana">
<emphasis box="[596,688,2006,2028]" italics="true" pageId="6" pageNumber="306">L. fouana</emphasis>
</taxonomicName>
, ♀, syntype, China (Yunnan), photoed by Christophe Rivier. 910.
<taxonomicName baseAuthorityName="Weise" baseAuthorityYear="1922" box="[1343,1426,2007,2028]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="306" phylum="Arthropoda" rank="species" species="neptis">
<emphasis box="[1343,1426,2007,2028]" italics="true" pageId="6" pageNumber="306">L. neptis</emphasis>
</taxonomicName>
, ♂, China (Zhejiang: Tianmu Shan). 7, 9. Dorsal view. 8, 10. Ventral view. Scale bars =5.0 mm.
</paragraph>
</caption>
<paragraph blockId="7.[136,1452,236,607]" pageId="7" pageNumber="307">
Remarks. The species is easy to recognize due to its small size, and complete but sparse punctures in each stria. It has been incorrectly placed in subgenus
<taxonomicName authority="Monros, 1959" authorityName="Monros" authorityYear="1959" box="[522,770,514,538]" class="Insecta" family="Chrysomelidae" genus="Chujoita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis box="[522,615,514,538]" italics="true" pageId="7" pageNumber="307">Chujoita</emphasis>
Monrós, 1959
</taxonomicName>
(Bezděk &amp; Schmitt, 2017). According to our observation on the
<typeStatus box="[136,182,549,573]" pageId="7" pageNumber="307">type</typeStatus>
specimen (
<figureCitation box="[306,371,548,572]" captionStart="Figures 710" captionStartId="6.[155,227,2006,2028]" captionTargetBox="[220,1371,482,1970]" captionTargetId="figure-143@6.[189,1399,479,1976]" captionTargetPageId="6" captionText="Figures 710. Habitus of Lilioceris spp. 78. L. fouana, ♀, syntype, China (Yunnan), photoed by Christophe Rivier. 910. L. neptis, ♂, China (Zhejiang: Tianmu Shan). 7, 9. Dorsal view. 8, 10. Ventral view. Scale bars=5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7176024" httpUri="https://zenodo.org/record/7176024/files/figure.png" pageId="7" pageNumber="307">Fig. 7</figureCitation>
), the species does not have obvious bulge on the elytral base, so it should be a member of subgenus
<taxonomicName authority="Reitter, 1913" authorityName="Reitter" authorityYear="1913" box="[136,387,583,607]" class="Insecta" family="Chrysomelidae" genus="Lilioceris" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis box="[136,238,583,607]" italics="true" pageId="7" pageNumber="307">Lilioceris</emphasis>
Reitter, 1913
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>