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<document ID-DOI="http://doi.org/10.5281/zenodo.4758476" ID-GBIF-Dataset="f0f8f265-10f1-448f-8b15-6c56fc140166" ID-GBIF-Taxon="182067785" ID-ISSN="1854-0392" ID-Zenodo-Dep="4758476" ID-ZooBank="8BFBE3F7-E0EF-4EDD-BA82-9EA087F74688" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1620834297829" checkinUser="felipe" docAuthor="Grubbs, Scott A." docDate="2015" docId="03DD8784FF9AA71DFEEB58CA62D9FA0C" docLanguage="en" docName="Illiesia.11.12.147-166.pdf" docOrigin="Illiesia 11 (12)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Leuctra carolinensis : Needham and Claassen 1925" docType="treatment" docVersion="6" lastPageId="5" lastPageNumber="152" masterDocId="FFE4FFFCFF98A718FFF65E23604AFFEB" masterDocTitle="Leuctra Schusteri, A New Stonefly Species (Plecoptera: Leuctridae) Of The L. Tenuis (Pictet) Group From The Southeastern Usa" masterLastPageNumber="166" masterPageNumber="147" pageId="2" pageNumber="149" updateTime="1632012907386" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Leuctra Schusteri, A New Stonefly Species (Plecoptera: Leuctridae) Of The L. Tenuis (Pictet) Group From The Southeastern Usa</mods:title>
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<mods:namePart>Grubbs, Scott A.</mods:namePart>
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<mods:title>Illiesia</mods:title>
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<taxonomicName authority="Claassen 1923" authorityName=": Needham and Claassen" authorityYear="1925" box="[285,701,1768,1792]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="carolinensis">
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<emphasis bold="true" box="[285,526,1768,1792]" italics="true" pageId="2" pageNumber="149">Leuctra carolinensis</emphasis>
<bibRefCitation author="Claassen, P. W." box="[533,701,1768,1792]" pageId="2" pageNumber="149" pagination="257 - 263" refId="ref9047" refString="Claassen, P. W. 1923. New species of North American Plecoptera. Canadian Entomologist, 55: 257 - 263." type="journal article" year="1923">Claassen 1923</bibRefCitation>
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<taxonomicName authority="Needlefly" authorityName="Needlefly" box="[381,605,1804,1828]" genus="Carolina" kingdom="Animalia" pageId="2" pageNumber="149" rank="genus">Carolina Needlefly</taxonomicName>
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<paragraph blockId="2.[189,790,1804,1936]" pageId="2" pageNumber="149">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName box="[579,710,1840,1864]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
.specie sfile.org:TaxonName:462471
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<subSubSection box="[433,553,1912,1936]" pageId="2" pageNumber="149" type="description">
<paragraph blockId="2.[189,790,1804,1936]" box="[433,553,1912,1936]" pageId="2" pageNumber="149">
(
<figureCitation box="[442,545,1912,1936]" captionStart-0="Figs" captionStart-1="Figs 49" captionStartId-0="2.[189,241,1373,1397]" captionStartId-1="3.[189,238,1552,1576]" captionTargetBox-0="[249,1383,470,1327]" captionTargetBox-1="[249,1383,218,1505]" captionTargetId-0="figure-268@2.[249,1383,470,1327]" captionTargetId-1="figure-210@3.[249,1383,218,1505]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." captionText-1="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi-0="http://doi.org/10.5281/zenodo.4752770" figureDoi-1="http://doi.org/10.5281/zenodo.4752774" httpUri-0="https://zenodo.org/record/4752770/files/figure.png" httpUri-1="https://zenodo.org/record/4752774/files/figure.png" pageId="2" pageNumber="149">Figs. 19</figureCitation>
)
</paragraph>
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<treatmentCitation author="Claassen, P. W." box="[873,1295,1587,1609]" page="258" pageId="2" pageNumber="149" year="1923">
<taxonomicName authority="Claassen 1923: 258" authorityName=": Needham and Claassen" authorityPageNumber="258" authorityYear="1925" box="[873,1295,1587,1609]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis box="[873,1077,1587,1609]" italics="true" pageId="2" pageNumber="149">Leuctra carolinensis</emphasis>
<bibRefCitation author="Claassen, P. W." box="[1093,1295,1587,1609]" pageId="2" pageNumber="149" pagination="257 - 263" refId="ref9047" refString="Claassen, P. W. 1923. New species of North American Plecoptera. Canadian Entomologist, 55: 257 - 263." type="journal article" year="1923">Claassen 1923:258</bibRefCitation>
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.
