260 lines
26 KiB
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260 lines
26 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1095.80097" ID-GBIF-Dataset="b3e3b39b-7242-4922-a644-7dc956ceb1df" ID-PMC="PMC9021148" ID-Pensoft-Pub="1313-2970-1095-123" ID-Pensoft-UUID="86ED0D023E645B38A2E138BC2A97D390" ID-PubMed="35836686" ID-ZooBank="FF352C2484774820808E324A32847888" ModsDocID="1313-2970-1095-123" checkinTime="1649881579121" checkinUser="pensoft" docAuthor="Murakami, Hiroyuki, Gerstmeier, Roland & Sakai, Kaoru" docDate="2022" docId="754546A10C645726B8464BAF91904BCC" docLanguage="en" docName="ZooKeys 1095: 123-142" docOrigin="ZooKeys 1095" docPubDate="2022-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1095.80097" docTitle="Tillicera spinosa Murakami & Gerstmeier & Sakai 2022, sp. nov." docType="treatment" docUuid="2E35F6F1-8F4A-4BB9-957F-82173B1EB70E" docUuidSource="ZooBank" docVersion="4" id="86ED0D023E645B38A2E138BC2A97D390" lastPageNumber="123" masterDocId="86ED0D023E645B38A2E138BC2A97D390" masterDocTitle="Description of two new species of the genus Tillicera Spinola (Coleoptera, Cleridae, Clerinae), with new synonyms, new distributional records, and an updated key" masterLastPageNumber="142" masterPageNumber="123" pageNumber="123" updateTime="1668151660150" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Description of two new species of the genus Tillicera Spinola (Coleoptera, Cleridae, Clerinae), with new synonyms, new distributional records, and an updated key</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Murakami, Hiroyuki</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4297-8221</mods:nameIdentifier>
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<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, 3 - 5 - 7 Tarumi, Matsuyama, 790 - 8566 Japan</mods:affiliation>
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<mods:nameIdentifier type="email">hiroyuki068@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gerstmeier, Roland</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0041-5703</mods:nameIdentifier>
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<mods:affiliation>Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, 81247 Muenchen, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sakai, Kaoru</mods:namePart>
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<mods:affiliation>Ota-ku, Tokyo, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-04-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1095</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>123</mods:start>
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<mods:end>142</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1095.80097</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1095.80097</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1095-123</mods:identifier>
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<mods:identifier type="ZooBank">FF352C2484774820808E324A32847888</mods:identifier>
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<mods:identifier type="Pensoft-UUID">86ED0D023E645B38A2E138BC2A97D390</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="194594647" LSID="urn:lsid:zoobank.org:act:2E35F6F1-8F4A-4BB9-957F-82173B1EB70E" httpUri="http://treatment.plazi.org/id/754546A10C645726B8464BAF91904BCC" lastPageNumber="123" pageId="0" pageNumber="123">
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<subSubSection pageId="0" pageNumber="123" type="nomenclature">
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<paragraph pageId="0" pageNumber="123">
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<taxonomicName LSID="http://zoobank.org/2E35F6F1-8F4A-4BB9-957F-82173B1EB70E" authority="Murakami & Gerstmeier & Sakai, 2022" authorityName="Murakami & Gerstmeier & Sakai" authorityYear="2022" class="Insecta" family="Cleridae" genus="Tillicera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tillicera spinosa" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">Tillicera spinosa</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="123">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="123" type="description">
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<paragraph pageId="0" pageNumber="123">
