treatments-xml/data/03/F5/6D/03F56D1E3225F60D57A3009CFCEC959B.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.5690709" ID-GBIF-Dataset="726bd2a8-d57a-4c4b-b2c0-053713adb218" ID-GBIF-Taxon="133186450" ID-ISSN="0936-9902" ID-Zenodo-Dep="5690709" checkinTime="1504699318837" checkinUser="plazi" docAuthor="Maurice Kottelat" docDate="2017" docId="03F56D1E3225F60D57A3009CFCEC959B" docLanguage="en" docName="2017 Vanmanenia orcicampus.pdf" docOrigin="Ichthyol. Explor. Freshwaters 28 (1)" docStyle="DocumentStyle{}" docTitle="Vanmanenia orcicampus" docType="treatment" docVersion="6" lastPageId="4" lastPageNumber="91" masterDocId="FFCC15663224F60957170530FFBA9030" masterDocTitle="Vanmanenia orcicampus ı a new species of loach from the Plain of Jarsı Laos (Teleostei: Gastromyzontidae)" masterLastPageNumber="95" masterPageNumber="87" pageId="1" pageNumber="88" updateTime="1636793188949" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Vanmanenia orcicampus ı a new species of loach from the Plain of Jarsı Laos (Teleostei: Gastromyzontidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maurice Kottelat</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Ichthyol. Explor. Freshwaters</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-08-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>28</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>95</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.886193</mods:identifier>
<mods:identifier type="GBIF-Dataset">726bd2a8-d57a-4c4b-b2c0-053713adb218</mods:identifier>
<mods:identifier type="ISSN">0936-9902</mods:identifier>
<mods:identifier type="Zenodo-Dep">886193</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5690709" ID-GBIF-Taxon="133186450" ID-Zenodo-Dep="5690709" LSID="urn:lsid:plazi:treatment:03F56D1E3225F60D57A3009CFCEC959B" httpUri="http://treatment.plazi.org/id/03F56D1E3225F60D57A3009CFCEC959B" lastPageId="4" lastPageNumber="91" pageId="1" pageNumber="88">
<subSubSection pageId="1" pageNumber="88" type="nomenclature">
<paragraph blockId="1.[180,589,1452,1503]" pageId="1" pageNumber="88">
<heading bold="true" centered="true" fontSize="9" level="1" pageId="1" pageNumber="88" reason="4">
<emphasis bold="true" box="[180,589,1452,1474]" pageId="1" pageNumber="88">
<taxonomicName box="[180,445,1452,1473]" class="Actinopterygii" family="Balitoridae" genus="Vanmanenia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="88" phylum="Chordata" rank="species" species="orcicampus">
<emphasis bold="true" box="[180,445,1452,1473]" italics="true" pageId="1" pageNumber="88">Vanmanenia orcicampus</emphasis>
</taxonomicName>
ı new species
</emphasis>
(
<figureCitation box="[338,432,1480,1503]" captionStart-0="Fig. 1" captionStart-1="Fig. 2" captionStartId-0="1.[129,161,1120,1139]" captionStartId-1="2.[98,130,1299,1318]" captionTargetBox-0="[156,1163,181,1078]" captionTargetBox-1="[127,1134,177,1250]" captionTargetId-0="figure@1.[129,1187,156,1104]" captionTargetId-1="figure@2.[98,1156,156,729]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Vanmanenia orcicampus ı MHNG 2767.094 ı holotypeı 47.5 mm SL; Laos: Nam Ngum on Plain of Jars." captionText-1="Fig. 2. Vanmanenia orcicampus ı CMK 22732 ı paratypes; Laos: Nam Ngum on Plain of Jars; aı 20.6 mm SL (reversed); and bı 23.2 mm SL." httpUri-0="https://zenodo.org/record/886195/files/figure.png" httpUri-1="https://zenodo.org/record/886197/files/figure.png" pageId="1" pageNumber="88">Figs. 1-2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="materials_examined">
<paragraph blockId="1.[129,641,1539,1676]" pageId="1" pageNumber="88">
<materialsCitation ID-GBIF-Occurrence="1611915197" collectingDate="2012-03-02" collectionCode="MHNG" collectorName="M. Kottelati &amp; T. Phommavong &amp; B. L. Amathi" country="Laos" elevation="1138" latitude="19.579723" location="Ban Khangviangi" municipality="Nam Ngum" pageId="1" pageNumber="88" specimenCode="MHNG 2767.094" specimenCount="1" stateProvince="Xiangkhouang Province" typeStatus="holotype">
<emphasis bold="true" box="[129,238,1539,1560]" pageId="1" pageNumber="88">
<typeStatus box="[129,233,1539,1560]" pageId="1" pageNumber="88">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[249,429,1539,1561]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="88">MHNG 2767.094</specimenCode>
ı 
<quantity box="[444,534,1539,1561]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" pageId="1" pageNumber="88" unit="mm" value="47.5">47.5 mm</quantity>
SL; 
<collectingCountry box="[585,638,1539,1560]" name="Laos" pageId="1" pageNumber="88">Laos</collectingCountry>
: 
<collectingRegion box="[129,402,1568,1590]" country="Laos" name="Xiangkhouang" pageId="1" pageNumber="88">Xiangkhouang Province</collectingRegion>
: 
<collectingMunicipality box="[417,554,1568,1589]" pageId="1" pageNumber="88">Nam Ngum</collectingMunicipality>
at 