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<typeStatus box="[1314,1414,1587,1609]" pageId="2" pageNumber="149">Holotype</typeStatus>
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.
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<taxonomicName authority=": Needham and Claassen 1925: 226" authorityName=": Needham and Claassen" authorityPageNumber="226" authorityYear="1925" box="[873,1068,1652,1674]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis box="[873,1068,1652,1674]" italics="true" pageId="2" pageNumber="149">Leuctra carolinensis</emphasis>
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:
<treatmentCitation author="Needham, J. G. &amp; P. W. Claassen" box="[1080,1427,1652,1674]" page="226" pageId="2" pageNumber="149" year="1925">
<bibRefCitation author="Needham, J. G. &amp; P. W. Claassen" box="[1080,1427,1652,1674]" pageId="2" pageNumber="149" pagination="1 - 397" refId="ref9603" refString="Needham, J. G. &amp; P. W. Claassen. 1925. A monograph of the Plecoptera or stoneflies of America north of Mexico. Thomas Say Foundation, Entomological Society of America, 2: 1 - 397. Available online: https: // archive. org / details / monographofpleco 00 need" type="journal article" year="1925">Needham and Claassen 1925:226</bibRefCitation>
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<taxonomicName authority=": Illies 1966: 84" authorityName=": Illies" authorityPageNumber="84" authorityYear="1966" box="[873,1068,1685,1706]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis box="[873,1068,1685,1706]" italics="true" pageId="2" pageNumber="149">Leuctra carolinensis</emphasis>
</taxonomicName>
:
<treatmentCitation author="Illies, J." box="[1080,1215,1685,1707]" page="84" pageId="2" pageNumber="149" year="1966">
<bibRefCitation author="Illies, J." box="[1080,1215,1685,1707]" pageId="2" pageNumber="149" refId="ref9520" refString="Illies, J. 1966. Katalog der rezenten Plecoptera. Das Tierreich 82. Walter de Gruyter and Company, Berlin. 632 pp." type="book" year="1966">Illies 1966:84</bibRefCitation>
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</paragraph>
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<subSubSection lastPageId="3" lastPageNumber="150" pageId="2" pageNumber="149" type="description">
<paragraph blockId="2.[835,1443,1754,1922]" lastBlockId="3.[835,1443,1804,1936]" lastPageId="3" lastPageNumber="150" pageId="2" pageNumber="149">
<emphasis bold="true" box="[835,904,1754,1778]" pageId="2" pageNumber="149">Male.</emphasis>
The 7th abdominal dorsal process is large and broadly rounded posteriorly; subtle lateral lobes are evident on central Appalachian populations and further north into New
<collectingCountry box="[1343,1442,1862,1886]" name="United Kingdom" pageId="2" pageNumber="149">England</collectingCountry>
(
<figureCitation box="[843,978,1898,1922]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="2" pageNumber="149">Figs. 12</figureCitation>
) but essentially absent from northwestern Alabama near the southwestern edge of this species range (
<figureCitation box="[467,540,1840,1864]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="3" pageNumber="150">Fig. 3</figureCitation>
). The specilla varies from triangular with a broad base (
<figureCitation box="[637,707,1876,1900]" captionStart="Figs 49" captionStartId="3.[189,238,1552,1576]" captionTargetBox="[249,1383,218,1505]" captionTargetId="figure-210@3.[249,1383,218,1505]" captionTargetPageId="3" captionText="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="3" pageNumber="150">Fig. 8</figureCitation>
) to ca. parallel-sided (
<figureCitation box="[378,453,1912,1936]" captionStart="Figs 49" captionStartId="3.[189,238,1552,1576]" captionTargetBox="[249,1383,218,1505]" captionTargetId="figure-210@3.[249,1383,218,1505]" captionTargetPageId="3" captionText="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="3" pageNumber="150">Fig. 6</figureCitation>
); distally the specillum is subtriangular with tubercles present and also along the distal outer margin (
<figureCitation box="[1150,1262,1840,1864]" captionStart="Figs 49" captionStartId="3.[189,238,1552,1576]" captionTargetBox="[249,1383,218,1505]" captionTargetId="figure-210@3.[249,1383,218,1505]" captionTargetPageId="3" captionText="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="3" pageNumber="150">Figs. 49</figureCitation>
). The subanal lobes extend nearly to tip of the specilla, recurved anteriorly in posterior half (
<figureCitation box="[1162,1291,1912,1936]" captionStart="Figs 49" captionStartId="3.[189,238,1552,1576]" captionTargetBox="[249,1383,218,1505]" captionTargetId="figure-210@3.[249,1383,218,1505]" captionTargetPageId="3" captionText="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="3" pageNumber="150">Figs. 4, 69</figureCitation>
).