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<figureCitation captionStart="Figures 23–27" captionStartId="F3" captionText="Figures 23 - 27. Habitus of Tillicera 23 Tillicera fortis sp. nov., Holotype, male 24 ditto, paratype, female 25 T. spinosa sp. nov., Holotype, male 26 ditto, paratype, male 27 ditto, paratype, female. Scale bars 2 mm." figureDoi="10.3897/zookeys.1095.80097.figures23-27" httpUri="https://binary.pensoft.net/fig/672267" pageId="0" pageNumber="123">Figs 25-27</figureCitation>
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<figureCitation captionStart="Figures 28–31" captionStartId="F4" captionText="Figures 28 - 31. Antennae of Tillicera spp. (28 dorsal view 29 - 31 ventral view) 28 male antennomere XI of T. spinosa sp. nov. 29 female antennomere X of T. spinosa sp. nov. 30 female antennomeres VII-IX of T. spinosa sp. nov. 31 male antennomeres IV and V of T. callosa. Abbreviations: P pit-like sensillum SB an area of sensilla basiconica." figureDoi="10.3897/zookeys.1095.80097.figures28-31" httpUri="https://binary.pensoft.net/fig/672268" pageId="0" pageNumber="123">, 28-30</figureCitation>
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<figureCitation captionStart="Figures 41–50" captionStartId="F7" captionText="Figures 41 - 50. Male terminal parts and aedeagus 41 - 45 Tillicera fortis sp. nov. 46 - 50 T. spinosa sp. nov. 41, 46 pygidium 42, 47 ventrite VI 43, 48 spicular fork 44, 49 tegmen in lateral (a), ventral (b) and dorsal (c) views 45, 50 median lobe in lateral (a), ventral (b), and dorsal (c) views." figureDoi="10.3897/zookeys.1095.80097.figures41-50" httpUri="https://binary.pensoft.net/fig/672271" pageId="0" pageNumber="123">, 46-50</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="123" type="types">
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<paragraph pageId="0" pageNumber="123">Types.</paragraph>
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<paragraph pageId="0" pageNumber="123">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Holotype.</emphasis>
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"Doi Pui, 1400-/ 1500 m, Chiang Mai, N. Thailand, 19-VI-1983, T. Shimomura leg". (RGCM, 1 male, erroneously cited in
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<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2359.1.1" author="Gerstmeier, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="123" pagination="1 - 34" refId="B5" refString="Gerstmeier, R, Bernhard, N, 2010. Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae). Zootaxa 2359 (1): 1 - 34, DOI: https://doi.org/10.11646/zootaxa.2359.1.1" title="Revision of the genus Tillicera Spinola, 1841 (Coleoptera: Cleridae, Clerinae)." url="https://doi.org/10.11646/zootaxa.2359.1.1" volume="2359" year="2010">Gerstmeier and Bernhard (2010)</bibRefCitation>
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under
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. auratofasciata" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="auratofasciata">
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<emphasis italics="true" pageId="0" pageNumber="123">T. auratofasciata</emphasis>
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</taxonomicName>
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).
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<emphasis bold="true" pageId="0" pageNumber="123">
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<emphasis italics="true" pageId="0" pageNumber="123">Paratypes</emphasis>
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. Laos
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</emphasis>
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: "NE LAOS/ Phu Pan, 1,750 m/ Ban Saleui, Xam Neua/ Houa Phan Prov./ 16-23.VI.2003/ Shinji Nagai leg." (KSCJ, 1 female).
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<emphasis bold="true" pageId="0" pageNumber="123">Myanmar</emphasis>
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: "near Kalaw/ 1,000-1,300 m in alt./ Shan Sta., Myanmar/ 10-25.V.2005/ Y. Kusakabe leg." (EUMJ, 1 male & 1 female; KSCJ, 1 male & 2 females); "Mt. Victoria/ (Natmataung N. P.)/ alt. 1500-2750m/ Kanpelet side", "Chin Sta., Myanmar/ 21-24. V. 2002/ Y. Kusakabe leg." (EUMJ, 1 male; KSCJ, 1 male & 1 female); "Mt. Victoria/ alt. 1500-2000 m/ Mindat side", "Chin Sta., Myanmar/ 13-14.VI.2002/ Y. Kusakabe leg.", (KSCJ, 1 female).
