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<mods:title id="CCD72D32ED086F6B422A5E1C3677AF0D">Reappraisal of species attributed to Halicarcinus White, 1846 (Crustacea: Decapoda: Brachyura: Hymenosomatidae) with diagnosis of four new genera and one new species from New Ireland, Papua New Guinea</mods:title>
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<emphasis id="CFE0EABC050DFD4FFF22FA80FE5AFA2B" bold="true" box="[151,404,1396,1422]" italics="true" pageId="11" pageNumber="491">Rhynchoplax messor</emphasis>
<bibRefCitation id="99054B5F050DFD4FFE2EFA80FD97FA2B" author="Stimpson" box="[411,601,1396,1422]" pageId="11" pageNumber="514" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in expeditione ad Oceanum Pacificum septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. Pars V. Crustacea Ocypodoida. Proceedings of the Academy of Natural Science of Philadelphia, 10, 93 - 110. Avaliable from: http: // biodiversitylibrary. org / page / 26289702 (Accessed 21 Mar. 2016)" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
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(
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)
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<emphasis id="CFE0EABC050DFD4FFF22FA2DFEA7FA4A" box="[151,361,1497,1519]" italics="true" pageId="11" pageNumber="491">Rhynchoplax messor</emphasis>
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<treatmentCitation id="7C3510BF050DFD4FFE3DFA2DFD99FA4A" author="Stimpson" box="[392,599,1497,1519]" page="109" pageId="11" pageNumber="491" year="1858">
<bibRefCitation id="99054B5F050DFD4FFE3DFA2DFDE4FA4A" author="Stimpson" box="[392,554,1497,1519]" pageId="11" pageNumber="514" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in expeditione ad Oceanum Pacificum septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. Pars V. Crustacea Ocypodoida. Proceedings of the Academy of Natural Science of Philadelphia, 10, 93 - 110. Avaliable from: http: // biodiversitylibrary. org / page / 26289702 (Accessed 21 Mar. 2016)" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
: 109
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.—
<treatmentCitation id="7C3510BF050DFD4FFDC0FA2DFC88FA4A" author="Stimpson" box="[629,838,1497,1519]" page="148" pageId="11" pageNumber="491" year="1907">
<bibRefCitation id="99054B5F050DFD4FFDC0FA2DFCDAFA4A" author="Stimpson" box="[629,788,1497,1519]" pageId="11" pageNumber="514" refString="Stimpson, W. (1907) Report on the Crustacea (Brachyura and Anomura) collected by the North Pacific Exploring Expedition, 1853 - 1856. Smithsonian Miscellaneous Collections, 49 (Art. III), 1 - 240. Avaliable from: http: // biodiversitylibrary. org / page / 14265890 (Accessed 21 Mar. 2016)" type="journal article" year="1907">Stimpson 1907</bibRefCitation>
: 148
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.
<emphasis id="CFE0EABC050DFD4FFCFFFA2EFCAEFA4A" box="[842,864,1498,1519]" italics="true" pageId="11" pageNumber="491"></emphasis>
<treatmentCitation id="7C3510BF050DFD4FFCD5FA2DFC33FA4A" author="Tesch" box="[864,1021,1497,1519]" page="18" pageId="11" pageNumber="491" year="1918">
<bibRefCitation id="99054B5F050DFD4FFCD5FA2DFC19FA4A" author="Tesch" box="[864,983,1497,1519]" pageId="11" pageNumber="514" refString="Tesch, J. J. (1918) Decapoda F. (Decapoda Brachyura continued) Hymenosomatidae, Retropilumnidae, Ocypodidae, Grapsidae and Gecarcinidae. Siboga-Expeditie, 39 c, 1 - 148, pls 1 - 6. Avaliable from: http: // biodiversitylibrary. org / page / 2037493 (Accessed 21 Mar. 2016)" type="journal article" year="1918">Tesch 1918</bibRefCitation>
: 18
</treatmentCitation>
.
<emphasis id="CFE0EABC050DFD4FFBB5FA2EFBD8FA4A" box="[1024,1046,1498,1519]" italics="true" pageId="11" pageNumber="491"></emphasis>
<treatmentCitation id="7C3510BF050DFD4FFBA2FA2DFB73FA4A" author="Sakai" box="[1047,1213,1497,1519]" page="289" pageId="11" pageNumber="491" year="1934">
<bibRefCitation id="99054B5F050DFD4FFBA2FA2DFB42FA4A" author="Sakai" box="[1047,1164,1497,1519]" pageId="11" pageNumber="514" refString="Sakai, T. (1934) Brachyura from the coast of Kyusyu, Japan. Science Reports of the Tokyo Bunrika Daigaku, Section B, 1 (25), 281 - 330." type="journal article" year="1934">Sakai 1934</bibRefCitation>
: 289
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, fig. 5a.
<emphasis id="CFE0EABC050DFD4FFAB8FA2EFAEDFA4A" box="[1293,1315,1498,1519]" italics="true" pageId="11" pageNumber="491"></emphasis>
<treatmentCitation id="7C3510BF050DFD4FFA91FA2DFF2DF9AB" author="Sakai" page="70" pageId="11" pageNumber="491" year="1935">
<bibRefCitation id="99054B5F050DFD4FFA91FA2DFA56FA4A" author="Sakai" box="[1316,1432,1497,1519]" pageId="11" pageNumber="514" refString="Sakai, T. (1935) Crabs of Japan, 66 Plates in Life Colours with Descriptions. Sanseido, Tokyo, x + 239 pp., 66 pls." type="book" year="1935">Sakai 1935</bibRefCitation>
: 70
</treatmentCitation>
, fig. 27.
<emphasis id="CFE0EABC050DFD4FFE84FA0DFE84F9AB" box="[305,330,1529,1550]" italics="true" pageId="11" pageNumber="491"></emphasis>
<treatmentCitation id="7C3510BF050DFD4FFEF9FA0CFE3EF9AB" author="Sakai" box="[332,496,1528,1550]" page="197" pageId="11" pageNumber="491" year="1938">
<bibRefCitation id="99054B5F050DFD4FFEF9FA0CFE0EF9AB" author="Sakai" box="[332,448,1528,1550]" pageId="11" pageNumber="514" refString="Sakai, T. (1938) Studies on the crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo C., Tokyo, pp. 193 - 364, pls 20 - 41." type="book chapter" year="1938">Sakai 1938</bibRefCitation>
: 197
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198, fig. 1, pl. 20 fig. 2.
<emphasis id="CFE0EABC050DFD4FFD59FA0DFCCCF9AB" box="[748,770,1529,1550]" italics="true" pageId="11" pageNumber="491"></emphasis>
<treatmentCitation id="7C3510BF050DFD4FFCB6FA0CFC52F9AB" author="Sakai" box="[771,924,1528,1550]" page="63" pageId="11" pageNumber="491" year="1965">
<bibRefCitation id="99054B5F050DFD4FFCB6FA0CFCB9F9AB" author="Sakai" box="[771,887,1528,1550]" pageId="11" pageNumber="514" refString="Sakai, T. (1965) The Crabs of Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen Co. Ltd, Tokyo, 206 pp. [English text], 100 plates, 92 pp. [Japanese text], 32 pp. of bibiliography and index." type="book" year="1965">Sakai 1965</bibRefCitation>
: 63
</treatmentCitation>
, fig. 9, pl. 25 fig. 2.
<treatmentCitation id="7C3510BF050DFD4FFB3CFA0CFAE0F9AB" author="Sakai" box="[1161,1326,1528,1550]" page="147" pageId="11" pageNumber="491" year="1976">
<bibRefCitation id="99054B5F050DFD4FFB3CFA0CFB31F9AB" author="Sakai" box="[1161,1279,1528,1550]" pageId="11" pageNumber="514" refString="Sakai, T. (1976) Crabs of Japan and Adjacent Seas. Kodansha Ltd, Tokyo, 773 pp [English text], volume of 251 pls." type="book" year="1976">Sakai 1976</bibRefCitation>
: 147
</treatmentCitation>
148, textfig. 76c (English text), pl. 46 fig. 2, 92 (Japanese text).
<treatmentCitation id="7C3510BF050DFD4FFCA2F9E2FC1CF989" author="Miyake" box="[791,978,1558,1580]" page="192" pageId="11" pageNumber="491" year="1983">
<bibRefCitation id="99054B5F050DFD4FFCA2F9E2FC6FF989" author="Miyake" box="[791,929,1558,1580]" pageId="11" pageNumber="513" refString="Miyake, S. (1983) Japanese Crustacean Decapods and Stomatopods in Color. Vol. II. Hoikusha Publishing Co., Ltd., Osaka. 277 pp., 64 pls." type="book" year="1983">Miyake 1983</bibRefCitation>
: 192
</treatmentCitation>
, pl. 64 fig. 6.
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</paragraph>
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<taxonomicName id="3A944D2D050DFD4FFF22F9C1FED4F9EE" ID-CoL="7PJ4L" box="[151,282,1589,1611]" class="Malacostraca" family="Hymenosomatidae" genus="Halicarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="491" phylum="Arthropoda" rank="genus">Halicarcinus</taxonomicName>
messor—
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<treatmentCitation id="7C3510BF050DFD4FFECBF9C2FDE6F9EE" author="Lucas" box="[382,552,1589,1611]" page="163" pageId="11" pageNumber="491" year="1980">
<bibRefCitation id="99054B5F050DFD4FFECBF9C2FE39F9EE" author="Lucas" box="[382,503,1589,1611]" pageId="11" pageNumber="513" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33, 148 - 247." type="journal article" year="1980">Lucas 1980</bibRefCitation>
: 163
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, fig. 9A.
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<treatmentCitation id="7C3510BF050DFD4FFD23F9C1FCB4F9EE" author="Dai" box="[662,890,1589,1611]" page="116" pageId="11" pageNumber="491" year="1991">
<bibRefCitation id="99054B5F050DFD4FFD23F9C1FC84F9EE" author="Dai" box="[662,842,1589,1611]" pageId="11" pageNumber="511" refString="Dai, A. Y. &amp; Yang, S. L. (1991) Crabs of the China Seas. China Oceaan Press, Beijing, 682 pp." type="book" year="1991">Dai &amp; Yang 1991</bibRefCitation>
: 116
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117, fig. 59.
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<treatmentCitation id="7C3510BF050DFD4FFBA0F9C1FB7FF9EE" author="Kim" box="[1045,1201,1589,1611]" page="257" pageId="11" pageNumber="491" year="1992">
<bibRefCitation id="99054B5F050DFD4FFBA0F9C1FBB1F9EE" author="Kim" box="[1045,1151,1589,1611]" pageId="11" pageNumber="512" refString="Kim, W. (1992) The brachyuran crabs (Crustacea: Decapoda) from Ulreung Island, Korea. Korean Journal of Zoology, 35, 256 - 261. Avaliable from: http: // koreascience. or. kr / article / ArticleFullRecord. jsp? cn = HGDMB 7 _ 1992 _ v 35 n 2 _ 256 &amp; ordernum = 15 (Accessed 21 Mar. 2016)" type="journal article" year="1992">Kim 1992</bibRefCitation>
: 257
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. Ng
<emphasis id="CFE0EABC050DFD4FFB4BF9C2FAE4F9EE" box="[1278,1322,1589,1611]" italics="true" pageId="11" pageNumber="491">et al</emphasis>
. 1999: 88 89, fig. 4.
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</paragraph>
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Not
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<emphasis id="CFE0EABC050DFD4FFF76F986FE5BF92D" box="[195,405,1650,1672]" italics="true" pageId="11" pageNumber="491">Rhynchoplax messor</emphasis>
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<treatmentCitation id="7C3510BF050DFD4FFE00F986FDA1F92D" author="Yokoya" box="[437,623,1650,1672]" page="761" pageId="11" pageNumber="491" year="1928">
<bibRefCitation id="99054B5F050DFD4FFE00F986FDF1F92D" author="Yokoya" box="[437,575,1650,1672]" pageId="11" pageNumber="514" refString="Yokoya, Y. (1928) Report of the biological survey of Mutsu Bay. 10. Brachyura and crab-shaped Anomura. Science Report of the Tohoku Imperial University, ser. 4, 3, 757 - 784." type="journal article" year="1928">Yokoya 1928</bibRefCitation>
: 761
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762, fig. 1 [=
<taxonomicName id="3A944D2D050DFD4FFCB2F986FB69F92D" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[775,1191,1650,1672]" class="Malacostraca" family="Hymenosomatidae" genus="Stimpsoplax" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="491" phylum="Arthropoda" rank="species" species="setirostris">
<emphasis id="CFE0EABC050DFD4FFCB2F986FC23F92D" box="[775,1005,1650,1672]" italics="true" pageId="11" pageNumber="491">Stimpsoplax setirostris</emphasis>
(
<bibRefCitation id="99054B5F050DFD4FFC4CF986FB51F92D" author="Stimpson" box="[1017,1183,1650,1672]" pageId="11" pageNumber="514" refString="Stimpson, W. (1858) Prodromus descriptionis animalium evertebratorum, quae in expeditione ad Oceanum Pacificum septentrionalem, a Republica Federata missa, Cadwaladaro Ringgold et Johanne Rodgers ducibus, observavit et descripsit. Pars V. Crustacea Ocypodoida. Proceedings of the Academy of Natural Science of Philadelphia, 10, 93 - 110. Avaliable from: http: // biodiversitylibrary. org / page / 26289702 (Accessed 21 Mar. 2016)" type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
)
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<paragraph id="FD2B36AE050DFD4FFF22F946FBB1F8F5" blockId="11.[151,1436,1714,1872]" pageId="11" pageNumber="491">
<emphasis id="CFE0EABC050DFD4FFF22F946FEB5F96E" bold="true" box="[151,379,1714,1739]" pageId="11" pageNumber="491">Material examined</emphasis>
.
<collectingCountry id="8583763E050DFD4FFE39F947FE01F96F" box="[396,463,1715,1738]" name="Japan" pageId="11" pageNumber="491">Japan</collectingCountry>
. Chiba Prefecture, Ubara, Katsuura, intertidal,
<date id="892A106E050DFD4FFC5EF947FBB4F96E" box="[1003,1146,1714,1739]" pageId="11" pageNumber="491" value="1996-06-03">3 June 1996</date>
, coll. E. Nishi, CBM-ZC 3507 (
<specimenCount id="EB92FD27050DFD4FFF57F920FEE1F948" box="[226,303,1748,1773]" pageId="11" pageNumber="491" type="male">1 male</specimenCount>
,
<quantity id="3A6C9B4B050DFD4FFE89F920FE59F949" box="[316,407,1748,1772]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="11" pageNumber="491" unit="mm" value="4.8">4.8 mm</quantity>
). Hasama, Tateyama, intertidal,
<date id="892A106E050DFD4FFCB0F920FC55F949" box="[773,923,1748,1773]" pageId="11" pageNumber="491" value="2001-07-22">22 July 2001</date>
, coll. T. Komai, CBM-ZC 6024 (1 ovigerous female,
<quantity id="3A6C9B4B050DFD4FFF47F902FE83F8AB" box="[242,333,1782,1806]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="11" pageNumber="491" unit="mm" value="4.3">4.3 mm</quantity>
). [Shizuoka Prefecture], S[h]imoda, AM P.10497 (
<specimenCount id="EB92FD27050DFD4FFC3EF902FC2BF8AB" box="[907,997,1781,1806]" pageId="11" pageNumber="491" type="male">3 males</specimenCount>
, largest
<quantity id="3A6C9B4B050DFD4FFBF1F902FB50F8A8" box="[1092,1182,1782,1806]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="11" pageNumber="491" unit="mm" value="3.3">3.3 mm</quantity>
;
<specimenCount id="EB92FD27050DFD4FFB1EF902FAD5F8AB" box="[1195,1307,1781,1806]" pageId="11" pageNumber="491" type="female">3 females</specimenCount>
, largest 3.0 mm, 1 ovigerous female, 3.0 mm). Wakayama Prefecture. Kushimoto, Shionomisaki, Naminoura,
<quantity id="3A6C9B4B050DFD4FFAB0F8E3FAFAF88B" box="[1285,1332,1815,1839]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="491" unit="m" value="9.0">9 m</quantity>
,
<date id="892A106E050DFD4FFAF6F8E2FF1CF8F5" pageId="11" pageNumber="491" value="1978-05-16">16 May 1978</date>
, SCUBA diving, coll. S. Yamaguchi, CBM-ZC 4163 (1 ovigerous female, 4.0 mm).
</paragraph>
<caption id="A9EB6626050AFD48FF22FA1AFDCCF9DB" httpUri="https://zenodo.org/record/258480/files/figure.svg" pageId="12" pageNumber="492" targetBox="[193,1390,200,1440]" targetPageId="12">
<paragraph id="FD2B36AE050AFD48FF22FA1AFDCCF9DB" blockId="12.[151,1436,1518,1662]" pageId="12" pageNumber="492">
<emphasis id="CFE0EABC050AFD48FF22FA1AFEC2F9A6" bold="true" box="[151,268,1518,1540]" pageId="12" pageNumber="492">FIGURE 3</emphasis>
.