</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4752774" ID-Zenodo-Dep="4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="3" pageNumber="150" startId="3.[189,238,1552,1576]" targetBox="[249,1383,218,1505]" targetPageId="3">
<paragraph blockId="3.[189,1444,1552,1720]" pageId="3" pageNumber="150">
Figs 49.
<taxonomicName authorityName=": Illies" authorityYear="1966" box="[303,522,1552,1576]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis box="[303,522,1552,1576]" italics="true" pageId="3" pageNumber="150">Leuctra carolinensis</emphasis>
</taxonomicName>
. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe.
</paragraph>
</caption>
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<paragraph blockId="4.[189,1443,1588,1684]" pageId="4" pageNumber="151">
Figs. 1015.
<taxonomicName authority="Grubbs, 2015" authorityName="Grubbs" authorityYear="2015" box="[340,529,1588,1612]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="151" phylum="Arthropoda" rank="species" species="schusteri" status="sp. n.">
<emphasis box="[340,529,1588,1612]" italics="true" pageId="4" pageNumber="151">Leuctra schusteri</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[539,607,1589,1612]" pageId="4" pageNumber="151" rank="species">sp. n.</taxonomicNameLabel>
, Kentucky, spring into Smyrna Creek. 1011, 7
<superScript attach="left" box="[1186,1201,1589,1603]" fontSize="6" pageId="4" pageNumber="151">th</superScript>
abdominal process, dorsal view, 350X; 12, paraprocts, caudal view, 350X; 13, paraprocts, caudal view, 1500X; 14, paraprocts, lateral view, 500X; 15, paraprocts, lateral view, 1000X. sp = specillum, sa = subanal lobe.
</paragraph>
</caption>
<subSubSection lastPageId="5" lastPageNumber="152" pageId="4" pageNumber="151" type="materials_examined">
<paragraph blockId="4.[189,797,1767,1936]" lastBlockId="5.[189,798,228,1511]" lastPageId="5" lastPageNumber="152" pageId="4" pageNumber="151">
<emphasis bold="true" box="[189,431,1767,1791]" pageId="4" pageNumber="151">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111206321" collectingDate="2008-05-04" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Colbert Co." latitude="34.8781" location="Ross Branch" longLatPrecision="6" longitude="-88.0102" pageId="4" pageNumber="151" specimenCount="17" specimenCount-male="17" stateProvince="Alabama">
<emphasis bold="true" box="[443,500,1767,1791]" pageId="4" pageNumber="151">
<collectingCountry box="[443,500,1767,1791]" name="United States of America" pageId="4" pageNumber="151">USA</collectingCountry>
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,
<emphasis bold="true" box="[521,630,1767,1791]" pageId="4" pageNumber="151">
<collectingRegion box="[521,630,1767,1791]" country="United States of America" name="Alabama" pageId="4" pageNumber="151">Alabama</collectingRegion>
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,
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, tributary to
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<quantity box="[541,611,1804,1828]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="151" unit="km" value="6.0">6 km</quantity>
SE Waterloo
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,
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,
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(
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)
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;
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<collectingCounty box="[319,516,1876,1900]" pageId="4" pageNumber="151">Lauderdale Co.</collectingCounty>
,
<collectingMunicipality pageId="4" pageNumber="151">South Fork Panther Creek</collectingMunicipality>
,
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<quantity box="[277,359,1912,1936]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="4" pageNumber="151" unit="km" value="7.5">7.5 km</quantity>
NW Waterloo
</locationDeviation>
,
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,
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,
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,