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<emphasis bold="true" pageId="0" pageNumber="123">Thailand</emphasis>
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: "Doi Pui Chieng Mai/ N - THAILAND/ 22.V.1986/ leg." (KSCJ, 1 female); QSBG-2014-0160-0010, Amnat Charoen, Chanuman Dist., Doi Inthanon NP,
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<geoCoordinate degrees="18" direction="south" minutes="32" orientation="latitude" precision="1" seconds="44.4" value="-18.545666">18°32'44.4"S</geoCoordinate>
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,
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<geoCoordinate degrees="98" direction="east" minutes="30" orientation="longitude" precision="15" seconds="53" value="98.514725">98°30'53"E</geoCoordinate>
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, 1376 m, 29.V.-1.VII.2014, Malaise trap, Wichai Srisuka et al. (RGCM, 1 ex.); Same with QSBG
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<normalizedToken originalValue="….">....</normalizedToken>
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-0008 (RGCM, 1 ex.); Same with QSBG
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<normalizedToken originalValue="….">....</normalizedToken>
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-0009 + 0011 (QSBG, 3 exs).
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<emphasis bold="true" pageId="0" pageNumber="123">China</emphasis>
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: S-Yunnan (Xishuangbanna), c. 30 km NW Jinghong, vic. Bameng, 1700-2000 m; Hua Zhuliangzi Mts.,
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<geoCoordinate degrees="22" direction="north" minutes="08.01" orientation="latitude" precision="9" value="22.1335">22°08.01'N</geoCoordinate>
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,
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<geoCoordinate degrees="100" direction="east" minutes="31.54" orientation="longitude" precision="9" value="100.525665">100°31.54'E</geoCoordinate>
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, 1700-2000 m, 30.V.2008, leg. A. Weigel, sec. forest (RGCM, 1 male).
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</paragraph>
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<caption doi="10.3897/zookeys.1095.80097.figures32-35" httpUri="https://binary.pensoft.net/fig/672269" pageId="0" pageNumber="123" start="Figures 32–35" startId="F5">
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<paragraph pageId="0" pageNumber="123">
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<emphasis bold="true" pageId="0" pageNumber="123">Figures 32-35.</emphasis>
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<taxonomicName class="Insecta" family="Cleridae" genus="Hemitrachys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hemitrachys tubericollis" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="tubericollis">
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<emphasis italics="true" pageId="0" pageNumber="123">Hemitrachys tubericollis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="123">32, 33</emphasis>
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habitus in dorsal (
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<emphasis bold="true" pageId="0" pageNumber="123">32</emphasis>
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) and ventral (
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<emphasis bold="true" pageId="0" pageNumber="123">33</emphasis>
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) views
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<emphasis bold="true" pageId="0" pageNumber="123">34</emphasis>
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male antennomeres IX-XI in ventral view
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<emphasis bold="true" pageId="0" pageNumber="123">35</emphasis>
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male antennomeres V-VIII in ventral view. Abbreviations:
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<emphasis bold="true" pageId="0" pageNumber="123">SB</emphasis>
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an area of sensilla basiconica.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="123" type="diagnosis">
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<paragraph pageId="0" pageNumber="123">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="123">
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This species is similar to
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<taxonomicName authorityName="Schenkling" authorityYear="1902" class="Insecta" family="Cleridae" genus="Tillicera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tillicera soror" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="soror">
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<emphasis italics="true" pageId="0" pageNumber="123">Tillicera soror</emphasis>
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</taxonomicName>
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Schenkling, 1902, but distinguished by the following characteristics: antennomere VI-X gradually broadened; metatibiae apically extended to a broad spine; posterior margin of abdominal ventrite V, with broadly V-shaped emargination; phallobase notched at apical 1/5 of total length of tegmen (vs.