<location LSID="urn:lsid:plazi:treatment:03F56D1E3225F60D57A3009CFCEC959B:8E838AD33225F60855400310FEB39662" country="Laos" latitude="19.579723" municipality="Nam Ngum" name="Ban Khangviangi" pageId="1" pageNumber="88" stateProvince="Xiangkhouang Province">Ban Khangviangı</location>
<geoCoordinate box="[268,386,1597,1618]" direction="north" orientation="latitude" pageId="1" pageNumber="88" precision="15" value="19.579723">19°34'47&quot;N</geoCoordinate>
103°17'42&quot;Eı 
<elevation box="[525,637,1596,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="1" pageNumber="88" unit="m" value="1138.0">
<quantity box="[525,600,1597,1618]" metricMagnitude="3" metricUnit="m" metricValue="1.138" pageId="1" pageNumber="88" unit="m" value="1138.0">1138 m</quantity>
asl
</elevation>
; 
<collectorName box="[129,256,1625,1647]" pageId="1" pageNumber="88">M. Kottelatı</collectorName>
<collectorName box="[265,445,1625,1647]" pageId="1" pageNumber="88">T. Phommavong</collectorName>
&amp; 
<collectorName box="[481,620,1625,1647]" pageId="1" pageNumber="88">B. L. Amathı</collectorName>
<collectingDate pageId="1" pageNumber="88" value="2012-03-02">
<date pageId="1" pageNumber="88" value="2012-03-02">2 March 2012</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph blockId="1.[675,1187,1192,1244]" pageId="1" pageNumber="88">
<emphasis bold="true" box="[675,787,1194,1215]" pageId="1" pageNumber="88">Paratypes.</emphasis>
CMK 22732 
<date box="[922,945,1194,1216]" pageId="1" pageNumber="88">ı 5</date>
ı 
<quantity box="[958,1103,1192,1216]" metricMagnitude="-2" metricUnit="m" metricValue="2.19" metricValueMax="2.32" metricValueMin="2.06" pageId="1" pageNumber="88" unit="mm" value="21.9" valueMax="23.2" valueMin="20.6">20.6-23.2 mm</quantity>
SL; collected with the holotype.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="88" type="diagnosis">
<paragraph blockId="1.[675,1187,1280,1676]" pageId="1" pageNumber="88">
<emphasis bold="true" box="[675,798,1280,1302]" pageId="1" pageNumber="88">Diagnosis.</emphasis>
<taxonomicName box="[810,1069,1281,1302]" class="Actinopterygii" family="Balitoridae" genus="Vanmanenia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="88" phylum="Chordata" rank="species" species="orcicampus">
<emphasis box="[810,1069,1281,1302]" italics="true" pageId="1" pageNumber="88">Vanmanenia orcicampus</emphasis>
</taxonomicName>
is distinguished from the other species of the genus by its unique colour patternı made of a midlateral row of six blotchesı anterior two longitudinally elongatedı 3rd to 5th roundishı 6th faintı at posterior extremity of caudal peduncle; a middorsal row of three predorsal and three postdorsal saddlesı 3rd predorsal one extending backwards along dorsalfin base; and vermiculations between saddles and blotches and on lower half of flank. It is further distinguished from congeners in having a slender caudal peduncle whose depth is 1.5 times in its length and 2.5 times in body depth; and 12-14 branched pectoral-fin rays.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="91" pageId="2" pageNumber="89" type="description">
<paragraph blockId="2.[106,118,703,724]" lastBlockId="2.[644,1157,1398,1679]" pageId="2" pageNumber="89">
<emphasis bold="true" pageId="2" pageNumber="89">a b Description.</emphasis>
Based on 
<typeStatus box="[356,450,1398,1420]" pageId="2" pageNumber="89">holotype</typeStatus>
only. See Figures 1-2 for general appearance and 
<tableCitation box="[495,573,1427,1449]" captionStart="Table 1" captionStartId="3.[129,184,732,751]" captionTargetBox="[129,598,817,1669]" captionText="Table 1. Morphometric data of holotype of Vanmanenia orcicampus." httpUri="http://table.plazi.org/id/DF238C803227F60A579607ECFF5D9338" pageId="2" pageNumber="89" tableUuid="DF238C803227F60A579607ECFF5D9338">Table 1</tableCitation>
for morphometric data of 
<typeStatus box="[349,446,1456,1478]" pageId="2" pageNumber="89">holotype</typeStatus>
. A moderately elongate 
<taxonomicName authorityName="Fowler" authorityYear="1905" box="[198,395,1485,1507]" class="Actinopterygii" family="Gastromyzontidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="89" phylum="Chordata" rank="family">Gastromyzontidae</taxonomicName>
with arched dorsal profileı body depth regularly increasing up to slightly in front of dorsal-fin originı decreasing under dorsal-fin base. Behind dorsal finı body depth decreasing regularly to slightly in front of caudal-fin base. Dorsal profile continuous between head and body. Ventral profile slightly convex. Preventral part of belly and lower surface of head flat. Head slightly depressed; body slightly compressed. Interorbital area flat. In lateral viewı eye flushed with dorsal profile of head. Snout rounded. Depth of caudal peduncle 1.5 times in its lengthı tapering posteriorlyı and 2.5 times in body depth. Largest recorded size 
<quantity box="[1015,1105,1571,1593]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" pageId="2" pageNumber="89" unit="mm" value="47.5">47.5 mm</quantity>
SL.