<date box="[1294,1439,1768,1793]" pageId="4" pageNumber="151" value="2008-05-04">
<collectingDate box="[1294,1439,1768,1793]" pageId="4" pageNumber="151" value="2008-05-04">4 May 2008</collectingDate>
</date>
,
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,
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(
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)
</materialsCitation>
;
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<location LSID="urn:lsid:plazi:treatment:03DD8784FF9AA71DFEEB58CA62D9FA0C:8EAB6049FF9CA71CFB72592F6560F8CF" box="[1156,1322,1804,1828]" country="United States of America" latitude="34.9745" longLatPrecision="6" longitude="-87.9942" name="Second Creek" pageId="4" pageNumber="151" stateProvince="Alabama">Second Creek</location>
,
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<quantity box="[1336,1396,1804,1828]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="151" unit="km" value="9.0">9 km</quantity>
NE Waterloo
</locationDeviation>
,
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,
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,
<date box="[1207,1371,1840,1865]" pageId="4" pageNumber="151" value="2008-05-04">
<collectingDate box="[1207,1371,1840,1865]" pageId="4" pageNumber="151" value="2008-05-04">4 May 2008</collectingDate>
</date>
,
<collectorName pageId="4" pageNumber="151">S.A. Grubbs</collectorName>
,
<specimenCount box="[943,976,1876,1900]" pageId="4" pageNumber="151" type="male">1♂</specimenCount>
(
<collectionCode box="[997,1088,1876,1900]" pageId="4" pageNumber="151">WKUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111206356" collectingDate="2007-06-16" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="New London Co." lastPageId="5" lastPageNumber="152" latitude="41.4244" location="Nehantic State Forest" longLatPrecision="6" longitude="-72.2685" municipality="Beaver Brook" pageId="4" pageNumber="151" specimenCount="10" specimenCount-male="10" stateProvince="Connecticut">
<emphasis bold="true" box="[1113,1260,1876,1900]" pageId="4" pageNumber="151">
<collectingRegion box="[1113,1260,1876,1900]" country="United States of America" name="Connecticut" pageId="4" pageNumber="151">Connecticut</collectingRegion>
</emphasis>
,
<collectingCounty pageId="4" pageNumber="151">New London Co.</collectingCounty>
, tributary to
<collectingMunicipality box="[1094,1276,1912,1936]" pageId="4" pageNumber="151">Beaver Brook</collectingMunicipality>
,
<locationDeviation lastPageId="5" lastPageNumber="152" location="Chesterfield" pageId="4" pageNumber="151">
<quantity box="[1309,1388,1912,1936]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="151" unit="km" value="4.0">4 km</quantity>
W Chesterfield
</locationDeviation>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9AA71DFEEB58CA62D9FA0C:8EAB6049FF9DA71DFE9C5EC762CCFF10" box="[362,646,228,252]" country="United States of America" county="New London Co." latitude="41.4244" longLatPrecision="6" longitude="-72.2685" municipality="Beaver Brook" name="Nehantic State Forest" pageId="5" pageNumber="152" stateProvince="Connecticut">Nehantic State Forest</location>
,
<geoCoordinate box="[672,762,228,252]" degrees="41.4244" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="41.4244">41.4244</geoCoordinate>
,
<geoCoordinate degrees="72.2685" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-72.2685">- 72.2685</geoCoordinate>
,
<date box="[293,451,264,288]" pageId="5" pageNumber="152" value="2007-06-16">
<collectingDate box="[293,451,264,288]" pageId="5" pageNumber="152" value="2007-06-16">16 June 2007</collectingDate>
</date>
,
<collectorName box="[465,614,264,288]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[630,676,264,288]" pageId="5" pageNumber="152" type="male">10♂</specimenCount>
(
<collectionCode box="[695,785,264,288]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206371" collectingDate="2007-06-17" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="41.5228" location="Pachaug State Forest" longLatPrecision="6" longitude="-71.8091" municipality="Green Fall River" pageId="5" pageNumber="152" specimenCount="1" specimenCount-male="1" stateProvince="Connecticut">
tributary to
<collectingMunicipality box="[372,608,300,324]" pageId="5" pageNumber="152">Green Fall River</collectingMunicipality>
,
<locationDeviation location="Voluntown" pageId="5" pageNumber="152">
<quantity box="[640,738,300,324]" metricMagnitude="3" metricUnit="m" metricValue="7.5" pageId="5" pageNumber="152" unit="km" value="7.5">7.5 km</quantity>
SE Voluntown
</locationDeviation>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9AA71DFEEB58CA62D9FA0C:8EAB6049FF9DA71DFEA75F736202FE83" box="[337,584,336,360]" country="United States of America" latitude="41.5228" longLatPrecision="6" longitude="-71.8091" municipality="Green Fall River" name="Pachaug State Forest" pageId="5" pageNumber="152" stateProvince="Connecticut">Pachaug State Forest</location>
,
<geoCoordinate box="[595,685,336,360]" degrees="41.5228" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="41.5228">41.5228</geoCoordinate>
,
<geoCoordinate box="[695,793,336,360]" degrees="71.8091" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-71.8091">-71.8091</geoCoordinate>
,
<date box="[189,341,372,396]" pageId="5" pageNumber="152" value="2007-06-17">
<collectingDate box="[189,341,372,396]" pageId="5" pageNumber="152" value="2007-06-17">17 June 2007</collectingDate>
</date>
,
<collectorName box="[353,498,372,396]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[511,545,371,396]" pageId="5" pageNumber="152" type="male">1♂</specimenCount>
(
<collectionCode box="[560,651,372,396]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111206391" collectingDate="1996-06-21" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Garrett Co." latitude="39.6314" location="Savage River State Forest" longLatPrecision="6" longitude="-79.1484" municipality="Little Laurel Run" pageId="5" pageNumber="152" specimenCount="17" specimenCount-female="8" specimenCount-male="9" stateProvince="Maryland">
<emphasis bold="true" box="[671,791,372,396]" pageId="5" pageNumber="152">
<collectingRegion box="[671,791,372,396]" country="United States of America" name="Maryland" pageId="5" pageNumber="152">Maryland</collectingRegion>
</emphasis>
,
<collectingCounty box="[189,327,408,432]" pageId="5" pageNumber="152">Garrett Co.</collectingCounty>
, spring into
<collectingMunicipality box="[487,698,408,432]" pageId="5" pageNumber="152">Little Laurel Run</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9AA71DFEEB58CA62D9FA0C:8EAB6049FF9DA71DFD3C5FBB61DCFE38" country="United States of America" county="Garrett Co." latitude="39.6314" longLatPrecision="6" longitude="-79.1484" municipality="Little Laurel Run" name="Savage River State Forest" pageId="5" pageNumber="152" stateProvince="Maryland">Savage River State Forest</location>
,
<geoCoordinate box="[421,508,444,468]" degrees="39.6314" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="39.6314">39.6314</geoCoordinate>
,
<geoCoordinate box="[524,622,444,468]" degrees="79.1484" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-79.1484">-79.1484</geoCoordinate>
,
<date box="[637,793,444,468]" pageId="5" pageNumber="152" value="1996-06-21">
<collectingDate box="[637,793,444,468]" pageId="5" pageNumber="152" value="1996-06-21">21 June 1996</collectingDate>
</date>
,
<collectorName box="[189,343,480,504]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[362,396,480,504]" pageId="5" pageNumber="152" type="male">9♂</specimenCount>
,
<specimenCount box="[418,451,480,504]" pageId="5" pageNumber="152" type="female">8♀</specimenCount>
(
<collectionCode box="[474,565,480,504]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206308" collectingDate="1996-05-25" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="39.4912" location="Garrett State Forest" longLatPrecision="6" longitude="-79.4584" municipality="Bull Glade Run" pageId="5" pageNumber="152" specimenCount="4" specimenCount-male="4" stateProvince="Maryland">
<collectingMunicipality box="[592,792,480,504]" pageId="5" pageNumber="152">Bull Glade Run</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9AA71DFEEB58CA62D9FA0C:8EAB6049FF9DA71DFF4B5C2761EFFDF0" box="[189,421,516,540]" country="United States of America" latitude="39.4912" longLatPrecision="6" longitude="-79.4584" municipality="Bull Glade Run" name="Garrett State Forest" pageId="5" pageNumber="152" stateProvince="Maryland">Garrett State Forest</location>
,
<geoCoordinate box="[432,521,516,540]" degrees="39.4912" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="39.4912">39.4912</geoCoordinate>
,
<geoCoordinate box="[534,632,516,540]" degrees="79.4584" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-79.4584">-79.4584</geoCoordinate>
,
<date box="[643,793,516,540]" pageId="5" pageNumber="152" value="1996-05-25">
<collectingDate box="[643,793,516,540]" pageId="5" pageNumber="152" value="1996-05-25">25 May 1996</collectingDate>
</date>
,
<collectorName box="[189,335,552,576]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[348,381,552,576]" pageId="5" pageNumber="152" type="male">4♂</specimenCount>
(
<collectionCode box="[397,485,552,576]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206382" collectingDate="1996-07-12" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" latitude="39.6597" location="Alexander Run" longLatPrecision="6" longitude="-79.2254" pageId="5" pageNumber="152" specimenCount="11" specimenCount-female="5" specimenCount-male="6" stateProvince="Maryland">
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9AA71DFEEB58CA62D9FA0C:8EAB6049FF9DA71DFE0A5C0B62F9FDD4" box="[508,691,552,576]" country="United States of America" latitude="39.6597" longLatPrecision="6" longitude="-79.2254" name="Alexander Run" pageId="5" pageNumber="152" stateProvince="Maryland">Alexander Run</location>
,
<geoCoordinate box="[704,791,552,576]" degrees="39.6597" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="39.6597">39.6597</geoCoordinate>
,
<geoCoordinate box="[189,285,588,612]" degrees="79.2254" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-79.2254">-79.2254</geoCoordinate>
,
<date box="[314,489,588,612]" pageId="5" pageNumber="152" value="1996-07-12">
<collectingDate box="[314,489,588,612]" pageId="5" pageNumber="152" value="1996-07-12">12 July 1996</collectingDate>
</date>
,
<collectorName box="[514,674,588,612]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[702,735,588,612]" pageId="5" pageNumber="152" type="male">6♂</specimenCount>
,
<specimenCount box="[764,797,588,612]" pageId="5" pageNumber="152" type="female">5♀</specimenCount>
(
<collectionCode box="[197,287,624,648]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="3111206343" collectingDate="1991-08-01" collectionCode="WKUC" collectorName="S. A. Grubbs" country="United States of America" county="Westmoreland Co." latitude="40.1675" location="Linn Camp Creek" longLatPrecision="6" longitude="-79.2307" pageId="5" pageNumber="152" specimenCount="1" specimenCount-male="1" stateProvince="Pennsylvania">
<emphasis bold="true" box="[307,471,623,647]" pageId="5" pageNumber="152">
<collectingRegion box="[307,471,623,647]" country="United States of America" name="Pennsylvania" pageId="5" pageNumber="152">Pennsylvania</collectingRegion>
</emphasis>
,
<collectingCounty box="[486,709,623,648]" pageId="5" pageNumber="152">Westmoreland Co.</collectingCounty>
, spring into
<location LSID="urn:lsid:plazi:treatment:03DD8784FF9AA71DFEEB58CA62D9FA0C:8EAB6049FF9DA71DFF025CB76186FD47" box="[244,460,660,684]" country="United States of America" county="Westmoreland Co." latitude="40.1675" longLatPrecision="6" longitude="-79.2307" name="Linn Camp Creek" pageId="5" pageNumber="152" stateProvince="Pennsylvania">Linn Camp Creek</location>
,
<geoCoordinate box="[474,564,659,683]" degrees="40.1675" direction="north" orientation="latitude" pageId="5" pageNumber="152" precision="5" value="40.1675">40.1675</geoCoordinate>
,
<geoCoordinate box="[577,674,660,684]" degrees="79.2307" direction="west" orientation="longitude" pageId="5" pageNumber="152" precision="5" value="-79.2307">-79.2307</geoCoordinate>
,
<date pageId="5" pageNumber="152" value="1991-08-01">
<collectingDate pageId="5" pageNumber="152" value="1991-08-01">1 August 1991</collectingDate>
</date>
,
<collectorName box="[256,401,696,720]" pageId="5" pageNumber="152">S.A. Grubbs</collectorName>
,
<specimenCount box="[411,445,696,720]" pageId="5" pageNumber="152" type="male">1♂</specimenCount>
(
<collectionCode box="[459,550,696,720]" pageId="5" pageNumber="152">WKUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="152" type="distribution">
<paragraph blockId="5.[189,798,228,1511]" pageId="5" pageNumber="152">
<emphasis bold="true" box="[189,348,731,755]" pageId="5" pageNumber="152">Distribution.</emphasis>
<collectingCountry box="[365,424,732,756]" name="United States of America" pageId="5" pageNumber="152">USA</collectingCountry>
: MD, MS, NC, NJ, TN, VA (
<bibRefCitation author="DeWalt R. E. &amp; M. D. Maehr &amp; U. Neu-Becker &amp; G. Steuber" box="[197,425,768,792]" pageId="5" pageNumber="152" refId="ref9097" refString="DeWalt R. E., M. D. Maehr, U. Neu-Becker, &amp; G. Steuber. 2015. Plecoptera species file online. Version 5.0 / 5.0. Available online: http: // Plecoptera. SpeciesFile. org [retrieved 19 July 2015]" type="url" year="2015">DeWalt et al. 2015</bibRefCitation>
), AL (
<bibRefCitation author="Grubbs, S. A." box="[510,668,768,792]" pageId="5" pageNumber="152" pagination="59 - 64" refId="ref9147" refString="Grubbs, S. A. 2010. Leuctra usdi n. sp., a new stonefly (Plecoptera: Leuctridae) of the L. tenuis (Pictet) species group from the southeastern U. S. A. plus three new Alabama state records. Zootaxa, 2498: 59 - 64. Available online: http: // 216.92.145.68 / zootaxa / 2010 / f / z 02498 p 064 f. pdf" type="journal article" year="2010">Grubbs 2010</bibRefCitation>
), CT (new state record), PA (
<bibRefCitation author="Masteller, E. C." box="[401,573,804,828]" pageId="5" pageNumber="152" pagination="107 - 120" refId="ref9546" refString="Masteller, E. C. 1996. New records of stoneflies (Plecoptera) with an annotated checklist of the species for Pennsylvania. Great Lakes Entomologist, 29: 107 - 120. Available online: http: // michentsoc. org / gle-pdfs / vol 29 no 3. pdf page = 17" type="journal article" year="1996">Masteller 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="152" type="discussion">
<paragraph blockId="5.[189,798,228,1511]" pageId="5" pageNumber="152">
<emphasis bold="true" box="[189,304,840,864]" pageId="5" pageNumber="152">Remarks.</emphasis>
<taxonomicName box="[314,523,840,864]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="carolinensi">
<emphasis box="[314,523,840,864]" italics="true" pageId="5" pageNumber="152">Leuctra carolinensi</emphasis>
</taxonomicName>
s and
<taxonomicName box="[601,704,840,864]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="tenella">
<emphasis box="[601,704,840,864]" italics="true" pageId="5" pageNumber="152">L. tenella</emphasis>
</taxonomicName>
appear to be sister species based on overall similarity in structure of the paraproct and the 7th abdominal dorsal process. The abdominal process resembles only
<taxonomicName box="[251,352,984,1008]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="tenella">
<emphasis box="[251,352,984,1008]" italics="true" pageId="5" pageNumber="152">L. tenella</emphasis>
</taxonomicName>
, a species with a similar distribution and springsummer emergence in the southern and central Appalachian Mountains. The two records of
<taxonomicName authorityName=": Illies" authorityYear="1966" box="[346,518,1092,1116]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis box="[346,518,1092,1116]" italics="true" pageId="5" pageNumber="152">L. carolinensis</emphasis>
</taxonomicName>
from low elevation streams in southeastern
<collectingRegion box="[500,642,1128,1152]" country="United States of America" name="Connecticut" pageId="5" pageNumber="152">Connecticut</collectingRegion>
represent a northeastern range extension and suggest that the distribution of this species is still poorly understood. The dorsal process of
<taxonomicName authorityName=": Illies" authorityYear="1966" box="[610,768,1235,1259]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis box="[610,768,1235,1259]" italics="true" pageId="5" pageNumber="152">L. carolinensis</emphasis>
</taxonomicName>
is broadly-rounded posteriorly (
<figureCitation box="[562,673,1272,1296]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="5" pageNumber="152">Figs. 13</figureCitation>
) whereas the process of
<taxonomicName box="[373,476,1307,1331]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="tenella">
<emphasis box="[373,476,1307,1331]" italics="true" pageId="5" pageNumber="152">L. tenella</emphasis>
</taxonomicName>
is subtriangular (
<figureCitation box="[698,784,1307,1332]" captionStart="Figs" captionStartId="8.[189,241,1152,1176]" captionTargetBox="[249,1383,254,1105]" captionTargetId="figure-432@8.[249,1383,254,1105]" captionTargetPageId="8" captionText="Figs. 1922. Leuctra tenella, Pennsylvania, spring into Powdermill Run. 19, 7th abdominal process, dorsal view, 350X; 20, paraprocts, caudal view, 1000X; 21, L. tenella, paraprocts, lateral view, 350X; 22, L. tenella, paraprocts, lateral view, 1500X. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752780" httpUri="https://zenodo.org/record/4752780/files/figure.png" pageId="5" pageNumber="152">Fig. 19</figureCitation>
). Although the specilla of
<taxonomicName authorityName=": Illies" authorityYear="1966" box="[483,640,1344,1368]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="carolinensis">
<emphasis box="[483,640,1344,1368]" italics="true" pageId="5" pageNumber="152">L. carolinensis</emphasis>
</taxonomicName>
vary slightly in overall shape (
<figureCitation box="[433,566,1379,1404]" captionStart="Figs 49" captionStartId="3.[189,238,1552,1576]" captionTargetBox="[249,1383,218,1505]" captionTargetId="figure-210@3.[249,1383,218,1505]" captionTargetPageId="3" captionText="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="5" pageNumber="152">Figs. 6, 8</figureCitation>
), the location of tubercles and the shape of the distal portion are similar among the material examined (
<figureCitation box="[678,789,1451,1476]" captionStart="Figs 49" captionStartId="3.[189,238,1552,1576]" captionTargetBox="[249,1383,218,1505]" captionTargetId="figure-210@3.[249,1383,218,1505]" captionTargetPageId="3" captionText="Figs 49. Leuctra carolinensis. 4, paraprocts, lateral view, 900X, Connecticut, tributary to Beaver Brook; 5, paraprocts, dorsal view, 1000X, Alabama, tributary to Ross Branch; 6, paraprocts, lateral view, 350X, Maryland, Bull Glade Run; 7, paraprocts, lateral view, 1000X, Maryland, Bull Glade Run; 8, paraprocts, lateral view, 350X, Alabama, tributary to Ross Branch; 9, paraprocts, lateral view, 1000X, Alabama, tributary to Ross Branch. sp = specillum, sa = subanal lobe." figureDoi="http://doi.org/10.5281/zenodo.4752774" httpUri="https://zenodo.org/record/4752774/files/figure.png" pageId="5" pageNumber="152">Figs. 49</figureCitation>
) and distinct from
<taxonomicName box="[400,499,1487,1511]" class="Insecta" family="Leuctridae" genus="Leuctra" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="152" phylum="Arthropoda" rank="species" species="tenella">
<emphasis box="[400,499,1487,1511]" italics="true" pageId="5" pageNumber="152">L. tenella</emphasis>
</taxonomicName>
(
<figureCitation box="[514,646,1487,1511]" captionStart="Figs" captionStartId="2.[189,241,1373,1397]" captionTargetBox="[249,1383,470,1327]" captionTargetId="figure-268@2.[249,1383,470,1327]" captionTargetPageId="2" captionText="Figs. 13. Leuctra carolinensis. 1, 7th abdominal process, dorsal view, 300X, Connecticut, tributary to Beaver Brook; 2, 7th abdominal process, dorsal view, 350X, Maryland, spring into Little Laurel Run; 3, 7th abdominal process, dorsal view, 350X, Alabama, tributary to Ross Branch." figureDoi="http://doi.org/10.5281/zenodo.4752770" httpUri="https://zenodo.org/record/4752770/files/figure.png" pageId="5" pageNumber="152">Figs. 2122</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>