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<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. soror" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="soror">
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<emphasis italics="true" pageId="0" pageNumber="123">T. soror</emphasis>
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</taxonomicName>
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: antennomere VIII-IX gradually broadened; hind tibiae not prominent at apex; posterior margin of abdominal sternite V broadly U-shaped and truncate; phallobase deeply notched at apical 1/3).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="123" type="description">
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<paragraph pageId="0" pageNumber="123">Description.</paragraph>
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<paragraph pageId="0" pageNumber="123">
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<emphasis bold="true" pageId="0" pageNumber="123">Male.</emphasis>
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Head, antennomeres IV-XI, base of pronotum and legs brownish black; antennomeres I-III and pronotum reddish. Elytra black except for reddish area near basal suture and yellowish area at basal 1/4 near lateral margin. Head at apex and legs covered with white setae; basal 1/2 of head and pronotum with mingled white and black setae; elytra densely vested with yellowish setae obliquely at suture of basal 1/3 and transversally at lateral margin basal 1/4 and apical 1/4, the remainder vested with black setae (Fig.
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<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Habitus of Tillicera species 1 Tillicera auratofasciata 2 T. aurivillosa 3 T. bibalteata 4 T. callosa 5, 6 T. cleroides 7, 8 T. hirsuta 9, 10 T. ihlei 11, 12 T. javana. Scale bars 2 mm." figureDoi="10.3897/zookeys.1095.80097.figures1-12" httpUri="https://binary.pensoft.net/fig/672265" pageId="0" pageNumber="123">10</figureCitation>
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). This species has two color patterns: the first, antennomeres IV-XI, apex of pronotum, elytra except for area covered with yellowish setae, and legs black (Fig.
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<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Habitus of Tillicera species 1 Tillicera auratofasciata 2 T. aurivillosa 3 T. bibalteata 4 T. callosa 5, 6 T. cleroides 7, 8 T. hirsuta 9, 10 T. ihlei 11, 12 T. javana. Scale bars 2 mm." figureDoi="10.3897/zookeys.1095.80097.figures1-12" httpUri="https://binary.pensoft.net/fig/672265" pageId="0" pageNumber="123">11</figureCitation>
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); the second, head black; apical 1/4 of pronotum brownish black; elytral setae white and black (Fig.
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<figureCitation captionStart="Figures 1–12" captionStartId="F1" captionText="Figures 1 - 12. Habitus of Tillicera species 1 Tillicera auratofasciata 2 T. aurivillosa 3 T. bibalteata 4 T. callosa 5, 6 T. cleroides 7, 8 T. hirsuta 9, 10 T. ihlei 11, 12 T. javana. Scale bars 2 mm." figureDoi="10.3897/zookeys.1095.80097.figures1-12" httpUri="https://binary.pensoft.net/fig/672265" pageId="0" pageNumber="123">12</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="123">
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Head including eyes as wide as pronotum; labrum incised at middle; maxillary terminal palpomeres digitiform; labial terminal palpomeres widely triangular; postgular plate narrow. Antennomere I claviform; II compact; III twice as long as II; IV-X triangular, becoming gradually widened; V-IX (Fig.
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<figureCitation captionStart="Figures 28–31" captionStartId="F4" captionText="Figures 28 - 31. Antennae of Tillicera spp. (28 dorsal view 29 - 31 ventral view) 28 male antennomere XI of T. spinosa sp. nov. 29 female antennomere X of T. spinosa sp. nov. 30 female antennomeres VII-IX of T. spinosa sp. nov. 31 male antennomeres IV and V of T. callosa. Abbreviations: P pit-like sensillum SB an area of sensilla basiconica." figureDoi="10.3897/zookeys.1095.80097.figures28-31" httpUri="https://binary.pensoft.net/fig/672268" pageId="0" pageNumber="123">30</figureCitation>
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) with an area vested with sensilla basiconica; X (Fig.