</paragraph>
<caption httpUri="https://zenodo.org/record/886197/files/figure.png" pageId="2" pageNumber="89" targetBox="[127,1134,177,1250]" targetPageId="2">
<paragraph blockId="2.[98,1157,1299,1344]" pageId="2" pageNumber="89">
<emphasis bold="true" box="[98,155,1299,1318]" pageId="2" pageNumber="89">Fig. 2.</emphasis>
<taxonomicName box="[166,376,1300,1318]" class="Actinopterygii" family="Balitoridae" genus="Vanmanenia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="89" phylum="Chordata" rank="species" species="orcicampus">
<emphasis box="[166,376,1300,1318]" italics="true" pageId="2" pageNumber="89">
Vanmanenia 
<taxonomicName box="[279,376,1300,1318]" pageId="2" pageNumber="89" rank="species" species="orcicampus">orcicampus</taxonomicName>
</emphasis>
</taxonomicName>
ı CMK 22732ı paratypes; Laos: Nam Ngum on Plain of Jars; 
<emphasis bold="true" box="[925,941,1299,1318]" pageId="2" pageNumber="89">aı</emphasis>
20.6 mm SL (reversed); and 
<emphasis bold="true" box="[140,158,1325,1344]" pageId="2" pageNumber="89">bı</emphasis>
23.2 mm SL.
</paragraph>
</caption>
<paragraph blockId="2.[644,1157,1398,1679]" lastBlockId="3.[128,641,162,674]" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89">
Dorsal fin with 4 unbranched and 7 (2) or 7 
<emphasis bold="true" box="[1137,1141,1599,1611]" pageId="2" pageNumber="89">1</emphasis>
/ 
<emphasis bold="true" box="[1149,1156,1607,1619]" pageId="2" pageNumber="89">2</emphasis>
(4*) branched rays; distal margin almost straight; 1 
<taxonomicName box="[654,724,1657,1679]" pageId="2" pageNumber="89" rank="subSpecies" subSpecies="and">st and</taxonomicName>
2nd branched rays longest. Pectoral fin with 1 unbranched and 12 (2) or 13 (4*) branched rays (including small last rayı unbranched)ı roundedı reaching 
<emphasis bold="true" box="[342,346,219,231]" pageId="3" pageNumber="90">3</emphasis>
/ 
<emphasis bold="true" box="[355,362,228,240]" pageId="3" pageNumber="90">4</emphasis>
of distance to pelvic-fin base. No axillary pectoral lobe. Pelvic fin with 1 unbranched and 8 (4) or 9 (2*) branched rays (including small last rayı unbranched); reaching about ¾ of distance to anus; rounded; origin below base of branched dorsal-fin rays 2-3; axillary lobe presentı freeı extending on 1 
<taxonomicName box="[424,490,392,414]" pageId="3" pageNumber="90" rank="subSpecies" subSpecies="and">st and</taxonomicName>
2nd branched rays. Anus situated at about 
<emphasis bold="true" box="[416,420,421,433]" pageId="3" pageNumber="90">1</emphasis>
/ 
<emphasis bold="true" box="[428,435,429,441]" pageId="3" pageNumber="90">3</emphasis>
of distance between pelvic-fin base and anal-fin origin. Anal fin with 3 unbranched and 5 
<emphasis bold="true" box="[348,352,478,490]" pageId="3" pageNumber="90">1</emphasis>
/ 
<emphasis bold="true" box="[360,367,487,499]" pageId="3" pageNumber="90">2</emphasis>
(6*) branched rays; distal margin convex. Caudal fin with 7+ 7 (6*) branched rays; about 6 dorsal and about 4 ventral procurrent rays; forkedı lobes roundedı of equal length. Upper 4 and lower 4 principal caudal-fin rays adnate (without membranes) until about first branching point.
</paragraph>
<paragraph blockId="3.[675,1188,162,1451]" pageId="3" pageNumber="90">Body entirely covered by scalesı except on belly in front of anus. Most scales with an acuminate projection on posterior marginı tip often fleshyı sometimes appearing as ‘soft tubercles’. Lateral line completeı not extending on caudal-fin baseı with 79 pored scales.</paragraph>
<paragraph blockId="3.[675,1188,162,1451]" pageId="3" pageNumber="90">
Anterior nostril pierced in front side of a pointed flap-like tube. Posterior nostril adjacent to anterior one. Mouth archedı gape about 2 times wider than long (
<figureCitation box="[944,1003,421,443]" captionStart="Fig. 3" captionStartId="5.[129,161,463,482]" captionTargetBox="[129,1187,159,445]" captionTargetId="figure@5.[129,1187,159,445]" captionTargetPageId="5" captionText="Fig. 3. Vanmanenia orcicampus ı mouth. aı CMK 22732 ı 20.6 mm SL; bı CMK 22732 ı 22.1 mm SL; and cı MHNG 2767.094 ı holotypeı 47.5 mm SL." httpUri="https://zenodo.org/record/886199/files/figure.png" pageId="3" pageNumber="90">Fig. 3</figureCitation>
b). Two pairs of rostral barbels and one barbel at each corner of mouth. Rostral groove shallow between rostral barbelsı deeper from base of outer rostral barbel backwards; edge of rostral fold with three fleshy lobes between rostral barbels. Lips very finely papillatedı continuous around corner of mouth. Upper lip fleshyı appearing as three transverse layers: two fleshly layers and a narrow intermediate oneı smoothı harderı slightly protrudingı not reaching corner of mouth. Lower lip in three partsı each about of equal lengthı separated by small notches; median part thinı papillated along anterior edge; lateral parts thick and fleshyı with a small projection at midlength; postlabial groove restricted to behind lateral parts. Mouth structure of juvenile (
<figureCitation box="[796,855,882,904]" captionStart="Fig. 3" captionStartId="5.[129,161,463,482]" captionTargetBox="[129,1187,159,445]" captionTargetId="figure@5.[129,1187,159,445]" captionTargetPageId="5" captionText="Fig. 3. Vanmanenia orcicampus ı mouth. aı CMK 22732 ı 20.6 mm SL; bı CMK 22732 ı 22.1 mm SL; and cı MHNG 2767.094 ı holotypeı 47.5 mm SL." httpUri="https://zenodo.org/record/886199/files/figure.png" pageId="3" pageNumber="90">Fig. 3</figureCitation>
a- b) as adultı except that mouth is narrowerı lower jaw exposed. Notches between three parts of lower lip not very marked; lateral parts close togetherı more or less in contact mesiallyı where fleshier and with two short projections and in some individuals forming a ridge with contralateralı wide and thin towards maxillary barbel with a short projection along median edge. Gill opening extending downwards in front and below base of anterior pectoral-fin ray.
</paragraph>
<caption ID-Table-UUID="DF238C803227F60A579607ECFF5D9338" httpUri="http://table.plazi.org/id/DF238C803227F60A579607ECFF5D9338" pageId="3" pageNumber="90" targetBox="[129,598,817,1669]" targetIsTable="true" targetPageId="3">
<paragraph blockId="3.[129,641,731,776]" pageId="3" pageNumber="90">
<emphasis bold="true" box="[129,204,731,751]" pageId="3" pageNumber="90">Table 1.</emphasis>
Morphometric data of holotype of 
<taxonomicName class="Actinopterygii" family="Balitoridae" genus="Vanmanenia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="90" phylum="Chordata" rank="species" species="orcicampus">
<emphasis italics="true" pageId="3" pageNumber="90">
Vanmanenia 
<taxonomicName box="[129,227,758,776]" pageId="3" pageNumber="90" rank="species" species="orcicampus">orcicampus</taxonomicName>
</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="3" pageNumber="90">
<table box="[129,598,817,1669]" gridcols="2" gridrows="33" pageId="3" pageNumber="90">
<tr box="[129,598,817,836]" gridrow="0" pageId="3" pageNumber="90">
<th box="[129,487,817,836]" gridcol="0" gridrow="0" pageId="3" pageNumber="90">Standard length (mm)</th>
<th box="[550,598,817,836]" gridcol="1" gridrow="0" pageId="3" pageNumber="90">47.5</th>
</tr>
<tr box="[129,598,846,865]" gridrow="1" pageId="3" pageNumber="90">
<td box="[129,487,846,865]" gridcol="0" gridrow="1" pageId="3" pageNumber="90">Total length (mm)</td>
<td box="[550,598,846,865]" gridcol="1" gridrow="1" pageId="3" pageNumber="90">61.3</td>
</tr>
<tr box="[129,598,879,899]" gridrow="2" pageId="3" pageNumber="90" rowspan-1="1">
<td box="[129,487,879,899]" gridcol="0" gridrow="2" pageId="3" pageNumber="90">
<emphasis bold="true" box="[129,416,879,899]" pageId="3" pageNumber="90">In percent of standard length</emphasis>
</td>
</tr>
<tr box="[129,598,905,924]" gridrow="3" pageId="3" pageNumber="90">
<td box="[129,487,905,924]" gridcol="0" gridrow="3" pageId="3" pageNumber="90">Total length</td>
<td box="[550,598,905,924]" gridcol="1" gridrow="3" pageId="3" pageNumber="90">129.2</td>
</tr>
<tr box="[129,598,930,949]" gridrow="4" pageId="3" pageNumber="90">
<td box="[129,487,930,949]" gridcol="0" gridrow="4" pageId="3" pageNumber="90">Head length (dorsal)</td>
<td box="[550,598,930,949]" gridcol="1" gridrow="4" pageId="3" pageNumber="90">23.5</td>
</tr>
<tr box="[129,598,956,975]" gridrow="5" pageId="3" pageNumber="90">
<td box="[129,487,956,975]" gridcol="0" gridrow="5" pageId="3" pageNumber="90">Head length (lateral)</td>
<td box="[550,598,956,975]" gridcol="1" gridrow="5" pageId="3" pageNumber="90">23.3</td>
</tr>
<tr box="[129,598,981,1000]" gridrow="6" pageId="3" pageNumber="90">
<td box="[129,487,981,1000]" gridcol="0" gridrow="6" pageId="3" pageNumber="90">Predorsal length</td>
<td box="[550,598,981,1000]" gridcol="1" gridrow="6" pageId="3" pageNumber="90">49.0</td>
</tr>
<tr box="[129,598,1006,1026]" gridrow="7" pageId="3" pageNumber="90">
<td box="[129,487,1006,1026]" gridcol="0" gridrow="7" pageId="3" pageNumber="90">Prepelvic length</td>
<td box="[550,598,1006,1026]" gridcol="1" gridrow="7" pageId="3" pageNumber="90">50.9</td>
</tr>
<tr box="[129,598,1032,1051]" gridrow="8" pageId="3" pageNumber="90">
<td box="[129,487,1032,1051]" gridcol="0" gridrow="8" pageId="3" pageNumber="90">Pre-anus length</td>
<td box="[550,598,1032,1051]" gridcol="1" gridrow="8" pageId="3" pageNumber="90">61.8</td>
</tr>
<tr box="[129,598,1057,1077]" gridrow="9" pageId="3" pageNumber="90">
<td box="[129,487,1057,1077]" gridcol="0" gridrow="9" pageId="3" pageNumber="90">Pre-anal length</td>
<td box="[550,598,1057,1077]" gridcol="1" gridrow="9" pageId="3" pageNumber="90">79.1</td>
</tr>
<tr box="[129,598,1083,1102]" gridrow="10" pageId="3" pageNumber="90">
<td box="[129,487,1083,1102]" gridcol="0" gridrow="10" pageId="3" pageNumber="90">Head depth</td>
<td box="[550,598,1083,1102]" gridcol="1" gridrow="10" pageId="3" pageNumber="90">13.3</td>
</tr>
<tr box="[129,598,1108,1128]" gridrow="11" pageId="3" pageNumber="90">
<td box="[129,487,1108,1128]" gridcol="0" gridrow="11" pageId="3" pageNumber="90">Body depth at dorsal-fin origin</td>
<td box="[550,598,1108,1128]" gridcol="1" gridrow="11" pageId="3" pageNumber="90">20.6</td>
</tr>
<tr box="[129,598,1134,1153]" gridrow="12" pageId="3" pageNumber="90">
<td box="[129,487,1134,1153]" gridcol="0" gridrow="12" pageId="3" pageNumber="90">Depth of caudal peduncle</td>
<td box="[550,598,1134,1153]" gridcol="1" gridrow="12" pageId="3" pageNumber="90">8.3</td>
</tr>
<tr box="[129,598,1159,1178]" gridrow="13" pageId="3" pageNumber="90">
<td box="[129,487,1159,1178]" gridcol="0" gridrow="13" pageId="3" pageNumber="90">Length of caudal peduncle</td>
<td box="[550,598,1159,1178]" gridcol="1" gridrow="13" pageId="3" pageNumber="90">12.8</td>
</tr>
<tr box="[129,598,1184,1204]" gridrow="14" pageId="3" pageNumber="90">
<td box="[129,487,1184,1204]" gridcol="0" gridrow="14" pageId="3" pageNumber="90">Head width</td>
<td box="[550,598,1184,1204]" gridcol="1" gridrow="14" pageId="3" pageNumber="90">17.8</td>
</tr>
<tr box="[129,598,1210,1229]" gridrow="15" pageId="3" pageNumber="90">
<td box="[129,487,1210,1229]" gridcol="0" gridrow="15" pageId="3" pageNumber="90">Body width at dorsal-fin origin</td>
<td box="[550,598,1210,1229]" gridcol="1" gridrow="15" pageId="3" pageNumber="90">16.8</td>
</tr>
<tr box="[129,598,1235,1254]" gridrow="16" pageId="3" pageNumber="90">
<td box="[129,487,1235,1254]" gridcol="0" gridrow="16" pageId="3" pageNumber="90">Snout length</td>
<td box="[550,598,1235,1254]" gridcol="1" gridrow="16" pageId="3" pageNumber="90">11.5</td>
</tr>
<tr box="[129,598,1261,1280]" gridrow="17" pageId="3" pageNumber="90">
<td box="[129,487,1261,1280]" gridcol="0" gridrow="17" pageId="3" pageNumber="90">Eye diameter</td>
<td box="[550,598,1261,1280]" gridcol="1" gridrow="17" pageId="3" pageNumber="90">5.0</td>
</tr>
<tr box="[129,598,1286,1305]" gridrow="18" pageId="3" pageNumber="90">
<td box="[129,487,1286,1305]" gridcol="0" gridrow="18" pageId="3" pageNumber="90">Interorbital width</td>
<td box="[550,598,1286,1305]" gridcol="1" gridrow="18" pageId="3" pageNumber="90">7.8</td>
</tr>
<tr box="[129,598,1311,1330]" gridrow="19" pageId="3" pageNumber="90">
<td box="[129,487,1311,1330]" gridcol="0" gridrow="19" pageId="3" pageNumber="90">Mouth width</td>
<td box="[550,598,1311,1330]" gridcol="1" gridrow="19" pageId="3" pageNumber="90">4.5</td>
</tr>
<tr box="[129,598,1337,1356]" gridrow="20" pageId="3" pageNumber="90">
<td box="[129,487,1337,1356]" gridcol="0" gridrow="20" pageId="3" pageNumber="90">Length of dorsal fin</td>
<td box="[550,598,1337,1356]" gridcol="1" gridrow="20" pageId="3" pageNumber="90">23.0</td>
</tr>
<tr box="[129,598,1362,1382]" gridrow="21" pageId="3" pageNumber="90">
<td box="[129,487,1362,1382]" gridcol="0" gridrow="21" pageId="3" pageNumber="90">Length of upper caudal-fin lobe</td>
<td box="[550,598,1362,1382]" gridcol="1" gridrow="21" pageId="3" pageNumber="90">28.7</td>
</tr>
<tr box="[129,598,1388,1407]" gridrow="22" pageId="3" pageNumber="90">
<td box="[129,487,1388,1407]" gridcol="0" gridrow="22" pageId="3" pageNumber="90">Length of median caudal-fin rays</td>
<td box="[550,598,1388,1407]" gridcol="1" gridrow="22" pageId="3" pageNumber="90">23.5</td>
</tr>
<tr box="[129,598,1413,1432]" gridrow="23" pageId="3" pageNumber="90">
<td box="[129,487,1413,1432]" gridcol="0" gridrow="23" pageId="3" pageNumber="90">Length of lower caudal-fin lobe</td>
<td box="[550,598,1413,1432]" gridcol="1" gridrow="23" pageId="3" pageNumber="90">27.4</td>
</tr>
<tr box="[129,598,1438,1458]" gridrow="24" pageId="3" pageNumber="90">
<td box="[129,487,1438,1458]" gridcol="0" gridrow="24" pageId="3" pageNumber="90">Length of anal fin</td>
<td box="[550,598,1438,1458]" gridcol="1" gridrow="24" pageId="3" pageNumber="90">18.8</td>
</tr>
<tr box="[129,598,1464,1483]" gridrow="25" pageId="3" pageNumber="90">
<td box="[129,487,1464,1483]" gridcol="0" gridrow="25" pageId="3" pageNumber="90">Length of pelvic fin</td>
<td box="[550,598,1464,1483]" gridcol="1" gridrow="25" pageId="3" pageNumber="90">23.5</td>
</tr>
<tr box="[129,598,1489,1508]" gridrow="26" pageId="3" pageNumber="90">
<td box="[129,487,1489,1508]" gridcol="0" gridrow="26" pageId="3" pageNumber="90">Length of pectoral fin</td>
<td box="[550,598,1489,1508]" gridcol="1" gridrow="26" pageId="3" pageNumber="90">27.2</td>
</tr>
<tr box="[129,598,1515,1534]" gridrow="27" pageId="3" pageNumber="90">
<td box="[129,487,1515,1534]" gridcol="0" gridrow="27" pageId="3" pageNumber="90">Distance between pelvic-fin bases</td>
<td box="[550,598,1515,1534]" gridcol="1" gridrow="27" pageId="3" pageNumber="90">5.2</td>
</tr>
<tr box="[129,598,1548,1567]" gridrow="28" pageId="3" pageNumber="90" rowspan-1="1">
<td box="[129,487,1548,1567]" gridcol="0" gridrow="28" pageId="3" pageNumber="90">
<emphasis bold="true" box="[129,446,1548,1567]" pageId="3" pageNumber="90">In percent of lateral head length</emphasis>
</td>
</tr>
<tr box="[129,598,1573,1593]" gridrow="29" pageId="3" pageNumber="90">
<td box="[129,487,1573,1593]" gridcol="0" gridrow="29" pageId="3" pageNumber="90">Snout length</td>
<td box="[550,598,1573,1593]" gridcol="1" gridrow="29" pageId="3" pageNumber="90">49</td>
</tr>
<tr box="[129,598,1599,1618]" gridrow="30" pageId="3" pageNumber="90">
<td box="[129,487,1599,1618]" gridcol="0" gridrow="30" pageId="3" pageNumber="90">Eye diameter</td>
<td box="[550,598,1599,1618]" gridcol="1" gridrow="30" pageId="3" pageNumber="90">21</td>
</tr>
<tr box="[129,598,1624,1643]" gridrow="31" pageId="3" pageNumber="90">
<td box="[129,487,1624,1643]" gridcol="0" gridrow="31" pageId="3" pageNumber="90">Interorbital width</td>
<td box="[550,598,1624,1643]" gridcol="1" gridrow="31" pageId="3" pageNumber="90">34</td>
</tr>
<tr box="[129,598,1650,1669]" gridrow="32" pageId="3" pageNumber="90">
<td box="[129,487,1650,1669]" gridcol="0" gridrow="32" pageId="3" pageNumber="90">Mouth width</td>
<td box="[550,598,1650,1669]" gridcol="1" gridrow="32" pageId="3" pageNumber="90">49</td>
</tr>
</table>
</paragraph>
<paragraph blockId="3.[675,1188,162,1451]" pageId="3" pageNumber="90">
Tubercles on all surfaces of headı finer on snout and top of headı larger but smooth on ventral surfaceı large and densely set on opercle. Unculiferous pads (sensu Conway et al. 
<date box="[1088,1151,1256,1279]" pageId="3" pageNumber="90" value="2012-01">ı 2012</date>
): in pectoral finı well developed along anterior edge of unbranched ray and behind unbranched and first seven branched raysı hardly distinct on remaining branched rays; in pelvic finı well developed along anterior edge of unbranched ray and behind unbranched and first three branched rays.
</paragraph>
<paragraph blockId="3.[675,1188,1487,1682]" pageId="3" pageNumber="90">
<emphasis bold="true" box="[675,798,1487,1509]" pageId="3" pageNumber="90">Coloration.</emphasis>
After about one month in formalin. Adult: Head and body background colour pale yellowish 
<taxonomicName box="[791,868,1544,1566]" pageId="3" pageNumber="90" rank="species" species="browni">brownı</taxonomicName>
throatı bellyı lower part of caudal peduncle whitish; except otherwise statedı markings dark brown. Head with a dark blotch in interorbital area and on posterior part of head; snoutı suborbital area and opercle variegated.
</paragraph>
<paragraph blockId="4.[98,611,162,1682]" pageId="4" pageNumber="91">
Body with three predorsal and three postdorsal saddles; third predorsal saddle extending backwards along (but not touching) dorsal-fin base; saddles restricted to middorsal areaı not extending down side. 
<collectionCodeEnum box="[263,282,277,298]" pageId="4" pageNumber="91">A</collectionCodeEnum>
midlateral row of six blotchesı anterior two longitudinally elongated; 3rd to 5th roundish; 6th faintı at posterior extremity of caudal peduncle. Vermiculations between rows of saddles and blotches and on lower half of flankı paler brown than saddles and blotches. 
<collectionCode box="[521,540,421,442]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="91">A</collectionCode>
blackish band between posterior extremity of second blotchı connecting it with its contralateral across base of dorsal fin.
</paragraph>
<paragraph blockId="4.[98,611,162,1682]" pageId="4" pageNumber="91">
Dorsal fin hyalineı with two irregular rows of spotsı most located on rays and membranes between branches at level of 1 
<taxonomicName box="[385,452,594,616]" pageId="4" pageNumber="91" rank="subSpecies" subSpecies="and">st and</taxonomicName>
2nd branching points (and at similar position on last unbranched ray). Caudal fin hyalineı with three vertical rows of spotsı first one near base of raysı second and third one on rays and membranes between branches near branching points; spots on first row appearing as a blotch on lower and upper most rays (not separated by membranes at this position). Anal fin hyalineı with an elongated mark on raysı mainly near branching points. Pelvic fin hyalineı with two irregular rows of spotsı one in proximal position on posterior raysı one on rays and membranes between branches near branching points. Pectoral fin hyalineı with three irregular rows of spotsı one in proximal position on posterior raysı one on rays and membranes between branches near branching points of all rays (faint on posterior rays)ı and one in similar position but absent on posterior rays.
</paragraph>
<paragraph blockId="4.[98,611,162,1682]" pageId="4" pageNumber="91">
Juveniles: Body background colour yellowish whiteı marking dark brown. Four juveniles (
<figureCitation box="[106,165,1199,1221]" captionStart="Fig. 2" captionStartId="2.[98,130,1299,1318]" captionTargetBox="[127,1134,177,1250]" captionTargetId="figure@2.[98,1156,156,729]" captionTargetPageId="2" captionText="Fig. 2. Vanmanenia orcicampus ı CMK 22732 ı paratypes; Laos: Nam Ngum on Plain of Jars; aı 20.6 mm SL (reversed); and bı 23.2 mm SL." httpUri="https://zenodo.org/record/886197/files/figure.png" pageId="4" pageNumber="91">Fig. 2</figureCitation>
a) with three large blotches along lateral lineı connected by a broad band over lateral line. First blotch connected with contralateral across back by a band of paler brown pigmentsı in position of second saddle in adult. Second and third blotches connected with contralaterals across back by band of similar pigmentsı in positionı respectivelyı of blackish band across dorsal-fin base and of 5th saddle in adult. Blotches and transverse bands resulting in four elliptic pale areas (anterior one not sharply contrasted)ı with a patch of pale brown pigments in centre of eachı in position of 1stı 3rdı 4th and 6th saddles in adult. A conspicuous black blotch on proximal 
<emphasis bold="true" box="[98,102,1601,1613]" pageId="4" pageNumber="91">1</emphasis>
/ 
<emphasis bold="true" box="[111,118,1610,1622]" pageId="4" pageNumber="91">3</emphasis>
of dorsal fin.
</paragraph>
<paragraph blockId="4.[98,611,162,1682]" lastBlockId="4.[644,1157,162,242]" pageId="4" pageNumber="91">
In largest juvenile (
<figureCitation box="[339,398,1631,1653]" captionStart="Fig. 2" captionStartId="2.[98,130,1299,1318]" captionTargetBox="[127,1134,177,1250]" captionTargetId="figure@2.[98,1156,156,729]" captionTargetPageId="2" captionText="Fig. 2. Vanmanenia orcicampus ı CMK 22732 ı paratypes; Laos: Nam Ngum on Plain of Jars; aı 20.6 mm SL (reversed); and bı 23.2 mm SL." httpUri="https://zenodo.org/record/886197/files/figure.png" pageId="4" pageNumber="91">Fig. 2</figureCitation>
b; only 
<quantity box="[476,536,1631,1652]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="91" unit="mm" value="3.0">3 mm</quantity>
longer than smallest)ı pattern still distinctı but blotches smallerı restricted to along lateral line and forming a very irregular stripe. Blotch on caudal fin fainter.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="91" type="distribution">
<paragraph blockId="4.[644,1157,277,1278]" pageId="4" pageNumber="91">
<emphasis bold="true" box="[644,910,277,299]" pageId="4" pageNumber="91">Distribution and habitat.</emphasis>
<taxonomicName box="[922,1156,278,299]" class="Actinopterygii" family="Balitoridae" genus="Vanmanenia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="orcicampus">
<emphasis box="[922,1156,278,299]" italics="true" pageId="4" pageNumber="91">Vanmanenia orcicampus</emphasis>
</taxonomicName>
has been collected only once in the upper Nam Ngum at the northern edge of the Plain of Jars. The Nam Ngum crosses the western part of the plain before descending westwards through 110-km long gorges until the Nam Ngum 2 reservoir and then to the Mekong. It was not present in the other 44 sites sampled in the Nam Ngum watershed downstream of the plain. The plain is also drained to the south by the Nam Ngiep (another Mekong tributary) and the Nam Neunı which flows to 
<collectingCountry box="[736,825,594,616]" name="Vietnam" pageId="4" pageNumber="91">Vietnam</collectingCountry>
(as Song Lam) and reaches the Gulf of 
<collectingCountry box="[725,804,623,645]" name="Vietnam" pageId="4" pageNumber="91">Tonkin</collectingCountry>
. It is possibly present in the Nam Ngiep and Nam Neun on the plain but sites with an appropriate topography could not be sampled. It was not present at 33 sites in the Nam Ngiep watershed (a Mekong tributary) and 11 sites in the Nam Neun drainage (which flows directly to the Gulf of 
<collectingCountry box="[794,871,796,818]" name="Vietnam" pageId="4" pageNumber="91">Tonkin</collectingCountry>
) downstream of the plain.
</paragraph>
<paragraph blockId="4.[644,1157,277,1278]" pageId="4" pageNumber="91">
At the 
<typeStatus box="[766,813,825,846]" pageId="4" pageNumber="91">type</typeStatus>
localityı the Nam Ngum was a sluggish stream in an open agricultural area (
<figureCitation box="[652,712,882,904]" captionStart="Fig. 4" captionStartId="5.[129,161,1606,1625]" captionTargetBox="[129,1187,885,1588]" captionTargetId="figure@5.[129,1187,885,1588]" captionTargetPageId="5" captionText="Fig. 4. Nam Ngum at Ban Khangviangı Xiangkhouang Provinceı Laos; type locality of Vanmanenia orcicampus. Juveniles were found among the pebbles and cobble in the foreground rightı adult among the bamboos of the weir." httpUri="https://zenodo.org/record/886201/files/figure.png" pageId="4" pageNumber="91">Fig. 4</figureCitation>
). The bottom was made of sand to cobbleı at places covered by mud. The current was slow. This is unlikely to be the original habitatı which was probably pebble to stone bottom. The juveniles were caught among gravel and pebble. The adult was obtained among bamboos used to build a weir. The other species present were a mix of species usually found in streams with fast current and clear water (
<taxonomicName baseAuthorityName="Pellegrin &amp; Fang" baseAuthorityYear="1940" box="[957,1151,1112,1134]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="laoensis">
<emphasis box="[957,1034,1113,1134]" italics="true" pageId="4" pageNumber="91">Devario</emphasis>
cf. 
<emphasis box="[1075,1151,1113,1134]" italics="true" pageId="4" pageNumber="91">laoensis</emphasis>
</taxonomicName>
ı 
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[644,865,1142,1163]" class="Actinopterygii" family="Cyprinidae" genus="Poropuntius" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="angustus">
<emphasis box="[644,865,1142,1163]" italics="true" pageId="4" pageNumber="91">Poropuntius angustus</emphasis>
</taxonomicName>
ı 
<taxonomicName baseAuthorityName="Fowler" baseAuthorityYear="1934" class="Actinopterygii" family="Cyprinidae" genus="Scaphiodonichthys" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="acanthopterus">
<emphasis italics="true" pageId="4" pageNumber="91">Scaphiodonichthys acanthopterus</emphasis>
</taxonomicName>
ı 
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[719,903,1170,1191]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="defectiva">
<emphasis box="[719,903,1170,1191]" italics="true" pageId="4" pageNumber="91">Schistura defectiva</emphasis>
</taxonomicName>
ı 
<taxonomicName authorityName="Kottelat" authorityYear="2000" box="[917,1021,1170,1192]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="quaesita">
<emphasis box="[917,1021,1170,1192]" italics="true" pageId="4" pageNumber="91">S. quaesita</emphasis>
</taxonomicName>
ı 
<taxonomicName class="Actinopterygii" family="Gobiidae" genus="Rhinogobius" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="milleri">
<emphasis italics="true" pageId="4" pageNumber="91">Rhinogobius milleri</emphasis>
</taxonomicName>
) and species typical of sluggish waters and swamps (
<taxonomicName baseAuthorityName="Bleeker" baseAuthorityYear="1849" box="[747,890,1228,1249]" class="Actinopterygii" family="Cyprinidae" genus="Puntius" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="brevis">
<emphasis box="[747,890,1228,1249]" italics="true" pageId="4" pageNumber="91">Puntius brevis</emphasis>
</taxonomicName>
ı 
<taxonomicName baseAuthorityName="Fowler" baseAuthorityYear="1958" box="[902,1151,1228,1249]" class="Actinopterygii" family="Cyprinidae" genus="Systomus" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="jacobusboehlkei">
<emphasis box="[902,1151,1228,1249]" italics="true" pageId="4" pageNumber="91">
<taxonomicName box="[902,997,1228,1249]" class="Actinopterygii" family="Cyprinidae" genus="Systomus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="91" phylum="Chordata" rank="genus">Systomus</taxonomicName>
jacobusboehlkei
</emphasis>
</taxonomicName>
ı 
<taxonomicName baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[644,795,1257,1278]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="gachua">
<emphasis box="[644,795,1257,1278]" italics="true" pageId="4" pageNumber="91">
<taxonomicName box="[644,720,1257,1278]" class="Actinopterygii" family="Channidae" genus="Channa" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="91" phylum="Chordata" rank="genus">Channa</taxonomicName>
gachua
</emphasis>
</taxonomicName>
ı 
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[808,1042,1257,1278]" class="Actinopterygii" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="91" phylum="Chordata" rank="species" species="opercularis">
<emphasis box="[808,1042,1257,1278]" italics="true" pageId="4" pageNumber="91">
<taxonomicName box="[808,928,1257,1278]" class="Actinopterygii" family="Osphronemidae" genus="Macropodus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="91" phylum="Chordata" rank="genus">Macropodus</taxonomicName>
opercularis
</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="91" type="etymology">
<paragraph blockId="4.[644,1157,1314,1451]" pageId="4" pageNumber="91">
<emphasis bold="true" box="[644,774,1314,1335]" pageId="4" pageNumber="91">Etymology.</emphasis>
From the Latin nouns orca (jar; dative plural: orcis) and campus (plainı field)ı literally the plain with the jarsı reference to the 
<typeStatus box="[644,691,1401,1422]" pageId="4" pageNumber="91">type</typeStatus>
locality on the Plain of Jars. A compound noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>