<taxonomicName id="3A944D2D050AFD48FEADFA1AFD5FF9A1" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[280,657,1518,1540]" class="Malacostraca" family="Hymenosomatidae" genus="Rhynchoplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="492" phylum="Arthropoda" rank="species" species="messor">
<emphasis id="CFE0EABC050AFD48FEADFA1AFE24F9A1" box="[280,490,1518,1540]" italics="true" pageId="12" pageNumber="492">Rhynchoplax messor</emphasis>
Stimpson, 1858
</taxonomicName>
. Male, cl 3.3 mm, AM P.10497: a, rostrum and eyestalks; b, anterior carapace and rostrum, right side; c, buccal region, ventral; d, thoracic sternum (penial process shown); e, pleon; f, left cheliped, lateral face; g, pereopod (uncertain); h, gonopod 1; i, gonopod 2; Female, cl 3.0 mm, AM P.10497: j, left lateral view, pleon slightly open; k, right thoracic sternites 7, 8, lateral margins of pleomeres 1, 2 (branchiosternal canal aperture shown). Upper scale bar = 1 mm; lower scale bar = 0. 5 mm.
</paragraph>
</caption>
<subSubSection id="B58E6525050AFD49FF72F951FB52FEC1" lastPageId="13" lastPageNumber="493" pageId="12" pageNumber="492" type="diagnosis">
<paragraph id="FD2B36AE050AFD49FF72F951FB52FEC1" blockId="12.[151,1437,1701,2014]" lastBlockId="13.[151,1437,151,608]" lastPageId="13" lastPageNumber="493" pageId="12" pageNumber="492">
<emphasis id="CFE0EABC050AFD48FF72F951FE8FF91B" bold="true" box="[199,321,1701,1726]" pageId="12" pageNumber="492">Diagnosis.</emphasis>
Carapace 1.1 times as wide as long; subhepatic region inflated, forming low elevation visible in dorsal view, with 2 conspicuous tubercles inferior to subhepatic elevation (not visible in dorsal view); anterolateral margin with minute tubercles (sometimes greatly reduced); epibranchial spine present, sharp; dorsum with gastrocardiac groove, cardiac median ridge. Male rostrum about 0.3 as long as carapace, spatulate with slightly convex lateral margins, with pair of small but sharp supraocular spines proximally; apex without long setae. Female rostrum narrower, tapered, length similar to that of male. Supraocular eave poorly developed; postocular margin with small acute spine. Supraocular spines small, at base of rostrum just mesial to base of eyestalk. Eyestalks compact, with small protuberance on anterior surface ventrally. Maxilliped 3 merus rounded distolaterally. Cheliped in male at most twice carapace length, with fairly swollen propodus, fixed finger with denticulate cutting edge; dactylus with simple proximal tooth, excavate in middle third; gape obscured by dense mat of setae; carpus short, cup-like; merus widened distally, with blunt dorsodistal tubercle. Pereopods 25 meri each with prominent dorsodistal tubercle; pereopod 2 dactylus weakly arcuate, not falcate, dactyli of pereopods 35 falcate. Male gonopore with prominent penial process. Gonopod 1 with swollen base, almost straight section, strongly curved tip terminating in fine apex. Gonopod 2 small, strongly flattened, terminating in short, non-tapered, terminally truncate tube-like process; lateral margin expanded into angular lobe, mesial margin gently concave.
</paragraph>
</subSubSection>
<subSubSection id="B58E6525050BFD49FF72FE9BFBE7FE22" box="[199,1065,367,392]" pageId="13" pageNumber="493" type="distribution">
<paragraph id="FD2B36AE050BFD49FF72FE9BFBE7FE22" blockId="13.[151,1437,151,608]" box="[199,1065,367,392]" pageId="13" pageNumber="493">
<emphasis id="CFE0EABC050BFD49FF72FE9BFE97FE2D" bold="true" box="[199,345,367,392]" pageId="13" pageNumber="493">Distribution</emphasis>
. East Asian endemic:
<collectingCountry id="8583763E050BFD49FDE1FE84FD59FE22" box="[596,663,368,391]" name="Japan" pageId="13" pageNumber="493">Japan</collectingCountry>
,
<collectingCountry id="8583763E050BFD49FD17FE84FD28FE22" box="[674,742,368,391]" name="South Korea" pageId="13" pageNumber="493">Korea</collectingCountry>
and
<collectingCountry id="8583763E050BFD49FCA8FE9BFCADFE2D" box="[797,867,367,392]" name="China" pageId="13" pageNumber="493">China</collectingCountry>
, intertidal to
<specimenCount id="EB92FD27050BFD49FC42FE84FBE7FE22" box="[1015,1065,368,392]" pageId="13" pageNumber="493" type="generic">
<quantity id="3A6C9B4B050BFD49FC42FE84FBEBFE22" box="[1015,1061,368,392]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="493" unit="m" value="9.0">9 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="B58E6525050BFD49FF72FE60FE1EFDC5" pageId="13" pageNumber="493" type="discussion">
<paragraph id="FD2B36AE050BFD49FF72FE60FE1EFDC5" blockId="13.[151,1437,151,608]" pageId="13" pageNumber="493">
<emphasis id="CFE0EABC050BFD49FF72FE60FEFAFE08" bold="true" box="[199,308,404,429]" pageId="13" pageNumber="493">Remarks</emphasis>
. This species has been mentioned in literature often and the dorsal habitus illustrated several times. Sakai (
<bibRefCitation id="99054B5F050BFD49FF51FE4CFEEEFE75" author="Sakai" box="[228,288,440,464]" pageId="13" pageNumber="514" refString="Sakai, T. (1934) Brachyura from the coast of Kyusyu, Japan. Science Reports of the Tokyo Bunrika Daigaku, Section B, 1 (25), 281 - 330." type="journal article" year="1934">1934</bibRefCitation>
,
<bibRefCitation id="99054B5F050BFD49FE9EFE43FEA8FE75" author="Sakai" box="[299,358,439,464]" pageId="13" pageNumber="514" refString="Sakai, T. (1935) Crabs of Japan, 66 Plates in Life Colours with Descriptions. Sanseido, Tokyo, x + 239 pp., 66 pls." type="book" year="1935">1935</bibRefCitation>
,
<bibRefCitation id="99054B5F050BFD49FEC5FE43FE67FE75" author="Sakai" box="[368,425,439,464]" pageId="13" pageNumber="514" refString="Sakai, T. (1976) Crabs of Japan and Adjacent Seas. Kodansha Ltd, Tokyo, 773 pp [English text], volume of 251 pls." type="book" year="1976">1976</bibRefCitation>
) and
<bibRefCitation id="99054B5F050BFD49FE52FE4CFDB2FE75" author="Lucas" box="[487,636,440,464]" pageId="13" pageNumber="513" refString="Lucas, J. S. (1980) Spider crabs of the family Hymenosomatidae (Crustacea; Brachyura) with particular reference to Australian species: systematics and biology. Records of the Australian Museum, 33, 148 - 247." type="journal article" year="1980">Lucas (1980)</bibRefCitation>
figured the gonopod 1.
<bibRefCitation id="99054B5F050BFD49FC32FE43FBD8FE75" author="Sakai" box="[903,1046,439,464]" pageId="13" pageNumber="514" refString="Sakai, T. (1938) Studies on the crabs of Japan. III. Brachygnatha, Oxyrhyncha. Yokendo C., Tokyo, pp. 193 - 364, pls 20 - 41." type="book chapter" year="1938">Sakai (1938)</bibRefCitation>
figured the dactylus of pereopod 4 and the male pleomeres 3 and 4 erroneously as free.
<bibRefCitation id="99054B5F050BFD49FD4BFE28FC11FE51" author="Sakai" box="[766,991,476,501]" pageId="13" pageNumber="514" refString="Sakai, T. (1965) The Crabs of Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen Co. Ltd, Tokyo, 206 pp. [English text], 100 plates, 92 pp. [Japanese text], 32 pp. of bibiliography and index." type="book" year="1965">Sakai (1965: fig. 9)</bibRefCitation>
figured the maxilliped 3 and the male pleon, and again stated that the male abdomen is composed of six distinct segments. Ng
<emphasis id="CFE0EABC050BFD49FB2DFDF5FB1FFDBD" box="[1176,1233,511,536]" italics="true" pageId="13" pageNumber="493">et al.</emphasis>
(1999) illustrated variation of the male rostrum. Re-examination of material from the
<typeStatus id="222F880C050BFD49FC61FDD1FBCAFD98" box="[980,1028,549,573]" pageId="13" pageNumber="493">type</typeStatus>
locality and elsewhere confirmed characters in earlier figures.
</paragraph>
</subSubSection>
</treatment>
</document>