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<figureCitation captionStart="Figures 28–31" captionStartId="F4" captionText="Figures 28 - 31. Antennae of Tillicera spp. (28 dorsal view 29 - 31 ventral view) 28 male antennomere XI of T. spinosa sp. nov. 29 female antennomere X of T. spinosa sp. nov. 30 female antennomeres VII-IX of T. spinosa sp. nov. 31 male antennomeres IV and V of T. callosa. Abbreviations: P pit-like sensillum SB an area of sensilla basiconica." figureDoi="10.3897/zookeys.1095.80097.figures28-31" httpUri="https://binary.pensoft.net/fig/672268" pageId="0" pageNumber="123">29</figureCitation>
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) with two areas vested with sensilla basiconica at apical margin in ventral view; XI (Fig.
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<figureCitation captionStart="Figures 28–31" captionStartId="F4" captionText="Figures 28 - 31. Antennae of Tillicera spp. (28 dorsal view 29 - 31 ventral view) 28 male antennomere XI of T. spinosa sp. nov. 29 female antennomere X of T. spinosa sp. nov. 30 female antennomeres VII-IX of T. spinosa sp. nov. 31 male antennomeres IV and V of T. callosa. Abbreviations: P pit-like sensillum SB an area of sensilla basiconica." figureDoi="10.3897/zookeys.1095.80097.figures28-31" httpUri="https://binary.pensoft.net/fig/672268" pageId="0" pageNumber="123">28</figureCitation>
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) with small pit-like sensillum in male.
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</paragraph>
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<paragraph pageId="0" pageNumber="123">Pronotum slightly longer than wide, widest at middle, with fine dense punctures. Mesoventrite with short anterior process.</paragraph>
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<paragraph pageId="0" pageNumber="123">Elytra oblong, parallel sided, covered with ten striae; striae I and II absent before basal oblique yellow fascia; III-V or VII extending after middle, sometimes absent before basal oblique yellow fascia; VI or VIII-X rudimentary.</paragraph>
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<paragraph pageId="0" pageNumber="123">Profemora stouter than meso- and metafemora; punctation of meso- and metafemora denser than that of profemora. Tibiae short prominent at apex, with distinct longitudinal carina on dorsal and ventral surfaces; tibial spur formula 1-2-0; tarsal pulvillar formula 4-4-2; pro- and mesotarsomeres I and II with large lobed pulvilli; III and IV with large bilobed pulvilli; metatarsomeres I and II without pulvilli; III and IV with large bilobed pulvilli; claws with basal denticles.</paragraph>
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<caption doi="10.3897/zookeys.1095.80097.figures36-40" httpUri="https://binary.pensoft.net/fig/672270" pageId="0" pageNumber="123" start="Figures 36–40" startId="F6">
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<paragraph pageId="0" pageNumber="123">
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<emphasis bold="true" pageId="0" pageNumber="123">Figures 36-40.</emphasis>
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Male terminal parts and aedeagus of
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<taxonomicName authorityName="Gerstmeier & Bernhard" authorityYear="2010" class="Insecta" family="Cleridae" genus="Tillicera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tillicera callosa" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="callosa">
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<emphasis italics="true" pageId="0" pageNumber="123">Tillicera callosa</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="123">36</emphasis>
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pygidium
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<emphasis bold="true" pageId="0" pageNumber="123">37</emphasis>
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ventrite VI
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<emphasis bold="true" pageId="0" pageNumber="123">38</emphasis>
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spicular fork
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<emphasis bold="true" pageId="0" pageNumber="123">39</emphasis>
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tegmen in lateral (a), ventral (b) and dorsal (c) views
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<emphasis bold="true" pageId="0" pageNumber="123">40</emphasis>
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median lobe in lateral (a), ventral (b), and dorsal (c) views.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="123">
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Abdominal ventrite V deeply emarginated at apical margin. Pygidium (Fig.
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<figureCitation captionStart="Figures 41–50" captionStartId="F7" captionText="Figures 41 - 50. Male terminal parts and aedeagus 41 - 45 Tillicera fortis sp. nov. 46 - 50 T. spinosa sp. nov. 41, 46 pygidium 42, 47 ventrite VI 43, 48 spicular fork 44, 49 tegmen in lateral (a), ventral (b) and dorsal (c) views 45, 50 median lobe in lateral (a), ventral (b), and dorsal (c) views." figureDoi="10.3897/zookeys.1095.80097.figures41-50" httpUri="https://binary.pensoft.net/fig/672271" pageId="0" pageNumber="123">46</figureCitation>
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) narrowly emarginated at apical margin; ventrite VI (Fig.
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<figureCitation captionStart="Figures 41–50" captionStartId="F7" captionText="Figures 41 - 50. Male terminal parts and aedeagus 41 - 45 Tillicera fortis sp. nov. 46 - 50 T. spinosa sp. nov. 41, 46 pygidium 42, 47 ventrite VI 43, 48 spicular fork 44, 49 tegmen in lateral (a), ventral (b) and dorsal (c) views 45, 50 median lobe in lateral (a), ventral (b), and dorsal (c) views." figureDoi="10.3897/zookeys.1095.80097.figures41-50" httpUri="https://binary.pensoft.net/fig/672271" pageId="0" pageNumber="123">47</figureCitation>
|
||
) almost transverse at apical margin; spicular fork long (Fig.
|
||
<figureCitation captionStart="Figures 41–50" captionStartId="F7" captionText="Figures 41 - 50. Male terminal parts and aedeagus 41 - 45 Tillicera fortis sp. nov. 46 - 50 T. spinosa sp. nov. 41, 46 pygidium 42, 47 ventrite VI 43, 48 spicular fork 44, 49 tegmen in lateral (a), ventral (b) and dorsal (c) views 45, 50 median lobe in lateral (a), ventral (b), and dorsal (c) views." figureDoi="10.3897/zookeys.1095.80097.figures41-50" httpUri="https://binary.pensoft.net/fig/672271" pageId="0" pageNumber="123">48</figureCitation>
|
||
), without intraspicular plate.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="123">
|
||
Tegmen (Fig.
|
||
<figureCitation captionStart="Figures 41–50" captionStartId="F7" captionText="Figures 41 - 50. Male terminal parts and aedeagus 41 - 45 Tillicera fortis sp. nov. 46 - 50 T. spinosa sp. nov. 41, 46 pygidium 42, 47 ventrite VI 43, 48 spicular fork 44, 49 tegmen in lateral (a), ventral (b) and dorsal (c) views 45, 50 median lobe in lateral (a), ventral (b), and dorsal (c) views." figureDoi="10.3897/zookeys.1095.80097.figures41-50" httpUri="https://binary.pensoft.net/fig/672271" pageId="0" pageNumber="123">49a-c</figureCitation>
|
||
) with dorsal and ventral sinus at apical 1/5; parameroid lobes tapered posteriorly at apex in lateral view; tegminal arms short, extending from middle to basal 1/4 of total length. Median lobe shorter than tegmen; plates with rows of denticles from apical 1/5 to 2/5 of total length on dorsal and ventral sides (Fig.
|
||
<figureCitation captionStart="Figures 41–50" captionStartId="F7" captionText="Figures 41 - 50. Male terminal parts and aedeagus 41 - 45 Tillicera fortis sp. nov. 46 - 50 T. spinosa sp. nov. 41, 46 pygidium 42, 47 ventrite VI 43, 48 spicular fork 44, 49 tegmen in lateral (a), ventral (b) and dorsal (c) views 45, 50 median lobe in lateral (a), ventral (b), and dorsal (c) views." figureDoi="10.3897/zookeys.1095.80097.figures41-50" httpUri="https://binary.pensoft.net/fig/672271" pageId="0" pageNumber="123">50a-c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="123">
|
||
<emphasis bold="true" pageId="0" pageNumber="123">Female.</emphasis>
|
||
Similar to male but distinguished by antennomere XI without pit-like sensillum, tibial spur formula 1-2-2 and apex of metatibiae not extended to a broad spine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="123" type="description">
|
||
<paragraph pageId="0" pageNumber="123">Measurements and ratios.</paragraph>
|
||
<paragraph pageId="0" pageNumber="123">
|
||
<emphasis bold="true" pageId="0" pageNumber="123">Male</emphasis>
|
||
(
|
||
<emphasis italics="true" pageId="0" pageNumber="123">N</emphasis>
|
||
= 4). BL 6.55-8.30 (7.14) mm; PL 1.75-2.30 (1.99) mm; PW 1.55-2.10 (1.78) mm; EL 4.60-6.00 (5.15) mm; EW 2.10-2.65 (2.29) mm; EyW 0.40-0.55 (0.49) mm; EyD 0.55-0.95 (0.76) mm; PL/PW 1.10-1.14 (1.12); EL/EW 2.09-2.36 (2.25); EL/PL 2.24-2.81 (2.60); EW/PW 1.22-1.35 (1.29); EyD/EyW 1.38-1.73 (1.56).
|
||
<emphasis bold="true" pageId="0" pageNumber="123">Female</emphasis>
|
||
(
|
||
<emphasis italics="true" pageId="0" pageNumber="123">N</emphasis>
|
||
= 4). BL 8.00-8.90 (8.49) mm; PL 2.20-2.50 (2.36) mm; PW 2.00-2.25 (2.10) mm; EL 5.80-6.50 (6.13) mm; EW 2.50-2.75 (2.65) mm; EyW 0.50-0.60 (0.54) mm; EyD 0.95-1.05 (1.00) mm; PL/PW 1.07-1.20 (1.13); EL/EW 2.23-2.36 (2.31); EL/PL 2.42-2.77 (2.59); EW/PW 1.22-1.31 (1.26); EyD/EyW 1.73-2.00 (1.87).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="123" type="etymology">
|
||
<paragraph pageId="0" pageNumber="123">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="123">
|
||
This specific name is derived from the Latin
|
||
<taxonomicName authorityName="Murakami & Gerstmeier & Sakai" authorityYear="2022" class="Insecta" family="Cleridae" genus="Tillicera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tillicera spinosa" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="spinosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="123">Tillicera spinosa</emphasis>
|
||
</taxonomicName>
|
||
(spine), referring to metatibiae apically extended to a broad spine.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1095.80097.figures41-50" httpUri="https://binary.pensoft.net/fig/672271" pageId="0" pageNumber="123" start="Figures 41–50" startId="F7">
|
||
<paragraph pageId="0" pageNumber="123">
|
||
<emphasis bold="true" pageId="0" pageNumber="123">Figures 41-50.</emphasis>
|
||
Male terminal parts and aedeagus
|
||
<emphasis bold="true" pageId="0" pageNumber="123">41-45</emphasis>
|
||
<taxonomicName authorityName="Murakami & Gerstmeier & Sakai" authorityYear="2022" class="Insecta" family="Cleridae" genus="Tillicera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tillicera fortis" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="fortis">
|
||
<emphasis italics="true" pageId="0" pageNumber="123">Tillicera fortis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="123">46-50</emphasis>
|
||
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="T. spinosa" order="Coleoptera" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="spinosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="123">T. spinosa</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="123">41, 46</emphasis>
|
||
pygidium
|
||
<emphasis bold="true" pageId="0" pageNumber="123">42, 47</emphasis>
|
||
ventrite VI
|
||
<emphasis bold="true" pageId="0" pageNumber="123">43, 48</emphasis>
|
||
spicular fork
|
||
<emphasis bold="true" pageId="0" pageNumber="123">44, 49</emphasis>
|
||
tegmen in lateral (a), ventral (b) and dorsal (c) views
|
||
<emphasis bold="true" pageId="0" pageNumber="123">45, 50</emphasis>
|
||
median lobe in lateral (a), ventral (b), and dorsal (c) views.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="123" type="distribution">
|
||
<paragraph pageId="0" pageNumber="123">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="123">Laos, Myanmar, Thailand.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |