186 lines
24 KiB
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186 lines
24 KiB
XML
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<mods:title id="D907CF8E080ADB15C13F154800E4FF85">An appraisal of the Middle-Late Miocene fossil decapod crustaceans of the ‘ Faluns’ (Anjou-Touraine, France)</mods:title>
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<mods:namePart id="DF1C4FE48C6CBD938798DB71A69806E1">Ossó, Àlex</mods:namePart>
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<taxonomicName id="4C784D649071FFBDFCB99BEE0E904687" ID-CoL="69c4474c-d6e5-40e1-ab8c-20c655ec9ac6" authority="Guinot, De Angeli & Garassino, 2007" authorityName="Guinot, De Angeli & Garassino, Paratethys" authorityYear="2007" box="[814,1454,216,244]" class="Malacostraca" family="Corystidae" genus="Hebertides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="372" phylum="Arthropoda" rank="species" species="jurassica">
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<emphasis id="B90CEAF59071FFBDFCB99BEE0F394687" bold="true" box="[814,1031,216,243]" italics="true" pageId="7" pageNumber="372">Hebertides jurassica</emphasis>
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Guinot, De Angeli & Garassino, 2007
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(
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<figureCitation id="13432A629071FFBDFBAA9BCF0F444767" box="[1085,1146,249,275]" captionStart="FIG" captionStartId="6.[132,143,1806,1824]" captionTargetBox="[132,1455,215,1763]" captionTargetId="figure@6.[132,1455,200,1774]" captionTargetPageId="6" captionText="FIG. 3. — A-C, Calappa praelata Lőrenthey in Lőrenthey & Beurlen, 1929, MS2006 FS 134a: A, dorsal view; B, frontal view; C, close-up of posterolateral margin; D, E, Calappa sp., MS2018-2-1: D, left chela, MS2018-2-2; E, right chela; F-J, Hebertides jurassica Guinot, De Angeli & Garassino, 2007; F-H, MS2018-1-1: F, right lateral view; G, frontal view; H, dorsal view; I, J, ULB-SO-1: I, fragment of right dorsal carapace; J, the same fragment showing bryozoans incrusted. Scale bars: A-H, 10 mm; I, J, 5 mm. Photographs by À. Ossó." figureDoi="http://doi.org/10.5281/zenodo.3700040" httpUri="https://zenodo.org/record/3700040/files/figure.png" pageId="7" pageNumber="372">Fig. 3</figureCitation>
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F-J)
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<subSubSection id="C362656C9071FFBDFCBA9A040FD847BD" pageId="7" pageNumber="372" type="reference_group">
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<treatmentCitation id="0AD910F69071FFBDFCBA9A0408684710" author="GUINOT D. & DE ANGELI A. & GARASSINO A." page="260" pageId="7" pageNumber="372" year="2007">
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<taxonomicName id="4C784D649071FFBDFCBA9A0408684710" ID-CoL="69c4474c-d6e5-40e1-ab8c-20c655ec9ac6" authority="Guinot, De Angeli & Garassino, 2007 b: 241 - 260" authorityName="Guinot, De Angeli & Garassino, Paratethys" authorityPageNumber="241 - 260" authorityYear="2007" class="Malacostraca" family="Corystidae" genus="Hebertides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="372" phylum="Arthropoda" rank="species" species="jurassica">
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<emphasis id="B90CEAF59071FFBDFCBA9A0408DA473E" box="[813,996,306,330]" italics="true" pageId="7" pageNumber="372">Hebertides jurassica</emphasis>
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<bibRefCitation id="EFE94B169071FFBDFC7C9A0408684710" author="GUINOT D. & DE ANGELI A. & GARASSINO A." pageId="7" pageNumber="372" pagination="241 - 260" refId="ref11864" refString="GUINOT D., DE ANGELI A. & GARASSINO A. 2007 b. - Hebertides jurassica n. gen., n. sp. (Crustacea, Decapoda, Brachyura) from the Middle Jurassic (Bathonian) of Normandy (France). Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale in Milano 14 (2): 241 - 260." type="journal article" year="2007">Guinot, De Angeli & Garassino, 2007b: 241- 260</bibRefCitation>
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, fig. 1-3. —
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<treatmentCitation id="0AD910F69071FFBDFC4C9A7B0FDE4710" author="VAN BAKEL B. W. M. & JAGT J. W. M. & ARTAL P. & FRAAIJE R. H. B." box="[987,1248,332,356]" page="80" pageId="7" pageNumber="372" year="2009">
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<bibRefCitation id="EFE94B169071FFBDFC4C9A7B0FDE4710" author="VAN BAKEL B. W. M. & JAGT J. W. M. & ARTAL P. & FRAAIJE R. H. B." box="[987,1248,332,356]" pageId="7" pageNumber="372" pagination="79 - 85" refId="ref15410" refString="VAN BAKEL B. W. M., JAGT J. W. M., ARTAL P. & FRAAIJE R. H. B. 2009. - Harenacorystes johanjansseni, a new Pliocene crab (Crustacea, Decapoda) from the Netherlands, and notes on Miocene-Pliocene corystoid crabs from the North Sea Basin. Bulletin of the Mizunami Fossil Museum 35: 79 - 85." type="journal article" year="2009">
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Van Bakel
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<emphasis id="B90CEAF59071FFBDFBDF9A7B0F444710" box="[1096,1146,332,356]" italics="true" pageId="7" pageNumber="372">et al.</emphasis>
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2009: 80
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</bibRefCitation>
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</treatmentCitation>
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. —
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||
<treatmentCitation id="0AD910F69071FFBDFA9A9A7A08A9470B" author="SCHWEITZER C. E. & FELDMANN R. M. & GARASSINO A. & KARASAWA H. & SCHWEIGERT G." page="104" pageId="7" pageNumber="372" year="2010">
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<bibRefCitation id="EFE94B169071FFBDFA9A9A7A08A9470B" author="SCHWEITZER C. E. & FELDMANN R. M. & GARASSINO A. & KARASAWA H. & SCHWEIGERT G." pageId="7" pageNumber="372" pagination="1 - 1222" refId="ref14815" refString="SCHWEITZER C. E., FELDMANN R. M., GARASSINO A., KARASAWA H. & SCHWEIGERT G. 2010. - Systematic list of fossil decapod crustacean species. Crustaceana Monograph 10: 1 - 1222. https: // doi. org / 10.1163 / ej. 9789004178915. i- 222" type="journal article" year="2010">
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Schweitzer
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<emphasis id="B90CEAF59071FFBDFAE89A7B0E8E4710" box="[1407,1456,332,356]" italics="true" pageId="7" pageNumber="372">et al.</emphasis>
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2010: 104
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</bibRefCitation>
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</treatmentCitation>
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. —
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||
<treatmentCitation id="0AD910F69071FFBDFC549A510FFE470B" author="TAYLOR P. D. & BRETON G. & GUINOT D. & DE ANGELI A. & GARASSINO A." box="[963,1216,359,383]" page="71" pageId="7" pageNumber="372" year="2012">
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<bibRefCitation id="EFE94B169071FFBDFC549A510FFE470B" author="TAYLOR P. D. & BRETON G. & GUINOT D. & DE ANGELI A. & GARASSINO A." box="[963,1216,359,383]" pageId="7" pageNumber="372" pagination="71 - 83" refId="ref15182" refString="TAYLOR P. D., BRETON G., GUINOT D., DE ANGELI A. & GARASSINO A. 2012. - The Cenozoic age of the supposed Jurassic crab Hebertides jurassica Guinot, De Angeli & Garassino, 2007 (Crustacea, Decapoda, Brachyura). Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale in Milano 153 (1): 71 - 83." type="journal article" year="2012">
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Taylor
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<emphasis id="B90CEAF59071FFBDFB9D9A510F05470B" box="[1034,1083,359,383]" italics="true" pageId="7" pageNumber="372">et al.</emphasis>
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2012: 71-83
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</bibRefCitation>
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</treatmentCitation>
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||
, fig. 1; — Taylor 2012: 9-11, fig. 2. —
|
||
<treatmentCitation id="0AD910F69071FFBDFC519AB40FB047ED" author="JAGT J. W. M. & VAN BAKEL B. W. M. & GUINOT D. & FRAAIJE R. H. B. & ARTAL P." box="[966,1166,385,410]" page="882" pageId="7" pageNumber="372" year="2015">
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<bibRefCitation id="EFE94B169071FFBDFC519AB40FB047ED" author="JAGT J. W. M. & VAN BAKEL B. W. M. & GUINOT D. & FRAAIJE R. H. B. & ARTAL P." box="[966,1166,385,410]" pageId="7" pageNumber="372" pagination="847 - 920" refId="ref12099" refString="JAGT J. W. M., VAN BAKEL B. W. M., GUINOT D., FRAAIJE R. H. B. & ARTAL P. 2015. - Fossil Brachyura, in CASTRO P., DAVIE P., GUINOT D., SCHRAM F. & VON VAUPEL KLEIN J. (eds), Treatise on Zoology - Anatomy, Taxonomy, Biology. The Crustacea, vol. 9, Part C-II, Chapter 71 - 15. Brill, Leiden / Boston: 847 - 920. https: // doi. org / 10.1163 / 9789004190832 _ 018" type="book chapter" year="2015">
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Jagt
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||
<emphasis id="B90CEAF59071FFBDFC669AB40F1E47ED" box="[1009,1056,385,409]" italics="true" pageId="7" pageNumber="372">et al.</emphasis>
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||
2015: 882
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
. —
|
||
<treatmentCitation id="0AD910F69071FFBDFB209AB40EAF47ED" author="EMMERSON W. D." box="[1207,1425,385,409]" page="323" pageId="7" pageNumber="372" year="2017">
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<bibRefCitation id="EFE94B169071FFBDFB209AB40EAF47ED" author="EMMERSON W. D." box="[1207,1425,385,409]" pageId="7" pageNumber="372" refId="ref10949" refString="EMMERSON W. D. 2017. - A Guide to, and Checklist for, the Decapoda of Namibia, South Africa and Mozambique. Volume 2. Cambridge Scholars Publishing, Cambridge, 650 p." type="book" year="2017">Emmerson 2017: 323</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph id="8BC736E79071FFBDFCBA9A870FD847BD" blockId="7.[813,1254,433,457]" box="[813,1254,433,457]" pageId="7" pageNumber="372">
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Corystid –
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<bibRefCitation id="EFE94B169071FFBDFC0C9A870F9A47BD" author="GUINOT D. & DE ANGELI A. & GARASSINO A." box="[923,1188,433,457]" pageId="7" pageNumber="372" pagination="53 - 55" refId="ref11779" refString="GUINOT D., DE ANGELI A. & GARASSINO A. 2007 a. - Discovery of the oldest eubrachyuran crab from the Middle Jurassic (Bathonian) of Normandy (France), in 3 rd Symposium on Mesozoic and Cenozoic Decapod Crustaceans - Museo di Storia Naturale di Milano, May 23 - 25, 2007. Memorie della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 35 (2): 53 - 55." type="journal article" year="2007">
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||
Guinot
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<emphasis id="B90CEAF59071FFBDFC7E9A840F2647BD" box="[1001,1048,433,457]" italics="true" pageId="7" pageNumber="372">et al.</emphasis>
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2007a: 53-55
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||
</bibRefCitation>
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||
, fig. 1.
|
||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C362656C9071FFBDFCBA9AD60E3644B3" pageId="7" pageNumber="372" type="materials_examined">
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<paragraph id="8BC736E79071FFBDFCBA9AD60FA94459" blockId="7.[813,1456,480,558]" pageId="7" pageNumber="372">
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MATERIAL EXAMINED AND MEASUREMENTS (in mm). —
|
||
<materialsCitation id="3B103CBA9071FFBDFAD89AD70FA94459" ID-GBIF-Occurrence="2574969788" pageId="7" pageNumber="372">
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||
Carapace, MS
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||
<date id="FFC610279071FFBDFCC29ACD088E4467" box="[853,944,507,531]" pageId="7" pageNumber="372" value="2018-01-01">2018-1-1</date>
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: L = 16.5, W = 13, FOW = 13. — Fragment of carapace, ULB-SO-1: L = 11.4; W = 5.8.
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</materialsCitation>
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</paragraph>
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||
<paragraph id="8BC736E79071FFBDFCBA99730E3644B3" blockId="7.[813,1455,581,711]" pageId="7" pageNumber="372">
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||
LOCALITY AND HORIZON. — Carapace: ‘La carrière-musée’ (Channay-sur-Lathan, Indre-et-Loire). Bioclastic calcarenite with
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<emphasis id="B90CEAF59071FFBDFAC89969084544E6" italics="true" pageId="7" pageNumber="372">Anadara turonica</emphasis>
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||
, Tortonian (Late Miocene). Fragment of carapace: ‘La Sonneterie’ quarry, Meigné-le-Vicomte (Maine-et-Loire). ‘Savignean facies’, Langhian-Serravallian (Middle Miocene).
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||
</paragraph>
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||
</subSubSection>
|
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<subSubSection id="C362656C9071FFBDFCBA99C10FCF40DB" pageId="7" pageNumber="372" type="description">
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<paragraph id="8BC736E79071FFBDFCBA99C10FCF40DB" blockId="7.[813,1456,759,1711]" pageId="7" pageNumber="372">
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DESCRIPTION (modified from
|
||
<bibRefCitation id="EFE94B169071FFBDFBE599C10E764565" author="GUINOT D. & DE ANGELI A. & GARASSINO A." box="[1138,1352,759,786]" pageId="7" pageNumber="372" pagination="241 - 260" refId="ref11864" refString="GUINOT D., DE ANGELI A. & GARASSINO A. 2007 b. - Hebertides jurassica n. gen., n. sp. (Crustacea, Decapoda, Brachyura) from the Middle Jurassic (Bathonian) of Normandy (France). Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale in Milano 14 (2): 241 - 260." type="journal article" year="2007">
|
||
Guinot
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||
<emphasis id="B90CEAF59071FFBDFB5F99C10FC54566" box="[1224,1275,759,786]" italics="true" pageId="7" pageNumber="372">et al.</emphasis>
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2007b
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||
</bibRefCitation>
|
||
) Carapace longitudinally ovate, longer than wide, convex in both sections; maximum width at the epibranchial teeth level; W/L ratio about 0.75. Cuticle well preserved. Regions relatively well defined, slightly swollen and separated by weak, smooth grooves; medial regions faintly distinct, forming a continuous medial area. Dorsal surface of carapace covered by rounded tubercles in anterior half, grouped and aligned in posterior half. Front narrow, broken, medially V-notched. Orbits broad; inner orbital tooth blunt, as a fold; supraorbital margin sinuous, finely serrated, with two V-shaped fissures; outer orbital tooth triangular, acute, prominent. Supraorbital area slightly depressed, smooth. Anterolateral margin convex, with three granular teeth (excluded the extraorbital one); the second sunken, placed at lower level that the rest; the third the larger; the fourth the smallest. Posterolateral margin slightly convex. Posterior margin convex, with marked granular rim. Protogastric regions slightly swollen, with two tubercles obliquely aligned at each lobe. Mesogastric region subpentagonal, slender and elongate anteriorly, broad posteriorly. Metagastric region indistinct from mesogastric region. Urogastric region subrectangular elongate, separated from meso-, metagastric regions by two gastric pits. Cardiac region subhexagonal, somewhat raised respect to urogastric region. Meta-, urogastric and cardiac regions, bounded laterally by well-marked branchiocardiac groove. Intestinal region indistinct. Hepatic region subtriangular, well delimited by cervical and hepatic groove. Branchial regions slightly swollen, epi- and mesobranchial lobes delimited by a weak oblique groove. Ventral parts and appendages not present.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C362656C9071FFB2FCBA9DE508364426" lastPageId="8" lastPageNumber="373" pageId="7" pageNumber="372" type="discussion">
|
||
<paragraph id="8BC736E79071FFBDFCBA9DE508AF4098" blockId="7.[813,1456,1747,2030]" box="[813,913,1747,1773]" pageId="7" pageNumber="372">REMARKS</paragraph>
|
||
<paragraph id="8BC736E79071FFB2FCBA9DC209C34727" blockId="7.[813,1456,1747,2030]" lastBlockId="8.[132,776,217,595]" lastPageId="8" lastPageNumber="373" pageId="7" pageNumber="372">
|
||
<taxonomicName id="4C784D649071FFBDFCBA9DC208C6417A" authorityName="Guinot, De Angeli & Garassino, Paratethys" authorityYear="2007" box="[813,1016,1780,1806]" class="Malacostraca" family="Corystidae" genus="Hebertides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="372" phylum="Arthropoda" rank="species" species="jurassica">
|
||
<emphasis id="B90CEAF59071FFBDFCBA9DC208C6417A" box="[813,1016,1780,1806]" italics="true" pageId="7" pageNumber="372">Hebertides jurassica</emphasis>
|
||
</taxonomicName>
|
||
was erected based on a unique specimen found as a loose block in a quarry of Ranville (
|
||
<collectingRegion id="49BCF8059071FFBDFAB09C220E9A415B" box="[1319,1444,1812,1839]" country="France" name="Haute-Normandie" pageId="7" pageNumber="372">Normandy</collectingRegion>
|
||
). In spite of the strikingly fine preservation of the specimen, its age was considered as the same as the rocks of the quarry, i.e., Bathonian (Middle Jurassic) (
|
||
<bibRefCitation id="EFE94B169071FFBDFB319C450EBF41F9" author="GUINOT D. & DE ANGELI A. & GARASSINO A." box="[1190,1409,1907,1933]" pageId="7" pageNumber="372" pagination="53 - 55" refId="ref11779" refString="GUINOT D., DE ANGELI A. & GARASSINO A. 2007 a. - Discovery of the oldest eubrachyuran crab from the Middle Jurassic (Bathonian) of Normandy (France), in 3 rd Symposium on Mesozoic and Cenozoic Decapod Crustaceans - Museo di Storia Naturale di Milano, May 23 - 25, 2007. Memorie della Societa italiana di Scienze naturali e del Museo civico di Storia naturale di Milano 35 (2): 53 - 55." type="journal article" year="2007">
|
||
Guinot
|
||
<emphasis id="B90CEAF59071FFBDFB689C420E0B41F9" box="[1279,1333,1907,1933]" italics="true" pageId="7" pageNumber="372">et al.</emphasis>
|
||
2007a
|
||
</bibRefCitation>
|
||
, b). Further examination of the matrix of this specimen revealed that the bryozoans (mainly cheilostomes) attached to the matrix and specimen were not Jurassic in age but younger, Cenozoic, and probably Miocene in age. Therefore, it was assumed that the specimen was accidentally discarded in the quarry by a fossil collector. Thus, the provenance of that specimen remained unknown (Taylor 2012;
|
||
<bibRefCitation id="EFE94B16907EFFB2FDB89A0F09D34727" author="TAYLOR P. D. & BRETON G. & GUINOT D. & DE ANGELI A. & GARASSINO A." box="[559,749,313,339]" pageId="8" pageNumber="373" pagination="71 - 83" refId="ref15182" refString="TAYLOR P. D., BRETON G., GUINOT D., DE ANGELI A. & GARASSINO A. 2012. - The Cenozoic age of the supposed Jurassic crab Hebertides jurassica Guinot, De Angeli & Garassino, 2007 (Crustacea, Decapoda, Brachyura). Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale in Milano 153 (1): 71 - 83." type="journal article" year="2012">
|
||
Taylor
|
||
<emphasis id="B90CEAF5907EFFB2FDEC9A0C09914727" box="[635,687,313,339]" italics="true" pageId="8" pageNumber="373">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC736E7907EFFB2FF0C9A6F08364426" blockId="8.[132,776,217,595]" pageId="8" pageNumber="373">
|
||
Despite this, complete specimens of
|
||
<taxonomicName id="4C784D64907EFFB2FD8C9A6F09DC4706" authorityName="Guinot, De Angeli & Garassino, Paratethys" authorityYear="2007" box="[539,738,345,371]" class="Malacostraca" family="Corystidae" genus="Hebertides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="373" phylum="Arthropoda" rank="species" species="jurassica">
|
||
<emphasis id="B90CEAF5907EFFB2FD8C9A6F09DC4706" box="[539,738,345,371]" italics="true" pageId="8" pageNumber="373">Hebertides jurassica</emphasis>
|
||
</taxonomicName>
|
||
are known among French collectors’ private collections, which regularly collect in the ‘Faluns’ of Anjou-Touraine. Fragmentary remains of their carapaces are relatively common among the typical shell grit of the ‘Faluns’ (
|
||
<figureCitation id="13432A62907EFFB2FD979AEE09004786" box="[512,574,472,498]" captionStart="FIG" captionStartId="6.[132,143,1806,1824]" captionTargetBox="[132,1455,215,1763]" captionTargetId="figure@6.[132,1455,200,1774]" captionTargetPageId="6" captionText="FIG. 3. — A-C, Calappa praelata Lőrenthey in Lőrenthey & Beurlen, 1929, MS2006 FS 134a: A, dorsal view; B, frontal view; C, close-up of posterolateral margin; D, E, Calappa sp., MS2018-2-1: D, left chela, MS2018-2-2; E, right chela; F-J, Hebertides jurassica Guinot, De Angeli & Garassino, 2007; F-H, MS2018-1-1: F, right lateral view; G, frontal view; H, dorsal view; I, J, ULB-SO-1: I, fragment of right dorsal carapace; J, the same fragment showing bryozoans incrusted. Scale bars: A-H, 10 mm; I, J, 5 mm. Photographs by À. Ossó." figureDoi="http://doi.org/10.5281/zenodo.3700040" httpUri="https://zenodo.org/record/3700040/files/figure.png" pageId="8" pageNumber="373">Fig. 3</figureCitation>
|
||
I-J). Therefore, the common occurrence of
|
||
<taxonomicName id="4C784D64907EFFB2FE1D9ACF09384466" authorityName="Guinot, De Angeli & Garassino, Paratethys" authorityYear="2007" box="[394,518,505,530]" class="Malacostraca" family="Corystidae" genus="Hebertides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="373" phylum="Arthropoda" rank="species" species="jurassica">
|
||
<emphasis id="B90CEAF5907EFFB2FE1D9ACF09384466" box="[394,518,505,530]" italics="true" pageId="8" pageNumber="373">H. jurassica</emphasis>
|
||
</taxonomicName>
|
||
in the ‘Faluns’ leads us to suggest that, likely, the
|
||
<typeStatus id="54C38845907EFFB2FE09992E0AC34446" box="[414,509,536,562]" pageId="8" pageNumber="373" type="holotype">holotype</typeStatus>
|
||
comes from a locality in this region whose age is confirmed as Middle-Late Miocene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |