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<document id="B10AA81CB08175C9A82E28E3CB572A15" ID-CLB-Dataset="263151" ID-DOI="10.11646/phytotaxa.598.2.4" ID-GBIF-Dataset="abaef93d-6b13-4c7c-bb40-569fed741453" ID-ISSN="1179-3163" ID-Zenodo-Dep="7961089" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1684824552020" checkinUser="plazi" docAuthor="Trovó, Marcelo &amp; Echternacht, Lívia" docDate="2023" docId="03F8A34BFFD214614ACBFF5A3FD6996F" docLanguage="en" docName="phytotaxa.598.2.4.pdf" docOrigin="Phytotaxa 598 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.598.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Paepalanthus balansae Ruhland 1903" docType="treatment" docVersion="5" lastPageNumber="150" masterDocId="FFC1DB33FFD714644A43FFAE3F1D9F09" masterDocTitle="Notes on Paepalanthus (Eriocaulaceae) from central Brazil and Paraguay" masterLastPageNumber="156" masterPageNumber="145" pageNumber="150" updateTime="1699515683115" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B4336EFEDE39FCCB2D0A19A93F7625DF">Notes on Paepalanthus (Eriocaulaceae) from central Brazil and Paraguay</mods:title>
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<mods:name id="EB2BD6615CE575D1EB06F3C7E9FDC0F9" type="personal">
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<mods:namePart id="C05C611DD244BBC54F6221454423A656">Trovó, Marcelo</mods:namePart>
<mods:affiliation id="4F2DDC1A52ACAF948EEF89A1BF15C6EC">Departamento de Botânica, Universidade Federal do Rio de Janeiro, Av. Carlos Chagas Filho 373, Cidade Universitária, 21941 - 590, Rio de Janeiro - RJ, Brazil.</mods:affiliation>
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<mods:namePart id="7054C9BC7B8CCC6F9FE0BDAB7560E612">Echternacht, Lívia</mods:namePart>
<mods:affiliation id="566F973E5D50B694CC71316373A3D7C3">Departamento de Biodiversidade, Evolução e Meio Ambiente, Universidade Federal de Ouro Preto, Campus Morro do Cruzeiro, ICEB III, Ouro Preto, MG 35400 - 000 Brazil.</mods:affiliation>
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<mods:title id="8BDCE0A2801885B88A5B91CB1B1DA841">Phytotaxa</mods:title>
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<mods:date id="785FDEDE7BC36F92F62DFD3BF838CAE1">2023</mods:date>
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<mods:number id="BE385AB54DF1746D392D8673D10B080F">598</mods:number>
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<treatment id="03F8A34BFFD214614ACBFF5A3FD6996F" ID-DOI="http://doi.org/10.5281/zenodo.7963116" ID-GBIF-Taxon="209628741" ID-Zenodo-Dep="7963116" LSID="urn:lsid:plazi:treatment:03F8A34BFFD214614ACBFF5A3FD6996F" httpUri="http://treatment.plazi.org/id/03F8A34BFFD214614ACBFF5A3FD6996F" lastPageNumber="150" pageId="5" pageNumber="150">
<subSubSection id="C34B41D6FFD214614ACBFF5A3D999E06" box="[136,644,244,271]" pageId="5" pageNumber="150" type="nomenclature">
<paragraph id="8BEE125DFFD214614ACBFF5A3D999E06" blockId="5.[136,1452,244,414]" box="[136,644,244,271]" pageId="5" pageNumber="150">
<taxonomicName id="4C5169DEFFD214614ACBFF5A3D9D9E06" ID-CoL="4BYGQ" ID-ENA="2776662" authority="Ruhland (1903: 151)" authorityName="Ruhland" authorityPageNumber="151" authorityYear="1903" box="[136,640,244,271]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="species" species="balansae">
<emphasis id="B925CE4FFFD214614ACBFF5A3E8E9E06" bold="true" box="[136,403,244,271]" italics="true" pageId="5" pageNumber="150">Paepalanthus balansae</emphasis>
<bibRefCitation id="EFC06FACFFD214614BD4FF5A3D9D9E06" author="Ruhland, W." box="[407,640,244,271]" pageId="5" pageNumber="150" pagination="1 - 294" refId="ref7200" refString="Ruhland, W. (1903) Eriocaulaceae. In: Engler, A. (ed.) Das Pflanzenreich. Regni vegetabilis conspectus 4 heft 30. Wilhelm Engelmann, Leipzig, pp. 1 - 294." type="book chapter" year="1903">Ruhland (1903: 151)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B41D6FFD2146148CBFF5A3D499E97" pageId="5" pageNumber="150" type="materials_examined">
<paragraph id="8BEE125DFFD2146148CBFF5A3D499E97" blockId="5.[136,1452,244,414]" pageId="5" pageNumber="150">
<location id="8E8E4486FFD2146148CBFF5A3DDD9E07" LSID="urn:lsid:plazi:treatment:03F8A34BFFD214614ACBFF5A3FD6996F:8E8E4486FFD2146148CBFF5A3DDD9E07" box="[648,704,244,270]" country="Paraguay" name="Type" pageId="5" pageNumber="150" stateProvince="Cordillera">Type</location>
:—
<collectingCountry id="F34652CDFFD2146148A3FF5A3C619E07" box="[736,892,244,270]" name="Paraguay" pageId="5" pageNumber="150">PARAGUAY</collectingCountry>
: [
<collectorName id="26A4778BFFD2146149C9FF5A3B089E07" box="[906,1045,244,270]" pageId="5" pageNumber="150">Pastoréo-mi</collectorName>
à
<collectorName id="26A4778BFFD214614E68FF5A3B759E06" box="[1067,1128,244,271]" pageId="5" pageNumber="150">la Est</collectorName>
de la
<collectingRegion id="4995DCBFFFD214614EE6FF5A3A0A9E07" box="[1189,1303,244,270]" country="Paraguay" name="Cordillera" pageId="5" pageNumber="150">Cordillère</collectingRegion>
<collectorName id="26A4778BFFD214614F5FFF5A3ABC9E07" box="[1308,1441,244,270]" pageId="5" pageNumber="150">de Villarica</collectorName>
],
<date id="FFEF349DFFD214614ACBFEB63E799E3B" box="[136,356,280,307]" pageId="5" pageNumber="150" value="1874-09-25">
<collectingDate id="EFABCD75FFD214614ACBFEB63E799E3B" box="[136,356,280,307]" pageId="5" pageNumber="150" value="1874-09-25">25 September 1874</collectingDate>
</date>
,
<emphasis id="B925CE4FFFD214614B2DFEB73D3C9E3B" box="[366,545,280,306]" italics="true" pageId="5" pageNumber="150">
<collectorName id="26A4778BFFD214614B2DFEB73EF49E3B" box="[366,489,280,306]" pageId="5" pageNumber="150">B. Balansa</collectorName>
568.
</emphasis>
<typeStatus id="54EAACFFFFD214614864FEB73D879E3A" box="[551,666,281,307]" pageId="5" pageNumber="150" type="lectotype">Lectotype</typeStatus>
, designated by
<bibRefCitation id="EFC06FACFFD214614906FEB63B5D9E3A" author="Moldenke, H. N." box="[837,1088,280,307]" pageId="5" pageNumber="150" pagination="339 - 368" refId="ref6940" refString="Moldenke, H. N. (1970) Additional notes on the Eriocaulaceae XXXIV. Phytologia 20: 339 - 368." type="journal article" year="1970">Moldenke (1970: 365)</bibRefCitation>
as “type” and corrected here, B! [B 10 0247854] fragment, isolectotypes BR! [BR0000008554048], F photo! [F0045750F] fragment, G! [G00004573], G! 2X[G00195500], K! [K000587311], LE! [LE00001137], P! [P00713450], P! [P00716650], P! [P00716651], NY! [NY00102796] fragment, S! [S-R-3866].
</paragraph>
</subSubSection>
<subSubSection id="C34B41D6FFD214614ACBFE7E3DE99EEF" box="[136,756,464,486]" pageId="5" pageNumber="150" type="reference_group">
<paragraph id="8BEE125DFFD214614ACBFE7E3DE99EEF" blockId="5.[136,1451,464,558]" box="[136,756,464,486]" pageId="5" pageNumber="150">
<emphasis id="B925CE4FFFD214614ACBFE7E3F889EEC" box="[136,149,464,485]" italics="true" pageId="5" pageNumber="150">=</emphasis>
<taxonomicName id="4C5169DEFFD214614AD6FE7E3DED9EEF" ID-CoL="5QFCZ" authority="Moldenke (1962: 162)" authorityName="Moldenke" authorityPageNumber="162" authorityYear="1962" box="[149,752,464,486]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="variety" species="balansae" variety="densiflorus">
<emphasis id="B925CE4FFFD214614AD6FE7E3E649EEC" box="[149,377,464,485]" italics="true" pageId="5" pageNumber="150">Paepalanthus balansae</emphasis>
var.
<emphasis id="B925CE4FFFD214614BE5FE7E3D0C9EEC" box="[422,529,464,485]" italics="true" pageId="5" pageNumber="150">densiflorus</emphasis>
<bibRefCitation id="EFC06FACFFD214614854FE7E3DED9EEF" author="Moldenke, H. N." box="[535,752,464,486]" pageId="5" pageNumber="150" pagination="159 - 164" refId="ref6916" refString="Moldenke, H. N. (1962) Notes on new and noteworthy plants XXX. Phytologia 8: 159 - 164." type="journal article" year="1962">Moldenke (1962: 162)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B41D6FFD2146148BAFE7E3A4E9D00" pageId="5" pageNumber="150" type="materials_examined">
<paragraph id="8BEE125DFFD2146148BAFE7E3A4E9D00" blockId="5.[136,1451,464,558]" pageId="5" pageNumber="150">
<location id="8E8E4486FFD2146148BAFE7E3C369EEC" LSID="urn:lsid:plazi:treatment:03F8A34BFFD214614ACBFF5A3FD6996F:8E8E4486FFD2146148BAFE7E3C369EEC" box="[761,811,464,485]" country="Brazil" name="Type" pageId="5" pageNumber="150" stateProvince="Parana">Type</location>
:—
<collectingCountry id="F34652CDFFD214614904FE7E3CBC9EEC" box="[839,929,464,485]" name="Brazil" pageId="5" pageNumber="150">BRAZIL</collectingCountry>
.
<collectingRegion id="4995DCBFFFD2146149EAFE7E3CF39EEC" box="[937,1006,464,485]" country="Brazil" name="Parana" pageId="5" pageNumber="150">Paraná</collectingRegion>
: [
<collectorName id="26A4778BFFD2146149BFFE7E3BA09EEF" box="[1020,1213,464,486]" pageId="5" pageNumber="150">Serra dos Dourados</collectorName>
,
<collectingRegion id="4995DCBFFFD214614E85FE7E3BFB9EEC" box="[1222,1254,464,485]" country="Brazil" name="Para" pageId="5" pageNumber="150">PR</collectingRegion>
. Cerradinho. Campo úmido],
<date id="FFEF349DFFD214614B4CFE5A3E899D03" box="[271,404,500,522]" pageId="5" pageNumber="150" value="1958-10">
<collectingDate id="EFABCD75FFD214614B4CFE5A3E899D03" box="[271,404,500,522]" pageId="5" pageNumber="150" value="1958-10">October 1958</collectingDate>
</date>
,
<emphasis id="B925CE4FFFD214614BDDFE5A3D959D00" box="[414,648,500,521]" italics="true" pageId="5" pageNumber="150">
R.
<taxonomicName id="4C5169DEFFD214614BFAFE5A3EE89D00" authority="R. Lange" authorityName="Schioedte &amp; Meinert" authorityYear="1881" box="[441,501,500,521]" class="Malacostraca" family="Cymothoidae" genus="Braga" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">Braga</taxonomicName>
&amp;
<collectorName id="26A4778BFFD214614850FE5A3D779D00" box="[531,618,500,521]" pageId="5" pageNumber="150">R. Lange</collectorName>
92
</emphasis>
.
<typeStatus id="54EAACFFFFD2146148D7FE5A3DF09D03" box="[660,749,500,522]" pageId="5" pageNumber="150" type="holotype">Holotype</typeStatus>
LL! [LL00374665], isotype UPCB! [UPCB0002273].
<emphasis id="B925CE4FFFD214614EBCFE5A3A4E9D00" box="[1279,1363,500,521]" italics="true" pageId="5" pageNumber="150">syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34B41D6FFD214614ACBFDBA3CD19D27" box="[136,972,532,558]" pageId="5" pageNumber="150" type="reference_group">
<paragraph id="8BEE125DFFD214614ACBFDBA3CD19D27" blockId="5.[136,1451,464,558]" box="[136,972,532,558]" pageId="5" pageNumber="150">
<emphasis id="B925CE4FFFD214614ACBFDBA3F8A9D27" box="[136,151,532,558]" italics="true" pageId="5" pageNumber="150">=</emphasis>
<taxonomicName id="4C5169DEFFD214614AD4FDBA3C459D27" ID-CoL="4BYGQ" authority="Ruhland ex Moldenke." authorityName="Moldenke." box="[151,856,532,558]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="variety" species="scholiophyllus" variety="balansae">
<emphasis id="B925CE4FFFD214614AD4FDBA3EC39D27" box="[151,478,532,558]" italics="true" pageId="5" pageNumber="150">Paepalanthus scholiophyllus</emphasis>
var.
<emphasis id="B925CE4FFFD21461484CFDBA3D699D27" box="[527,628,532,558]" italics="true" pageId="5" pageNumber="150">balansae</emphasis>
Ruhland ex Moldenke.
</taxonomicName>
<emphasis id="B925CE4FFFD21461491FFDBB3CD19D27" box="[860,972,532,558]" italics="true" pageId="5" pageNumber="150">nom. nud.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34B41D6FFD214614ACBFDF23FD6996F" pageId="5" pageNumber="150" type="discussion">
<paragraph id="8BEE125DFFD214614ACBFDF23A309C7B" blockId="5.[136,1452,604,1638]" pageId="5" pageNumber="150">
<emphasis id="B925CE4FFFD214614ACBFDF23FCF9D7F" bold="true" box="[136,210,604,630]" pageId="5" pageNumber="150">
<emphasis id="B925CE4FFFD214614ACBFDF23FD49D7F" bold="true" box="[136,201,604,630]" italics="true" pageId="5" pageNumber="150">Notes</emphasis>
:
</emphasis>
—
<bibRefCitation id="EFC06FACFFD214614AACFDF23EB99D7F" author="Ruhland, W." box="[239,420,604,630]" pageId="5" pageNumber="150" pagination="1 - 294" refId="ref7200" refString="Ruhland, W. (1903) Eriocaulaceae. In: Engler, A. (ed.) Das Pflanzenreich. Regni vegetabilis conspectus 4 heft 30. Wilhelm Engelmann, Leipzig, pp. 1 - 294." type="book chapter" year="1903">Ruhland (1903)</bibRefCitation>
described
<taxonomicName id="4C5169DEFFD21461486BFDF23C2E9D7F" authorityName="Ruhland" authorityYear="1903" box="[552,819,604,630]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="species" species="balansae">
<emphasis id="B925CE4FFFD21461486BFDF23C2E9D7F" box="[552,819,604,630]" italics="true" pageId="5" pageNumber="150">Paepalanthus balansae</emphasis>
</taxonomicName>
based on the specimens
<emphasis id="B925CE4FFFD214614E21FDF33A059D7F" box="[1122,1304,604,630]" italics="true" pageId="5" pageNumber="150">B. Balansa 568</emphasis>
collected in
<collectingCountry id="F34652CDFFD214614ACBFD2F3FED9D92" box="[136,240,641,667]" name="Paraguay" pageId="5" pageNumber="150">Paraguay</collectingCountry>
and distributed trough many European and North American herbaria.
<bibRefCitation id="EFC06FACFFD214614E71FD2E3BE79D93" author="Moldenke, H. N." box="[1074,1274,640,666]" pageId="5" pageNumber="150" pagination="339 - 368" refId="ref6940" refString="Moldenke, H. N. (1970) Additional notes on the Eriocaulaceae XXXIV. Phytologia 20: 339 - 368." type="journal article" year="1970">Moldenke (1970)</bibRefCitation>
stated that the specimen at B is the type and the specimens in other herbaria are isotypes. Although Wilhelm Ruhland annotated on the exsiccate labels that the specimen in BR is a type specimen and the specimen at B is a fragment of the specimen from BR,
<bibRefCitation id="EFC06FACFFD214614AB8FD423ECC9C0F" author="Moldenke, H. N." box="[251,465,748,774]" pageId="5" pageNumber="150" pagination="339 - 368" refId="ref6940" refString="Moldenke, H. N. (1970) Additional notes on the Eriocaulaceae XXXIV. Phytologia 20: 339 - 368." type="journal article" year="1970">Moldenke’s (1970)</bibRefCitation>
designation is not in serious conflict with the published protologue, as the specimen at B has enough material to enable us to distinguish the species, and the designation satisfies the ICN articles 7.10, 7.11, 9.10, and 9.23 (
<bibRefCitation id="EFC06FACFFD214614B74FC9A3D099C47" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. H. &amp; Li, D. Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[311,532,820,847]" pageId="5" pageNumber="150" refId="ref7648" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. H., Li, D. Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glash ¸ tten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B925CE4FFFD214614BDBFC9B3ED49C47" box="[408,457,820,846]" italics="true" pageId="5" pageNumber="150">et al</emphasis>
. 2018
</bibRefCitation>
). Therefore, although it is not the best option, the specimen at B should be treated as the
<typeStatus id="54EAACFFFFD214614A93FCF63E2A9C7B" box="[208,311,856,882]" pageId="5" pageNumber="150" type="lectotype">lectotype</typeStatus>
(
<bibRefCitation id="EFC06FACFFD214614B06FCF63EFB9C7B" author="McNeill, J." box="[325,486,856,882]" pageId="5" pageNumber="150" pagination="1112 - 1113" refId="ref6860" refString="McNeill, J. (2014) Holotype specimens and type citations: General issues. Taxon 63: 1112 - 1113. https: // doi. org / 10.12705 / 635.7" type="journal article" year="2014">McNeill 2014</bibRefCitation>
,
<bibRefCitation id="EFC06FACFFD214614BB2FCF63DA99C7A" author="Prado, J. &amp; Hirai, R. Y. &amp; Moran, R. C." box="[497,692,856,883]" pageId="5" pageNumber="150" pagination="651" refId="ref7110" refString="Prado, J., Hirai, R. Y. &amp; Moran, R. C. (2015) Proposals concerning inadvertent lectotypifications (and neotypifications). Taxon 64: 651. https: // doi. org / 10.12705 / 643.29" type="journal article" year="2015">
Prado
<emphasis id="B925CE4FFFD21461487AFCF73D6C9C7B" box="[569,625,856,882]" italics="true" pageId="5" pageNumber="150">et al.</emphasis>
2015
</bibRefCitation>
), and
<bibRefCitation id="EFC06FACFFD2146148B4FCF63CA79C7B" author="Moldenke, H. N." box="[759,954,856,882]" pageId="5" pageNumber="150" pagination="339 - 368" refId="ref6940" refString="Moldenke, H. N. (1970) Additional notes on the Eriocaulaceae XXXIV. Phytologia 20: 339 - 368." type="journal article" year="1970">Moldenke (1970)</bibRefCitation>
“type” citation is here corrected.
</paragraph>
<paragraph id="8BEE125DFFD214614AFEFCD23D2D9BD2" blockId="5.[136,1452,604,1638]" pageId="5" pageNumber="150">
<bibRefCitation id="EFC06FACFFD214614AFEFCD23E9B9C9F" author="Moldenke, H. N." box="[189,390,892,918]" pageId="5" pageNumber="150" pagination="159 - 164" refId="ref6916" refString="Moldenke, H. N. (1962) Notes on new and noteworthy plants XXX. Phytologia 8: 159 - 164." type="journal article" year="1962">Moldenke (1962)</bibRefCitation>
described
<taxonomicName id="4C5169DEFFD214614849FCD23CC99C9F" authorityName="Moldenke" authorityYear="1962" box="[522,980,892,919]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="variety" species="balansae" variety="densiflorus">
<emphasis id="B925CE4FFFD214614849FCD23C0B9C9F" box="[522,790,892,918]" italics="true" pageId="5" pageNumber="150">Paepalanthus balansae</emphasis>
var.
<emphasis id="B925CE4FFFD21461491BFCD23CC99C9F" box="[856,980,892,918]" italics="true" pageId="5" pageNumber="150">densiflorus</emphasis>
</taxonomicName>
based on the specimen
<emphasis id="B925CE4FFFD214614EB9FCD33FE89CB3" italics="true" pageId="5" pageNumber="150">
R.
<taxonomicName id="4C5169DEFFD214614F5CFCD33A789C9F" authority="R. Lange" authorityName="Schioedte &amp; Meinert" authorityYear="1881" box="[1311,1381,893,918]" class="Malacostraca" family="Cymothoidae" genus="Braga" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">Braga</taxonomicName>
&amp;
<collectorName id="26A4778BFFD214614FD0FCD33FD39CB3" pageId="5" pageNumber="150">R. Lange</collectorName>
92
</emphasis>
collected in Paraná (Brazil) and said to be deposited in his personal herbarium in Yonkers,
<collectingRegion id="4995DCBFFFD214614F6CFC0F3AB49CB3" box="[1327,1449,928,955]" country="United States of America" name="New York" pageId="5" pageNumber="150">New York</collectingRegion>
. The only specimen matching the collection number, date, and place is the specimen now housed at LL, and it also bears the typical Moldenke “type” label. There are however specimens deposited in NY [NY00501296] and in US [
<collectingCountry id="F34652CDFFD214614AD2FBA23FA99B2E" box="[145,180,1036,1063]" name="United States of America" pageId="5" pageNumber="150">US</collectingCountry>
01932715] holding labels with the same collector number, but with a posterior date and from different collection sites. We consider thus that the only specimen fully matching the protolog is the specimen at LL with a duplicate found at UPCB. These specimens should be treated as the
<typeStatus id="54EAACFFFFD214614886FBFA3C3A9B67" box="[709,807,1108,1134]" pageId="5" pageNumber="150" type="holotype">holotype</typeStatus>
and isotype of the species, respectively.
<bibRefCitation id="EFC06FACFFD214614EAAFBFA3AB69B67" author="Moldenke, H. N." box="[1257,1451,1108,1134]" pageId="5" pageNumber="150" pagination="159 - 164" refId="ref6916" refString="Moldenke, H. N. (1962) Notes on new and noteworthy plants XXX. Phytologia 8: 159 - 164." type="journal article" year="1962">Moldenke (1962)</bibRefCitation>
distinguished
<taxonomicName id="4C5169DEFFD214614B65FBD73D439B9B" authorityName="Moldenke" authorityYear="1962" box="[294,606,1144,1171]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="variety" species="balansae" variety="densiflorus">
<emphasis id="B925CE4FFFD214614B65FBD73EB59B9B" box="[294,424,1144,1170]" italics="true" pageId="5" pageNumber="150">P. balansae</emphasis>
var.
<emphasis id="B925CE4FFFD214614BA1FBD63D439B9B" box="[482,606,1144,1170]" italics="true" pageId="5" pageNumber="150">densiflorus</emphasis>
</taxonomicName>
based on the globose and many-flowered capitula of ca.
<quantity id="4CA9BFB8FFD214614EAEFBD63A009B9B" box="[1261,1309,1144,1170]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="150" unit="cm" value="1.0">1cm</quantity>
in diameter. Such circumscription falls within the range of morphological variation of the typical
<taxonomicName id="4C5169DEFFD214614E73FB323BB19BBF" box="[1072,1196,1180,1206]" pageId="5" pageNumber="150" rank="variety" variety="and">variety and</taxonomicName>
the heterotypic variety is thus proposed here as synonymous.
</paragraph>
<paragraph id="8BEE125DFFD214614AFEFB4A3D7C9ABB" blockId="5.[136,1452,604,1638]" pageId="5" pageNumber="150">
When detailing
<taxonomicName id="4C5169DEFFD214614B2DFB4A3C5E9BF7" authority=", Moldenke (1970)" authorityName=", Moldenke" authorityYear="1970" box="[366,835,1252,1279]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="species" species="balansae">
<emphasis id="B925CE4FFFD214614B2DFB4A3D6E9BF7" box="[366,627,1252,1278]" italics="true" pageId="5" pageNumber="150">Paepalanthus balansae</emphasis>
,
<bibRefCitation id="EFC06FACFFD2146148C3FB4A3C5E9BF7" author="Moldenke, H. N." box="[640,835,1252,1278]" pageId="5" pageNumber="150" pagination="339 - 368" refId="ref6940" refString="Moldenke, H. N. (1970) Additional notes on the Eriocaulaceae XXXIV. Phytologia 20: 339 - 368." type="journal article" year="1970">Moldenke (1970)</bibRefCitation>
</taxonomicName>
cited the name
<taxonomicName id="4C5169DEFFD2146149B3FB4B3E009A2B" authority="Ruhland ex Moldenke" authorityName="Moldenke" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="variety" species="scholiophyllus" variety="balansae">
<emphasis id="B925CE4FFFD2146149B3FB4B3BB19BF7" box="[1008,1196,1252,1278]" italics="true" pageId="5" pageNumber="150">P. scholiophyllus</emphasis>
var.
<emphasis id="B925CE4FFFD214614EA2FB4A3A5B9BF7" box="[1249,1350,1252,1278]" italics="true" pageId="5" pageNumber="150">balansae</emphasis>
Ruhland ex Moldenke
</taxonomicName>
as its synonym with additional references. The name indeed appears only in lists of species and in the label of a few specimens numbered as
<emphasis id="B925CE4FFFD214614803FA823DCA9A4F" box="[576,727,1324,1350]" italics="true" pageId="5" pageNumber="150">Hassler 9499</emphasis>
(e.g.
<collectingCountry id="F34652CDFFD21461495AFA823C209A4F" box="[793,829,1324,1350]" name="United States of America" pageId="5" pageNumber="150">US</collectingCountry>
[
<collectingCountry id="F34652CDFFD21461490DFA823C6F9A4F" box="[846,882,1324,1350]" name="United States of America" pageId="5" pageNumber="150">US</collectingCountry>
01932714]). As no proper description or diagnosis was published, the name is considered as
<emphasis id="B925CE4FFFD21461481AFAFE3DB89A63" box="[601,677,1360,1386]" italics="true" pageId="5" pageNumber="150">nudum</emphasis>
according to article 38 of the ICN (
<bibRefCitation id="EFC06FACFFD214614E71FAFE3A139A63" author="Turland, N. J. &amp; Wiersema, J. H. &amp; Barrie, F. R. &amp; Greuter, W. &amp; Hawksworth, D. L. &amp; Herendeen, P. S. &amp; Knapp, S. &amp; Kusber, W. H. &amp; Li, D. Z. &amp; Marhold, K. &amp; May, T. W. &amp; McNeill, J. &amp; Monro, A. M. &amp; Prado, J. &amp; Price, M. J. &amp; Smith, G. F." box="[1074,1294,1360,1386]" pageId="5" pageNumber="150" refId="ref7648" refString="Turland, N. J., Wiersema, J. H., Barrie, F. R., Greuter, W., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Kusber, W. H., Li, D. Z., Marhold, K., May, T. W., McNeill, J., Monro, A. M., Prado, J., Price, M. J. &amp; Smith, G. F. (Eds.) (2018) International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glash ¸ tten, 254 pp. https: // doi. org / 10.12705 / Code. 2018" type="book" year="2018">
Turland
<emphasis id="B925CE4FFFD214614ED0FAFF3BD69A63" box="[1171,1227,1360,1386]" italics="true" pageId="5" pageNumber="150">et al.</emphasis>
2018
</bibRefCitation>
). As indicated by
<bibRefCitation id="EFC06FACFFD214614AEEFADA3E699A87" author="Moldenke, H. N." box="[173,372,1396,1422]" pageId="5" pageNumber="150" pagination="339 - 368" refId="ref6940" refString="Moldenke, H. N. (1970) Additional notes on the Eriocaulaceae XXXIV. Phytologia 20: 339 - 368." type="journal article" year="1970">Moldenke (1970)</bibRefCitation>
, the specimen
<emphasis id="B925CE4FFFD214614863FADA3DA59A87" box="[544,696,1396,1422]" italics="true" pageId="5" pageNumber="150">Hassler 9499</emphasis>
is within the range of morphological variation of
<taxonomicName id="4C5169DEFFD214614EBBFADB3A679A87" authorityName=", Moldenke" authorityYear="1970" box="[1272,1402,1396,1422]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="5" pageNumber="150" phylum="Tracheophyta" rank="species" species="balansae">
<emphasis id="B925CE4FFFD214614EBBFADB3A679A87" box="[1272,1402,1396,1422]" italics="true" pageId="5" pageNumber="150">P. balansae</emphasis>
</taxonomicName>
and should be treated only at the species level.
</paragraph>
<paragraph id="8BEE125DFFD214614AFEFA123FD6996F" blockId="5.[136,1452,604,1638]" pageId="5" pageNumber="150">
<emphasis id="B925CE4FFFD214614AFEFA123E429ADF" bold="true" box="[189,351,1468,1495]" pageId="5" pageNumber="150">
<emphasis id="B925CE4FFFD214614AFEFA123E4B9ADE" bold="true" box="[189,342,1468,1495]" italics="true" pageId="5" pageNumber="150">Exsiccate list</emphasis>
:
</emphasis>
—
<emphasis id="B925CE4FFFD214614B38FA123EEE9ADF" box="[379,499,1468,1494]" italics="true" pageId="5" pageNumber="150">Arbo 8857</emphasis>
(HUEFS, MO);
<emphasis id="B925CE4FFFD2146148F1FA123C5D9ADF" box="[690,832,1468,1494]" italics="true" pageId="5" pageNumber="150">Balansa 568</emphasis>
(B, BR, F, G, LE, K, P, NY, S);
<emphasis id="B925CE4FFFD214614EF1FA123A419ADF" box="[1202,1372,1468,1494]" italics="true" pageId="5" pageNumber="150">Basualdo 4694</emphasis>
(MO);
<emphasis id="B925CE4FFFD214614ACBFA4E3FDD9AF3" box="[136,192,1504,1530]" italics="true" pageId="5" pageNumber="150">6447</emphasis>
(MO);
<emphasis id="B925CE4FFFD214614B57FA4F3E619AF3" box="[276,380,1504,1530]" italics="true" pageId="5" pageNumber="150">
<taxonomicName id="4C5169DEFFD214614B57FA4F3E479AF3" authorityName="Schioedte &amp; Meinert" authorityYear="1881" box="[276,346,1505,1530]" class="Malacostraca" family="Cymothoidae" genus="Braga" kingdom="Animalia" order="Isopoda" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">Braga</taxonomicName>
92
</emphasis>
(LL, UPCB);
<emphasis id="B925CE4FFFD214614858FA4E3D2A9AF3" box="[539,567,1504,1530]" italics="true" pageId="5" pageNumber="150">92</emphasis>
(NY,
<collectingCountry id="F34652CDFFD21461483AFA4E3DBC9AF3" box="[633,673,1504,1530]" name="United States of America" pageId="5" pageNumber="150">US</collectingCountry>
);
<emphasis id="B925CE4FFFD2146148F7FA4F3C529AF3" box="[692,847,1504,1530]" italics="true" pageId="5" pageNumber="150">Brunner 1510</emphasis>
(MO, SPF);
<emphasis id="B925CE4FFFD21461499CFA4E3B7F9AF3" box="[991,1122,1504,1530]" italics="true" pageId="5" pageNumber="150">
<taxonomicName id="4C5169DEFFD21461499CFA4E3B3E9AF3" box="[991,1059,1504,1530]" class="Insecta" family="Coleophoridae" genus="Casas" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="150" phylum="Arthropoda" rank="genus">Casas</taxonomicName>
3865
</emphasis>
(NY);
<emphasis id="B925CE4FFFD214614EF2FA4E3BF49AF3" box="[1201,1257,1504,1530]" italics="true" pageId="5" pageNumber="150">4006</emphasis>
(NY);
<emphasis id="B925CE4FFFD214614F7BFA4E3A6D9AF3" box="[1336,1392,1504,1530]" italics="true" pageId="5" pageNumber="150">7539</emphasis>
(NY, MO);
<emphasis id="B925CE4FFFD214614A8FF9AA3E559917" box="[204,328,1540,1566]" italics="true" pageId="5" pageNumber="150">Hahn 1774</emphasis>
(MO);
<emphasis id="B925CE4FFFD214614BD8F9AA3D2D9917" box="[411,560,1540,1566]" italics="true" pageId="5" pageNumber="150">Hassler 4448</emphasis>
(NY, MO, P,
<collectingCountry id="F34652CDFFD214614887F9AA3DF19917" box="[708,748,1540,1566]" name="United States of America" pageId="5" pageNumber="150">US</collectingCountry>
);
<emphasis id="B925CE4FFFD2146148BCF9AA3C2A9917" box="[767,823,1540,1566]" italics="true" pageId="5" pageNumber="150">9499</emphasis>
(NY, MO, P,
<collectingCountry id="F34652CDFFD214614988F9AA3CEE9917" box="[971,1011,1540,1566]" name="United States of America" pageId="5" pageNumber="150">US</collectingCountry>
);
<emphasis id="B925CE4FFFD214614E45F9AA3B579917" box="[1030,1098,1540,1566]" italics="true" pageId="5" pageNumber="150">11345</emphasis>
(P);
<emphasis id="B925CE4FFFD214614E3CF9AA3A4C9917" box="[1151,1361,1540,1566]" italics="true" pageId="5" pageNumber="150">Hatschbach 48461</emphasis>
(MBM, MO,
<collectingCountry id="F34652CDFFD214614A86F9863FF0994B" box="[197,237,1576,1602]" name="United States of America" pageId="5" pageNumber="150">US</collectingCountry>
);
<emphasis id="B925CE4FFFD214614B40F9863E54994B" box="[259,329,1576,1602]" italics="true" pageId="5" pageNumber="150">58708</emphasis>
(MBM);
<emphasis id="B925CE4FFFD214614BF9F9863D49994B" box="[442,596,1576,1602]" italics="true" pageId="5" pageNumber="150">Mereles 1051</emphasis>
(MO);
<emphasis id="B925CE4FFFD2146148EDF9863C78994B" box="[686,869,1576,1602]" italics="true" pageId="5" pageNumber="150">Pedersen 15045</emphasis>
(L, MBM, MO);
<emphasis id="B925CE4FFFD214614E77F9873BA8994B" box="[1076,1205,1576,1602]" italics="true" pageId="5" pageNumber="150">Rojas 4067</emphasis>
(MO);
<emphasis id="B925CE4FFFD214614F4CF9863AB6994B" box="[1295,1451,1576,1602]" italics="true" pageId="5" pageNumber="150">Schinini 5820</emphasis>
(NY).
</paragraph>
</subSubSection>
</treatment>
</document>
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