432 lines
62 KiB
XML
432 lines
62 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1168.100798" ID-Pensoft-Pub="1313-2970-1168-295" ID-Pensoft-UUID="F19054F21F3D516FA0292C896F566182" ID-ZooBank="745FFCD5E3E84DC1B76CC82AD1AC5C64" ModsDocID="1313-2970-1168-295" checkinTime="1688439520051" checkinUser="pensoft" docAuthor="Hou, Yanmeng, Zhao, Lingchen & Zhang, Feng" docDate="2023" docId="5BA0B899E7565AA9852F35CCF22EC2E1" docLanguage="en" docName="ZooKeys 1168: 295-327" docOrigin="ZooKeys 1168" docPubDate="2023-07-03" docSource="http://dx.doi.org/10.3897/zookeys.1168.100798" docTitle="Catatemnus scaber Hou & Zhao & Zhang 2023, sp. nov." docType="treatment" docUuid="84384592-433C-4CF6-9093-5C519173C6FA" docUuidSource="ZooBank" docVersion="1" id="F19054F21F3D516FA0292C896F566182" lastPageNumber="295" masterDocId="F19054F21F3D516FA0292C896F566182" masterDocTitle="First record of the genus Catatemnus Beier, 1932 from China, with the description of six new species (Pseudoscorpiones, Atemnidae)" masterLastPageNumber="327" masterPageNumber="295" pageNumber="295" updateTime="1688439520051" updateUser="pensoft">
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<mods:titleInfo id="B0544CFF57C183D69B1C9440BF27A43E">
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<mods:title id="8D9796637303B26D9DE39108028596AB">First record of the genus Catatemnus Beier, 1932 from China, with the description of six new species (Pseudoscorpiones, Atemnidae)</mods:title>
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<mods:name id="3C7B56CA1139B9E169A06553C9E11B9E" type="personal">
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<mods:namePart id="B684E1C9975D5538D967FF7ECF736C95">Hou, Yanmeng</mods:namePart>
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<mods:affiliation id="DED3A1B369D549E7CD98503D22203F38">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="C7ED2D326F616911B165B2C4F4256946">Zhao, Lingchen</mods:namePart>
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<mods:nameIdentifier id="7E655465B5B0B7C49919C2318A3E8437" type="ORCID">https://orcid.org/0000-0001-8477-836X</mods:nameIdentifier>
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<mods:affiliation id="4FF87AF5BE73215AC1BAF21373193F39">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China & Changzhi Tunliu District No. 6 Middle School, Changzhi, Shanxi 046100, China</mods:affiliation>
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<mods:namePart id="AA385BCF72C8E6FB82BCD47DFE53B24E">Zhang, Feng</mods:namePart>
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<mods:nameIdentifier id="6ABE2725C35FFACD92130E4A3F283F6F" type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation id="9ECE02786C9AAA31B852CB0FBFFC5C6F">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier id="299608B61EA62BA4F592FA2A3E944E06" type="email">dudu06042001@163.com</mods:nameIdentifier>
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<mods:title id="F6ABB4A5E79C1952B995C8AECE8AD9AD">ZooKeys</mods:title>
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<mods:date id="EB26F36A41ED406503C58052DA5BFE0C">2023</mods:date>
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<mods:number id="5658F8AE4A96021C9467061C14B1A5DB">2023-07-03</mods:number>
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<mods:number id="A6717165064D97FCFE81EA8A36584A60">1168</mods:number>
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<mods:identifier id="F3AE0AEF48766AF3AE4E8B6B010DC0BE" type="DOI">http://dx.doi.org/10.3897/zookeys.1168.100798</mods:identifier>
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<treatment id="5BA0B899E7565AA9852F35CCF22EC2E1" LSID="urn:lsid:zoobank.org:act:84384592-433C-4CF6-9093-5C519173C6FA" httpUri="http://treatment.plazi.org/id/5BA0B899E7565AA9852F35CCF22EC2E1" lastPageNumber="295" pageId="0" pageNumber="295">
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<subSubSection id="E3BF7CB60DF80A605469685ED4FF98E9" pageId="0" pageNumber="295" type="nomenclature">
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<paragraph id="F1674D8CC7C25DE2706D02B832039885" pageId="0" pageNumber="295">
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<taxonomicName id="475EB1D7568F69B2F17169282381F7E3" LSID="https://zoobank.org/84384592-433C-4CF6-9093-5C519173C6FA" authority="Hou & Zhao & Zhang, 2023" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus scaber" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="scaber" status="sp. nov.">Catatemnus scaber</taxonomicName>
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<taxonomicNameLabel id="081766C6DD537B7475CE10E63D796E68" pageId="0" pageNumber="295">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="80762FB19952589F1B9E0557F841245F" pageId="0" pageNumber="295" type="description">
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<paragraph id="5388779FAB8CBF7923FB765042887FA7" pageId="0" pageNumber="295">
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<figureCitation id="0F139DB58CE3CD080057EA2D16FC2894" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Catatemnus scaber sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 2.00 mm." figureDoi="10.3897/zookeys.1168.100798.figure10" httpUri="https://binary.pensoft.net/fig/872309" pageId="0" pageNumber="295">Figs 10</figureCitation>
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<figureCitation id="C7F9BF5872DA36FF0DF62ED3DC63C9BF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Catatemnus scaber sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295">, 11</figureCitation>
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<figureCitation id="7A073F59A5ED75221CE5EB5E317988AD" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">, 12 (粗糙猫伪蝎)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7A92C1CF7B7B2333339668F6BBB310D5" pageId="0" pageNumber="295" type="type material">
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<paragraph id="60776F75B0DBC5DD775356CE61389616" pageId="0" pageNumber="295">Type material.</paragraph>
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<paragraph id="FC2B14B622E7EF1040CF9FD5ED1E60C5" pageId="0" pageNumber="295">
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<emphasis id="565BA1437DD43DC4C740708B1FCA7E8D" bold="true" italics="true" pageId="0" pageNumber="295">Holotype</emphasis>
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: China • ♂; Yunnan Province, Longyang District, Lujiang Town, Bawan Village;
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<geoCoordinate id="1BE212DDB41E6DED0E86AE022FCEA090" degrees="24" direction="north" minutes="56.871" orientation="latitude" precision="1" value="24.94785">24°56.871'N</geoCoordinate>
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,
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<geoCoordinate id="E728E977839AEAE2D3E43363FD299BBB" degrees="98" direction="east" minutes="49.969" orientation="longitude" precision="1" value="98.83282">98°49.969'E</geoCoordinate>
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; 1025 m a.s.l.; 29 May. 2017; Chi Jin, Beibei Zhou and Xiao Zang leg.; Ps.-MHBU-YNBS170101.
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<emphasis id="C7BF310FF21AEDD0A4587123393861CE" bold="true" italics="true" pageId="0" pageNumber="295">Paratypes</emphasis>
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: • 8♂10♀; same data as the holotype; Ps.-MSWU-YNBS170102-19.
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</paragraph>
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</subSubSection>
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<subSubSection id="EFF52954295A9E79B0F09F7DC1E0B71F" pageId="0" pageNumber="295" type="diagnosis">
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<paragraph id="FDD8EBA1FC3A6E1FBE6117029F6902ED" pageId="0" pageNumber="295">Diagnosis (♂♀).</paragraph>
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<paragraph id="421B68D8EE19BBE7C8C22DBC10321684" pageId="0" pageNumber="295">
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This new species is characteristic by: palpal femur (♂) 2.38-2.65
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<normalizedToken id="530FC08B99ED1AE3793AEEC21F3757BE" originalValue="×">x</normalizedToken>
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, (♀) 2.44-2.50
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<normalizedToken id="EB5AC98906D661B6B4D4898E15F45DBD" originalValue="×">x</normalizedToken>
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, chela with pedicel (♂) 2.76-2.83
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<normalizedToken id="4581BCA5CE9FD7B84872B058AC0D236C" originalValue="×">x</normalizedToken>
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, (♀) 2.61
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<normalizedToken id="241EC29433634BA0BC3F2BA075F09F36" originalValue="×">x</normalizedToken>
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, without pedicel (♂) 2.51-2.60
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<normalizedToken id="66318AB80E57248DC8691191BF2301B5" originalValue="×">x</normalizedToken>
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, (♀) 1.57-1.65
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<normalizedToken id="00A0321B8D40C552DAC5F7B585E84DC9" originalValue="×">x</normalizedToken>
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longer than broad; distance between est and esb farther to that of ist and isb, st midway between sb and t; male genitalia: the longitudinal fold of medial diverticula (d) with a projection midway along its length, forming a lobe bulge.
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</paragraph>
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</subSubSection>
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<subSubSection id="775B650905C50C01086DCC4874ED22AE" pageId="0" pageNumber="295" type="description">
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<paragraph id="E5F47654715E20B104ED7FBD44200169" pageId="0" pageNumber="295">Description.</paragraph>
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<paragraph id="D9253C2CC25BF4139BE253A7960DA624" pageId="0" pageNumber="295">
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<emphasis id="3E001241086E7F370E05BC4116AE2486" bold="true" pageId="0" pageNumber="295">Adult males</emphasis>
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(Figs
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<figureCitation id="F8CA12F974451057780D974254EA133B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Catatemnus scaber sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 2.00 mm." figureDoi="10.3897/zookeys.1168.100798.figure10" httpUri="https://binary.pensoft.net/fig/872309" pageId="0" pageNumber="295">10A</figureCitation>
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,
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<figureCitation id="A1CB2C628968934D458EECF36862858B" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Catatemnus scaber sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295">11A-E, G-I</figureCitation>
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,
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<figureCitation id="C6BCA203CCC3B89E1C2DE674F180B3A3" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12A-C, E, F, H, I, K-M</figureCitation>
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).
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<emphasis id="823B95044B8758BB75AD615B8997AEDC" bold="true" italics="true" pageId="0" pageNumber="295">Color</emphasis>
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: carapace and tergites dark brown; pedipalps reddish brown, but paler in trochanter; legs pale yellow.
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<emphasis id="80E2BEF5FC0923952028F9C87DB18663" bold="true" italics="true" pageId="0" pageNumber="295">Carapace</emphasis>
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(Figs
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<figureCitation id="4358272AB8B7B75FC5AA3F2BBA3EBC76" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Catatemnus scaber sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295">11A</figureCitation>
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,
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<figureCitation id="EA5B7BE78934015BFFF313C96CCF72C6" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12A</figureCitation>
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): 1.16-1.33
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<normalizedToken id="D107F9D0C41D4384B3A50C1ACFD0FD34" originalValue="×">x</normalizedToken>
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longer than broad; surface smooth, but with a medial furrow; with two distinct eyespots situated near anterior margin of carapace; anterior margin with six setae, posterior margin with eight setae, each seta acicular and very slightly curved.
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<emphasis id="0B96371C2118BA0EBE406CEC22F0FFD0" bold="true" italics="true" pageId="0" pageNumber="295">Chelicera</emphasis>
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(Fig.
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<figureCitation id="5E54B6D94F3B1FD77C28D1AE438263A2" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12B, C, E</figureCitation>
|
||
): much smaller than carapace length; surface smooth; four setae (sbs absent) and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand; movable finger with one galeal seta (shorter than others); bs and es dentate apically, is and ls long and acute. Fixed finger with five large retrorse teeth and two or three small apical teeth, movable finger with one large teeth; galea present, shorter and with seven or eight branchlets (Fig.
|
||
<figureCitation id="DF5DEE23AA2C8210D665F2CDE8D40EAC" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12C</figureCitation>
|
||
). Serrula interior connected to fixed finger for entire length, proximally modified to form velum, serrula exterior with 26 lamellae, the basal one longest; lamina exterior present. Rallum composed of four blades, the basal two blades shorter than others, the distal two dentated anteriorly, remainder smooth (Fig.
|
||
<figureCitation id="407023C7E9366CD451DB2C06087EA4AF" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12E</figureCitation>
|
||
).
|
||
<emphasis id="7C67C572CF48FDBE921FF161FEAD805C" bold="true" italics="true" pageId="0" pageNumber="295">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation id="76D77EBBA289DC156A6935CEA9E5925D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Catatemnus scaber sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295">11B-D</figureCitation>
|
||
,
|
||
<figureCitation id="6052770F925126F632D563962AD961D4" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12K, M</figureCitation>
|
||
): stout, trochanter 1.62-1.79
|
||
<normalizedToken id="31A35F920F0A8B278AADCB4ED15C3E39" originalValue="×">x</normalizedToken>
|
||
, femur 2.38-2.65
|
||
<normalizedToken id="BA38FAA0FE9C890AB4BF8BCEC50FABF1" originalValue="×">x</normalizedToken>
|
||
, patella 2.08-2.11
|
||
<normalizedToken id="6636F627CF305F381446C2EAB896205C" originalValue="×">x</normalizedToken>
|
||
, chela with pedicel (without pedicel) 2.76-2.83
|
||
<normalizedToken id="6C7121BB3A01F810B57ABA2B08E66456" originalValue="×">x</normalizedToken>
|
||
(2.51-2.60
|
||
<normalizedToken id="714757B9C4F785DFD87567C9C06C0EFD" originalValue="×">x</normalizedToken>
|
||
), hand with pedicel (without pedicel) 1.65-1.73
|
||
<normalizedToken id="5FFE688294779E1A53133FB08CBBDB37" originalValue="×">x</normalizedToken>
|
||
(1.41-1.50
|
||
<normalizedToken id="24ACD0FD28B2A88EA5101E2A8B5A1751" originalValue="×">x</normalizedToken>
|
||
) longer than broad; movable chelal finger 0.64-0.67
|
||
<normalizedToken id="B40141C15E454C1E50DAE032BF8397F5" originalValue="×">x</normalizedToken>
|
||
(0.74-0.78
|
||
<normalizedToken id="4AB863D880105BF70BF7D8F89D0EC4FA" originalValue="×">x</normalizedToken>
|
||
) longer than hand with pedicel (without pedicel) and 0.39-0.40
|
||
<normalizedToken id="929C5705369E893A22111727B6D8EBAB" originalValue="×">x</normalizedToken>
|
||
(0.42-0.44
|
||
<normalizedToken id="674A1CEDA797C691F5F257EE494DFFA8" originalValue="×">x</normalizedToken>
|
||
) longer than chela with pedicel (without pedicel). Setae generally long and acuminate. Retrolateral surface of trochanter, surface of femur and hand granular; trochanter with two inconspicuous tubercles. Fixed chelal finger with eight trichobothria, movable chelal finger with four trichobothria: eb and esb situated at base of fixed finger on retrolateral face, esb slightly distal to eb; ib and isb situated at base of fixed finger on prolateral face, isb slightly distal to ib; est in the middle of fixed finger; et near sub-distal of fixed finger; est closer to et than to esb; it slightly distal to est and proximal to et; ist situated proximal to est and distal to isb; it closer to ist than to fingertip; distance between est and esb farther to that of ist and isb; distance between it and fingertip shorter to distance between ist and isb; b and sb situated at base of movable finger on retrolateral face; t in the middle of movable finger and at same level as est; sb closer to b than to st; st midway between t and sb (Fig.
|
||
<figureCitation id="9475B5F76B0A7D4C908C40454F73CAA0" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12M</figureCitation>
|
||
). Venom apparatus only present in fixed chelal finger, venom ducts curved and short, terminating in inflated nodus ramosus between et and est, closer to et. Both chelal fingers with a row of acute teeth, spaced contiguously along the margin, slightly rounded proximally: fixed chelal finger with 23 or 25 teeth; movable chelal finger with 32 teeth (nearly as large as teeth on fixed chelal finger); without accessory teeth (Fig.
|
||
<figureCitation id="073DD5AF33C6702B1C5E9B9B88D48CA8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12M</figureCitation>
|
||
). Femur without long tactile setae. Movable chelal finger slightly curved in lateral view (Figs
|
||
<figureCitation id="06BB67E10FFF35534B5F5347D70BA48D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Catatemnus scaber sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295">11B</figureCitation>
|
||
,
|
||
<figureCitation id="BBAFD081C52304CC0D2DEBEA063DE601" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12M</figureCitation>
|
||
).
|
||
<emphasis id="63284DC54B710295F567DC04174542E1" bold="true" italics="true" pageId="0" pageNumber="295">Opisthosoma</emphasis>
|
||
: typical, all setae long and acuminate, setal bases distinct larger; pleural membrane longitudinally striate, without setae. Tergites I-III and XI undivided and other tergites incompletely divided, tergites I-III each with nine setae, uniseriate; other tergites with 13 setae, biseriate; tergites X and XI each with a pair of tactile setae. Sternites completely divided, each half sternites with seven or eight setae; sternites X and XI each with two pairs of long tactile setae. Anus (tergite XII and sternite XII) without raised rim. Anterior genital operculum with 18 setae. Male genitalia (Figs
|
||
<figureCitation id="9C19097F3B19B209FFB6FA35DE2E9021" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Catatemnus scaber sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295">11E</figureCitation>
|
||
,
|
||
<figureCitation id="00B8B46B9BB4E557A05D6AFB4349AEE7" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12F, L</figureCitation>
|
||
): the distal part (l) of lateral apodemes with a distinctive inner ridge curved into semicircle; the hooked branch (br) bowed distally and terminated in a plate-like tip; the proximal part with a distinct dark sclerotized bar (c); the longitudinal fold of medial diverticula (d) with a projection midway along its length, forming a lobe bulge; the ejaculatory canal atrium (e) not well-developed, curved distally; the lateral rods (f) long and diverging proximally; the tip of dorsal apodeme (g) completely joined; the ventral diverticulum (h) bilobed; genital atrium without genital setae.
|
||
<emphasis id="232796B4205E6D9CB297EC6877329A97" bold="true" italics="true" pageId="0" pageNumber="295">Legs</emphasis>
|
||
(Figs
|
||
<figureCitation id="9E0BB63A607CF1E7A4F765BDCF7A38EF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Catatemnus scaber sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295">11G-I</figureCitation>
|
||
,
|
||
<figureCitation id="8A90302110E0B0958FA7B3635C9E1B1A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12H, I</figureCitation>
|
||
): typical, fairly smooth, slightly stout; junction between femora and patellae I and II oblique. Femoropatella of leg IV 2.86-3.07
|
||
<normalizedToken id="493083A42600DC3AE40B98E20BFA5875" originalValue="×">x</normalizedToken>
|
||
longer than deep; tibia 3.47-3.71
|
||
<normalizedToken id="296E29626A412464EA5275306F43C748" originalValue="×">x</normalizedToken>
|
||
longer than deep; with basal tactile setae on tarsal segment: tarsus 4.09-4.18
|
||
<normalizedToken id="1D97E27A6749C8FA618EDDC42DA795CB" originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.11); subterminal tarsal setae arcuate and acute. Arolium slightly shorter than claws, not divided; claws smooth.
|
||
</paragraph>
|
||
<caption id="A4564261137F2F22210D918753E2E81A" doi="10.3897/zookeys.1168.100798.figure10" httpUri="https://binary.pensoft.net/fig/872309" pageId="0" pageNumber="295" start="Figure 10" startId="F10">
|
||
<paragraph id="D70048BB15C9E844238E121B999C4462" pageId="0" pageNumber="295">
|
||
<emphasis id="D1AECD80C326222A00DBCDD711C0F575" bold="true" pageId="0" pageNumber="295">Figure 10.</emphasis>
|
||
<taxonomicName id="2D997A80FA89B2E886D355CF431B304B" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus scaber" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="scaber">
|
||
<emphasis id="317E6B4B58D6D2DFC369E03EBD84010B" italics="true" pageId="0" pageNumber="295">Catatemnus scaber</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="1C1EA62DA2976D40DB70AE1BD98A4B06" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
holotype male, habitus (dorsal view)
|
||
<emphasis id="87A05EDFF01145F5A89AF4677C5F2CF9" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
paratype female, habitus (dorsal view). Scale bars: 2.00 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="771AB08858A34C1CB9D2D2BF7F039E43" doi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295" start="Figure 11" startId="F11">
|
||
<paragraph id="2852E10D9C0B718195AEB22F79263280" pageId="0" pageNumber="295">
|
||
<emphasis id="119424F0C281BAD2324990E0BD28182B" bold="true" pageId="0" pageNumber="295">Figure 11.</emphasis>
|
||
<taxonomicName id="DC24BD92472DA01B7DB7930CDD3A8186" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus scaber" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="scaber">
|
||
<emphasis id="9E038F76233248843E518823D5136DEB" italics="true" pageId="0" pageNumber="295">Catatemnus scaber</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis id="495B642D0DE4DBF473370ECABFBEB6B4" bold="true" pageId="0" pageNumber="295">A-E, G-I</emphasis>
|
||
), paratype female (
|
||
<emphasis id="CE4685F801E4E209632734026EFC575D" bold="true" pageId="0" pageNumber="295">F</emphasis>
|
||
)
|
||
<emphasis id="82137E20BF6D0375D3CB656ACEF3E039" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis id="D06A11006EF8B150B74EA7FCBC702961" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
left chelal fingers (lateral view)
|
||
<emphasis id="EA63D7C3C1FFBC8425EA4CD286A4AB36" bold="true" pageId="0" pageNumber="295">C</emphasis>
|
||
left chela (lateral view)
|
||
<emphasis id="685ABFDEF3F191C074803A888F00874A" bold="true" pageId="0" pageNumber="295">D</emphasis>
|
||
left pedipalp (dorsal view)
|
||
<emphasis id="E6D2059E0BED2F489E33A946A5B9FC8F" bold="true" pageId="0" pageNumber="295">E</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis id="3C7A55BD9B82C8DC4C35741CA14D2297" bold="true" pageId="0" pageNumber="295">F</emphasis>
|
||
female genital area (ventral view)
|
||
<emphasis id="F3687952BD01B6EE56FB0D8C59ABC4D9" bold="true" pageId="0" pageNumber="295">G</emphasis>
|
||
tarsus of left leg IV (lateral view)
|
||
<emphasis id="04184EAA220EDFDDF4843882E1DC9C2E" bold="true" pageId="0" pageNumber="295">H</emphasis>
|
||
left leg I (lateral view)
|
||
<emphasis id="6CC5D8D1999442FDE770E20A955BF20C" bold="true" pageId="0" pageNumber="295">I</emphasis>
|
||
left leg IV (lateral view). Scale bars: 0.50 mm (
|
||
<emphasis id="2428DECD9233AE8FEB08DB88506FEE34" bold="true" pageId="0" pageNumber="295">A, C, D</emphasis>
|
||
); 0.20 mm (
|
||
<emphasis id="12BCEAE0C4D1F2FE487FC04CF53A6CC6" bold="true" pageId="0" pageNumber="295">B, E-I</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="AD79CDB7CB1629750080080B51E52E21" doi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295" start="Figure 12" startId="F12">
|
||
<paragraph id="3203795960FA1701FA0704A9314B4000" pageId="0" pageNumber="295">
|
||
<emphasis id="E52D34313AB64F0C98E206A0D75DD119" bold="true" pageId="0" pageNumber="295">Figure 12.</emphasis>
|
||
<taxonomicName id="41189D41F174D4AA80C08FFA8648E84D" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus scaber" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="scaber">
|
||
<emphasis id="7E256CE64D809330CEA3F296621CBB21" italics="true" pageId="0" pageNumber="295">Catatemnus scaber</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis id="DF54F6D8E80B2948513B27444E615B45" bold="true" pageId="0" pageNumber="295">A-C, E, F, H, I, K-M</emphasis>
|
||
), paratype female (
|
||
<emphasis id="87800DE54540B70A4B1AB81BFC3947AA" bold="true" pageId="0" pageNumber="295">D, G, J</emphasis>
|
||
)
|
||
<emphasis id="5A20391095D34514D7E5E17ACBD4DCE3" bold="true" pageId="0" pageNumber="295">A</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis id="F0D4A56D10646B1EB2C2355E12E526AC" bold="true" pageId="0" pageNumber="295">B</emphasis>
|
||
left chelicera (dorsal view)
|
||
<emphasis id="61525CB33A1B7FEBFE558AB57F4514FC" bold="true" pageId="0" pageNumber="295">C</emphasis>
|
||
male galea
|
||
<emphasis id="EC44A1B46D27FFE6150796C4CD0A0433" bold="true" pageId="0" pageNumber="295">D</emphasis>
|
||
female galea
|
||
<emphasis id="B5C15A1F98937D2C393C67EE1463F0B1" bold="true" pageId="0" pageNumber="295">E</emphasis>
|
||
rallum
|
||
<emphasis id="8A35329AE2E2763125A2F5936AA22A71" bold="true" pageId="0" pageNumber="295">F</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis id="5EF60184FECC393BADCF03F2DC0A7AB7" bold="true" pageId="0" pageNumber="295">G</emphasis>
|
||
female genital area (ventral view)
|
||
<emphasis id="31FD6BDCAFFCAC6C97D2CE3116533306" bold="true" pageId="0" pageNumber="295">H</emphasis>
|
||
left leg I (lateral view)
|
||
<emphasis id="286AF72A33BD6B1F979043FF75F789CE" bold="true" pageId="0" pageNumber="295">I</emphasis>
|
||
left leg IV (lateral view)
|
||
<emphasis id="4FCA825AFFE22AC0DCE02B668AB4FEA0" bold="true" pageId="0" pageNumber="295">J</emphasis>
|
||
spermatheca
|
||
<emphasis id="325396F29069C270F5E022DA91E13CCE" bold="true" pageId="0" pageNumber="295">K</emphasis>
|
||
left pedipalp (dorsal view)
|
||
<emphasis id="8286230FCA9D8210DEF35283D77818B6" bold="true" pageId="0" pageNumber="295">L</emphasis>
|
||
male genital organ
|
||
<emphasis id="7EE8F409D2AFBE98FA50975668F9D146" bold="true" pageId="0" pageNumber="295">M</emphasis>
|
||
left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (
|
||
<emphasis id="52C5B8BE0F169AC2512615860781B44B" bold="true" pageId="0" pageNumber="295">I, K</emphasis>
|
||
); 0.25 mm (
|
||
<emphasis id="A52F69CC7B331C70F0B56B4ED0C7A1B2" bold="true" pageId="0" pageNumber="295">A, F-H, J, L, M</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis id="BBB1B54F58F390646DDD302FA8CCAED0" bold="true" pageId="0" pageNumber="295">B-E</emphasis>
|
||
). Abbreviations:
|
||
<emphasis id="125372E1C77B17229A06081508735841" bold="true" pageId="0" pageNumber="295">b</emphasis>
|
||
= basal;
|
||
<emphasis id="0F0233EAC31FF71B3C65550877504613" bold="true" pageId="0" pageNumber="295">sb</emphasis>
|
||
= sub-basal;
|
||
<emphasis id="8272BB8C991FF96F32F97D58A16AA330" bold="true" pageId="0" pageNumber="295">st</emphasis>
|
||
= sub-terminal;
|
||
<emphasis id="FCF488B7E7F5A7F1B6D2CF65ABB36E49" bold="true" pageId="0" pageNumber="295">t</emphasis>
|
||
= terminal;
|
||
<emphasis id="99FDA429DB2BBC3D9F6747792470F7BB" bold="true" pageId="0" pageNumber="295">ib</emphasis>
|
||
= interior basal;
|
||
<emphasis id="C9409362700C70F26925DF54AEEBF524" bold="true" pageId="0" pageNumber="295">isb</emphasis>
|
||
= interior sub-basal;
|
||
<emphasis id="C895A8E8369CAF8173709A57540B30A2" bold="true" pageId="0" pageNumber="295">ist</emphasis>
|
||
= interior sub-terminal;
|
||
<emphasis id="F620E94630AC461662B6347F33C2DF02" bold="true" pageId="0" pageNumber="295">it</emphasis>
|
||
= interior terminal;
|
||
<emphasis id="DE6863B527036A7AD366358C2D5E153F" bold="true" pageId="0" pageNumber="295">eb</emphasis>
|
||
= exterior basal;
|
||
<emphasis id="EE4E2C540ADFD00F92DB2D0CF73CAB2F" bold="true" pageId="0" pageNumber="295">esb</emphasis>
|
||
= exterior sub-basal;
|
||
<emphasis id="FD1B1D57C050803C63800726085E1631" bold="true" pageId="0" pageNumber="295">est</emphasis>
|
||
= exterior sub-terminal;
|
||
<emphasis id="771023A8FC3175DCE9DFBD64A8E46C3B" bold="true" pageId="0" pageNumber="295">et</emphasis>
|
||
= exterior terminal;
|
||
<emphasis id="CD3222C3B6BEB597C04621C97871CCA0" bold="true" pageId="0" pageNumber="295">a</emphasis>
|
||
= lateral apodeme;
|
||
<emphasis id="F33F6ADC120C3E35D73325FABBF7E164" bold="true" pageId="0" pageNumber="295">br</emphasis>
|
||
= hooked branch;
|
||
<emphasis id="20FD539F79E1B54B38CBCB4CCE6E090A" bold="true" pageId="0" pageNumber="295">c</emphasis>
|
||
= sclerotized bar;
|
||
<emphasis id="C05BCD0130B525D2FDE9DBF9227AFE12" bold="true" pageId="0" pageNumber="295">d</emphasis>
|
||
= longitudinal fold of medial diverticulum;
|
||
<emphasis id="6E0C9F0173D97F95C3A8A8C95D037EB1" bold="true" pageId="0" pageNumber="295">e</emphasis>
|
||
= ejaculatory canal atrium;
|
||
<emphasis id="369BA057DEFCC779386C49F633338C6C" bold="true" pageId="0" pageNumber="295">f</emphasis>
|
||
= lateral rods;
|
||
<emphasis id="C620635ABDAE120C67BB4D1FE07D5897" bold="true" pageId="0" pageNumber="295">g</emphasis>
|
||
= dorsal apodeme;
|
||
<emphasis id="1508B7B5877D72EA44864D853B7FD7FA" bold="true" pageId="0" pageNumber="295">h</emphasis>
|
||
= ventral diverticulum;
|
||
<emphasis id="38AAE69BD9CB04D591A6AEEBAAF74506" bold="true" pageId="0" pageNumber="295">l</emphasis>
|
||
= lateral lip of lateral apodeme.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6294BEA5E0920F9CB0A296D27E158D62" pageId="0" pageNumber="295">
|
||
<emphasis id="C5C45B87459A1A290F184AF275E8C4BE" bold="true" pageId="0" pageNumber="295">Adult females</emphasis>
|
||
(Figs
|
||
<figureCitation id="0D3E15356BE6684FE3534F209FA83C35" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Catatemnus scaber sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 2.00 mm." figureDoi="10.3897/zookeys.1168.100798.figure10" httpUri="https://binary.pensoft.net/fig/872309" pageId="0" pageNumber="295">10B</figureCitation>
|
||
,
|
||
<figureCitation id="4BB52942F34FFD32A8DAF801FA1CCAA6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Catatemnus scaber sp. nov., holotype male (A-E, G-I), paratype female (F) A carapace (dorsal view) B left chelal fingers (lateral view) C left chela (lateral view) D left pedipalp (dorsal view) E male genital area (ventral view) F female genital area (ventral view) G tarsus of left leg IV (lateral view) H left leg I (lateral view) I left leg IV (lateral view). Scale bars: 0.50 mm (A, C, D); 0.20 mm (B, E-I)." figureDoi="10.3897/zookeys.1168.100798.figure11" httpUri="https://binary.pensoft.net/fig/872310" pageId="0" pageNumber="295">11F</figureCitation>
|
||
,
|
||
<figureCitation id="D03480793EA7BC6B5623AB45C26C27CF" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12G, J</figureCitation>
|
||
): mostly same as the male, but a little larger in body size and paler; pedipalps paler, reddish brown, and paler from fingertip to trochanter; carapace 1.16-1.25
|
||
<normalizedToken id="B67821D178382534C45C4752E22F9BC5" originalValue="×">x</normalizedToken>
|
||
longer than broad.
|
||
<emphasis id="17F76A0CFCC9AD349499B140AF08F919" bold="true" italics="true" pageId="0" pageNumber="295">Chelicera</emphasis>
|
||
: hand with four setae, gs shorter than others; galea distinct relatively short, with ten branchlets (Fig.
|
||
<figureCitation id="04BFD63620A25218377ADBF7528D09C8" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Catatemnus scaber sp. nov., holotype male (A-C, E, F, H, I, K-M), paratype female (D, G, J) A carapace (dorsal view) B left chelicera (dorsal view) C male galea D female galea E rallum F male genital area (ventral view) G female genital area (ventral view) H left leg I (lateral view) I left leg IV (lateral view) J spermatheca K left pedipalp (dorsal view) L male genital organ M left chelal fingers (lateral view), with details of trichobothrial pattern. Scale bars: 0.50 mm (I, K); 0.25 mm (A, F-H, J, L, M); 0.10 mm (B-E). Abbreviations: b = basal; sb = sub-basal; st = sub-terminal; t = terminal; ib = interior basal; isb = interior sub-basal; ist = interior sub-terminal; it = interior terminal; eb = exterior basal; esb = exterior sub-basal; est = exterior sub-terminal; et = exterior terminal; a = lateral apodeme; br = hooked branch; c = sclerotized bar; d = longitudinal fold of medial diverticulum; e = ejaculatory canal atrium; f = lateral rods; g = dorsal apodeme; h = ventral diverticulum; l = lateral lip of lateral apodeme." figureDoi="10.3897/zookeys.1168.100798.figure12" httpUri="https://binary.pensoft.net/fig/872311" pageId="0" pageNumber="295">12D</figureCitation>
|
||
); serrula exterior with 20 or 21 lamellae; rallum composed of four blades, the distal one dentate anteriorly.
|
||
<emphasis id="6B907EA06693151010C3A6D2BD53FCA2" bold="true" italics="true" pageId="0" pageNumber="295">Pedipalp</emphasis>
|
||
: stout, trochanter 1.57-1.62, femur 2.44-2.50, patella 2.11-2.14, chela (with pedicel) 2.61, chela (without pedicel) 2.41-2.43, hand (with pedicel) 1.57-1.65, hand (without pedicel) 1.39-1.45
|
||
<normalizedToken id="2961035AD0671E91F8831C6327E49EEE" originalValue="×">x</normalizedToken>
|
||
longer than broad, movable chelal finger 0.60-0.65
|
||
<normalizedToken id="37A13F77CDBF61FCCF701CDDA07E9D62" originalValue="×">x</normalizedToken>
|
||
(0.69-0.73
|
||
<normalizedToken id="C41B9D2F0BAF735A7ADA57A3229E786B" originalValue="×">x</normalizedToken>
|
||
) longer than hand with pedicel (without pedicel) and 0.38-0.39
|
||
<normalizedToken id="53367092DB3866B982F796403A40B2A5" originalValue="×">x</normalizedToken>
|
||
(0.41-0.42
|
||
<normalizedToken id="459251D712D6EBCC63BF65C1183C99AE" originalValue="×">x</normalizedToken>
|
||
) longer than chela with pedicel (without pedicel).
|
||
<emphasis id="29B3CF515E97D1ED3D609B62A9A6DE9B" bold="true" italics="true" pageId="0" pageNumber="295">Opisthosoma</emphasis>
|
||
: genitalia simple, spermathecae provided with separated median cribriform plates; anterior genital operculum with ten setae.
|
||
<emphasis id="98FDB24900A75673602B46B54ED60A2B" bold="true" italics="true" pageId="0" pageNumber="295">Legs</emphasis>
|
||
: femoropatella of leg IV 2.79-2.89
|
||
<normalizedToken id="7A3680F1FF4D0D91571A6333080740D9" originalValue="×">x</normalizedToken>
|
||
longer than deep; tibia 3.33-3.47
|
||
<normalizedToken id="292458B09C8EA926FA0F1558850727AE" originalValue="×">x</normalizedToken>
|
||
longer than deep; with basal tactile setae on tarsal segment: tarsus 3.75-4.00
|
||
<normalizedToken id="338BCBBA244528476B34689A1327441C" originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.11).
|
||
</paragraph>
|
||
<paragraph id="9E261021D327284D1BC78CFA0EBC20A5" pageId="0" pageNumber="295">
|
||
<emphasis id="503D391507C406B05730BFB9D293DEC3" bold="true" pageId="0" pageNumber="295">Dimensions</emphasis>
|
||
(length/breadth or, in the case of the legs, length/depth in mm). Males (females in parentheses): body length 2.99-3.35 (4.11-4.64). Carapace 0.93-0.96/0.72-0.80 (0.95-1.00/0.80-0.82). Pedipalp: trochanter 0.47-0.52/0.29 (0.44-0.47/0.28-0.29), femur 0.76-0.82/0.31-0.32 (0.75-0.78/0.30-0.32), patella 0.80-0.83/0.38-0.40 (0.77-0.78/0.36-0.37), chela (with pedicel) 1.35-1.36/0.48-0.49 (1.33-1.41/0.51-0.54), chela (without pedicel) 1.23-1.25 (1.23-1.31), hand (with pedicel) 0.81-0.83 (0.84-0.85), hand (without pedicel) 0.69-0.72 (0.74-0.75), movable finger length 0.53-0.54 (0.50-0.55). Leg I: trochanter 0.17-0.19/0.14-0.16 (0.16-0.18/0.14-0.15), femur 0.28-0.29/0.20-0.21 (0.25-0.26/0.20), patella 0.42-0.43/0.18-0.19 (0.41/0.18), tibia 0.42-0.45/0.13 (0.42-0.43/0.12-0.13), tarsus 0.40-0.42/0.09 (0.35-0.38/0.10-0.11). Leg IV: trochanter 0.32-0.33/0.19-0.22 (0.34/0.21), femoropatella 0.80-0.83/0.27-0.28 (0.81/0.28-0.29), tibia 0.59-0.63/0.17 (0.59-0.60/0.17-0.18), tarsus 0.45-0.46/0.11 (0.44-0.45/0.11-0.12).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9B3EE455CEB83C92B32CBFCB7E933116" pageId="0" pageNumber="295" type="remarks">
|
||
<paragraph id="C4F160CF8108D5A1AF0105703070F296" pageId="0" pageNumber="295">Remarks.</paragraph>
|
||
<paragraph id="42EE0AB97EEC9DF8674EB05766BB9EB4" pageId="0" pageNumber="295">
|
||
<taxonomicName id="FAB9899BF39940D8D00FD57B795CBE60" authorityName="Hou & Zhao & Zhang" authorityYear="2023" class="Arachnida" family="Atemnidae" genus="Catatemnus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Catatemnus scaber" order="Pseudoscorpiones" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="scaber">
|
||
<emphasis id="7DD94FAC23EEBE06EE4443DF05522688" italics="true" pageId="0" pageNumber="295">Catatemnus scaber</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName id="6DA78C3623A20D7DCDEDADA607D87A5B" class="Arachnida" kingdom="Animalia" lsidName="C. birmanicus" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="birmanicus">
|
||
<emphasis id="5EA6E8FC241F0061E9E99FEF17794A1A" italics="true" pageId="0" pageNumber="295">C. birmanicus</emphasis>
|
||
</taxonomicName>
|
||
by the smaller body size (♂) (e.g., body length 2.99-3.35 mm vs 4.27 mm; palpal femur length 0.76-0.82 mm vs 1.09 mm; movable chelal finger length 0.53-0.54 mm vs 0.78 mm) and the trait of eyespots (with two distinct eyespots vs with two faint eyespots); from
|
||
<taxonomicName id="C14781CC3E30A5D91EFBE189FE24BC9B" class="Arachnida" kingdom="Animalia" lsidName="C. concavus" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="concavus">
|
||
<emphasis id="B4E6FBAB2D02B993C5549D524B623C8B" italics="true" pageId="0" pageNumber="295">C. concavus</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs with two faint eyespots), the stouter pedipalps (♀) (palpal femur 2.44-2.50
|
||
<normalizedToken id="DB0B24D66DCA7CEE390DD55110846DAA" originalValue="×">x</normalizedToken>
|
||
vs 2.29
|
||
<normalizedToken id="CDB32B699176B508F70E53CED05DD410" originalValue="×">x</normalizedToken>
|
||
longer than broad, length 0.75-0.78 mm vs 0.90 mm) and the traits of the three first tergites (regularly convex from the one side to the other vs with a descending lateral portion, perpendicular on the dorsal part); from
|
||
<taxonomicName id="440FCEDAD028C3C62BDC4BAC247A04FC" class="Arachnida" kingdom="Animalia" lsidName="C. monitor" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="monitor">
|
||
<emphasis id="066FC32BDB4E849D4EC2368A93037B76" italics="true" pageId="0" pageNumber="295">C. monitor</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs no eyespots visible) and the stouter pedipalps (e.g., palpal femur (♂) 2.38-2.65
|
||
<normalizedToken id="B8B55C19BFA4BE4B986584D471A540FE" originalValue="×">x</normalizedToken>
|
||
vs 2.25
|
||
<normalizedToken id="1FFB1CF841DC1B89BFDEE0650DD5620D" originalValue="×">x</normalizedToken>
|
||
, (♀) 2.44-2.50
|
||
<normalizedToken id="0D16B3A52AE85E270C65501C0EE83D02" originalValue="×">x</normalizedToken>
|
||
vs 2.27
|
||
<normalizedToken id="44334B4750D82F790A0A5565B59A3DF7" originalValue="×">x</normalizedToken>
|
||
longer than broad, length (♂) 0.76-0.82 mm vs 1.01 mm, (♀) 0.75-0.78 mm vs 0.95 mm); from
|
||
<taxonomicName id="30E136FEFF76E9F0FFF912CB18CC9996" class="Arachnida" kingdom="Animalia" lsidName="C. nicobarensis" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="nicobarensis">
|
||
<emphasis id="DB9DF8C6890E74CBA38216DBA74A0FBE" italics="true" pageId="0" pageNumber="295">C. nicobarensis</emphasis>
|
||
</taxonomicName>
|
||
by the trait of eyespots (with two distinct eyespots vs with two indistinct eyespots) and the traits of the palpal trochanter (anterior surface granular vs smooth or almost smooth); from
|
||
<taxonomicName id="9B385A405F95AB3A86CC71BFA96EA9B4" class="Arachnida" kingdom="Animalia" lsidName="C. thorelli" order="Pseudoscorpionida" pageId="0" pageNumber="295" phylum="Arthropoda" rank="species" species="thorelli">
|
||
<emphasis id="5F4259A45BDD73CB9D0F36F278867345" italics="true" pageId="0" pageNumber="295">C. thorelli</emphasis>
|
||
</taxonomicName>
|
||
by the traits of pedipalps (surface of all segments except chelal fingers granular vs surface of palps smooth; palpal femur 2.38-2.65
|
||
<normalizedToken id="1209875CCA3C9734D787BF531B54040B" originalValue="×">x</normalizedToken>
|
||
vs 2.40
|
||
<normalizedToken id="71F3F7AC4E0BF547C0ACFEF245642EA2" originalValue="×">x</normalizedToken>
|
||
longer than broad, length 0.76-0.82 mm vs 1.15 mm; movable finger teeth length 0.53-0.54 mm vs 0.70 mm) (
|
||
<bibRefCitation id="DF9BF9D79C2CB4C62F18169177C1DBA8" DOI="https://doi.org/10.5962/bhl.title.10400" author="Thorell, T" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="295" pagination="521 - 729" refId="B9" refString="Thorell, T, 1889. Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885-1887. Annali del Museo Civico di Storia Naturale di Genova 7 (2): 521 - 729, DOI: https://doi.org/10.5962/bhl.title.10400" title="Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885 - 1887." url="https://doi.org/10.5962/bhl.title.10400" volume="7" year="1889">Thorell 1889</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5C5E9F5C2F0BA70FDA36BC6928AE75D7" author="Balzan, L" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="295" pagination="497 - 512" refId="B1" refString="Balzan, L, 1891. Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones). Annales de la Societe Entomologique de France 60: 497 - 512" title="Voyage de M. E. Simon au Venezuela (Decembre 1887 - Avril 1888). Arachnides. Chernetes (Pseudoscorpiones)." volume="60" year="1891">Balzan 1891</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="47F9F3B8C54FBB7874B3CCFD0D3A593F" author="With, CJ" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" pageId="0" pageNumber="295" refId="B10" refString="With, CJ, 1906. The Danish expedition to Siam 1899-1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7)3: 1-214." title="The Danish expedition to Siam 1899 - 1900. III. Chelonethi. An account of the Indian false-scorpions together with studies on the anatomy and classification of the order. Oversigt over det Konigelige Danske Videnskabernes Selskabs Forhandlinger (7) 3: 1 - 214." year="1906">With 1906</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="0FE5965C9879BF2C8C9582B91FEDA58D" DOI="https://doi.org/10.1515/9783111385402" author="Beier, M" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="295" refId="B2" refString="Beier, M, 1932. Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: [i-xxi,] 1-294. https://doi.org/10.1515/9783111385402" title="Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich 58: [i-xxi,] 1 - 294." url="https://doi.org/10.1515/9783111385402" year="1932">Beier 1932</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B03EE465D584BC7F369C133D392A69F9" pageId="0" pageNumber="295" type="distribution">
|
||
<paragraph id="819D0B0E7DEEBAC121D0219C3982FCA8" pageId="0" pageNumber="295">Distribution.</paragraph>
|
||
<paragraph id="0C4E71D96D9D191AFF090FC089721449" pageId="0" pageNumber="295">Known only from the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="182E840FB756000D390AE5143A72FF09" pageId="0" pageNumber="295" type="etymology">
|
||
<paragraph id="ED0F88543FC47E6939F65BDFEF70BF77" pageId="0" pageNumber="295">Etymology.</paragraph>
|
||
<paragraph id="BB728459BD65E7E1833DAA06951EACF5" pageId="0" pageNumber="295">
|
||
The specific name is derived from the Latin word
|
||
<emphasis id="71D221FD09C1F47FAD51C50BDCB572CA" italics="true" pageId="0" pageNumber="295">scaber</emphasis>
|
||
(rough) and refers to the large number of granulations present on the surface of the pedipalp.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |