389 lines
30 KiB
XML
389 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.814.28869" ID-GBIF-Dataset="b8296d03-56e2-4efa-89ec-2f8cbe851740" ID-PMC="PMC6333729" ID-Pensoft-Pub="1313-2970-814-53" ID-PubMed="30651712" ID-ZBK="25C7041051D8445C99A1A7E158CFD507" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-814-53" ModsDocOrigin="ZooKeys 814" ModsDocTitle="New species of bone-eating worm Osedax from the abyssal South Atlantic Ocean (Annelida, Siboglinidae)" checkinTime="1547042491050" checkinUser="pensoft" docAuthor="Fujiwara, Yoshihiro, Jimi, Naoto, Sumida, Paulo Y. G., Kawato, Masaru & Hiroshi Kitazato," docDate="2019" docId="F1CB4DA529B8EEBE075E618E954F70D2" docLanguage="en" docName="ZooKeys 814: 53-69" docOrigin="ZooKeys 814" docSource="http://dx.doi.org/10.3897/zookeys.814.28869" docTitle="Osedax braziliensis Fujiwara, Jimi, Sumida, Kawato & Kitazato, 2019, sp. n." docType="treatment" docUuid="7AD6CE45-4585-42C6-BB5E-22587BB2307F" docUuidSource="ZooBank" docVersion="5" lastPageNumber="62" masterDocId="FFE9FFA1C814550ABB08FF91DE49FFC7" masterDocTitle="New species of bone-eating worm Osedax from the abyssal South Atlantic Ocean (Annelida, Siboglinidae)" masterLastPageNumber="69" masterPageNumber="53" pageNumber="58" updateTime="1668166724051" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of bone-eating worm Osedax from the abyssal South Atlantic Ocean (Annelida, Siboglinidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fujiwara, Yoshihiro</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jimi, Naoto</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sumida, Paulo Y. G.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kawato, Masaru</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hiroshi Kitazato,</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>814</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>53</mods:start>
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<mods:end>69</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.814.28869</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.814.28869</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-814-53</mods:identifier>
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<mods:identifier type="ZBK">25C7041051D8445C99A1A7E158CFD507</mods:identifier>
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<mods:identifier type="ZooBank">25C7041051D8445C99A1A7E158CFD507</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154127468" LSID="urn:lsid:zoobank.org:act:7AD6CE45-4585-42C6-BB5E-22587BB2307F" httpUri="http://treatment.plazi.org/id/F1CB4DA529B8EEBE075E618E954F70D2" lastPageId="9" lastPageNumber="62" pageId="5" pageNumber="58">
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<subSubSection pageId="5" pageNumber="58" type="nomenclature">
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<paragraph pageId="5" pageNumber="58">
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<taxonomicName LSID="http://zoobank.org/7AD6CE45-4585-42C6-BB5E-22587BB2307F" class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax braziliensis" order="Sabellida" pageId="5" pageNumber="58" phylum="Annelida" rank="species" species="braziliensis">Osedax braziliensis</taxonomicName>
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<taxonomicNameLabel pageId="5" pageNumber="58">sp. n.</taxonomicNameLabel>
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Figures 3, 4 New Japanese name: Burajiru-honekuihanamushi
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="58" type="reference_group">
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<paragraph pageId="5" pageNumber="58">
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="5" pageNumber="58" phylum="Annelida" rank="genus">Osedax</taxonomicName>
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sp. nov.:
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<bibRefCitation author="Sumida, PY" journalOrPublisher="Bioinformatics" pageId="12" pageNumber="65" url="https://doi.org/10.1038/srep22139" year="2016">Sumida et al. 2016</bibRefCitation>
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: 1-6, figs 3-4, Table 1.
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="5" pageNumber="58" phylum="Annelida" rank="genus">Osedax</taxonomicName>
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:
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<bibRefCitation author="Alfaro-Lucas, JM" journalOrPublisher="Deep-Sea Research Part II: Topical Studies in Oceanography" pageId="9" pageNumber="62" pagination="4 - 12" title="Bone-eating Osedax worms (Annelida: Siboglinidae) regulate biodiversity of deep-sea whale-fall communities." url="https://doi.org/10.1016/j.dsr2.2017.04.011" volume="146" year="2017">Alfaro-Lucas et al. 2017</bibRefCitation>
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: 1-9, fig. 2B.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58" type="type material">
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<paragraph pageId="5" pageNumber="58">Type material.</paragraph>
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<paragraph lastPageId="6" lastPageNumber="59" pageId="5" pageNumber="58">
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Holotype: NMST-Pol H-685, trunk 14 mm long, 2 mm wide, female, 4,203 m depth, 26 April 2013, collected by YF, DDBJ No. LC381421,
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<pageBreakToken pageId="6" pageNumber="59" start="start">LC</pageBreakToken>
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381422, LC381424. Paratypes (14 specimens): NSMT-Pol P-686-690 and JAMSTEC-1130038806, 1130039105, 1130039113, 1130039116, 1130039146, 1130039163, 11 specimens, female, 24 and 26 April 2013, collected by YF, DDBJ No. LC381777, LC381766, LC381767, LC381768, LC381769, LC381771; JAMSTEC-1130057454, 1130057457, 1130057458, 3 specimens, male, 26 April 2013, collected by YF.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="60" type="type locality">
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<paragraph pageId="7" pageNumber="60">
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<pageBreakToken pageId="7" pageNumber="60" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph pageId="7" pageNumber="60">
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Paulo Ridge, Brazil, 4,204 m depth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="60" type="diagnosis">
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<paragraph pageId="7" pageNumber="60">Diagnosis.</paragraph>
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<paragraph pageId="7" pageNumber="60">Trunk length long. Gelatinous hemispherical tube encases trunk and base of palps. Yellow bump or patch present (absent in some specimens). Pinnules on inner margin of palps. Root lobulated without branching.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="61" pageId="7" pageNumber="60" type="description">
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<paragraph pageId="7" pageNumber="60">Description.</paragraph>
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<paragraph pageId="7" pageNumber="60">
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Genetic data (COI, 16S, and 18S) deposited in DDBJ (LC106303, LC381421, LC381422, LC381424, and LC381765-LC381787). Trunk length up to 22 mm, width at collar 0.5 mm, reddish purple while alive and whitish after fixation (Fig. 3
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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); gelatinous hemispherical tube encases trunk and base of palps, 1-2 mm thick, contains eggs and dwarf males (Fig. 3A, C). Prostomium whitish while alive, present at top of trunk. Yellow bump or patch present behind prostomium: this yellow bump or patch size varies among individuals, biggest bump reaches top of trunk, and is absent in some specimens (Fig. 3
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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). Crown consists of four palps; palps about 1.5 mm length, red colour while alive with two whitish stripes on the inner side, fused for about 30% of length; pinnules on inner margin of palps, about 50-250
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, 7-8 pinnules in transverse rows (Figs 3A, C, 4B, C). Oviduct free to base, adjoined to the trunk at opposite side of prostomium region, reaching up to 20-30% of palp length (Fig. 4C). Ovisac whitish;
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<normalizedToken originalValue="trunk–ovisac">trunk-ovisac</normalizedToken>
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junction about 15% of trunk length, light green while alive (Fig. 3B). Root lobulated without branching, yellow greenish while alive and whitish after fixation; intracellular symbiotic bacteria in root tissue (Fig. 3B). Eggs about 150
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<normalizedToken originalValue="µm">µm</normalizedToken>
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diameter (n = 20), whitish while alive (Fig. 3B).
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</paragraph>
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<caption pageId="7" pageNumber="60">
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<paragraph pageId="7" pageNumber="60">
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Figure 3. Photographs of unfixed
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax braziliensis" order="Sabellida" pageId="7" pageNumber="60" phylum="Annelida" rank="species" species="braziliensis">Osedax braziliensis</taxonomicName>
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sp. n. A Palps (pa), trunk (tr) and a gelatinous hemispherical tube (tu)B lateral view of palps (pa), trunk (tr), ovary (o), and root (r)C ventral view of palps (pa) with pinnules (pi) and trunk (tr). Dwarf males (m) inhabit a gelatinous tube (tu), and D ventro-lateral view of an individual possessing a yellow bump (b) present behind prostomium E Ventral view of holotype (NMST-Pol H-685) possessing a yellow patch (pt)F Ventral view of an individual without yellow bump.
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</paragraph>
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</caption>
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<paragraph pageId="8" pageNumber="61">
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<pageBreakToken pageId="8" pageNumber="61" start="start">Dwarf</pageBreakToken>
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male about 250
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<normalizedToken originalValue="µm">µm</normalizedToken>
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in length (n = 20), fusiform, whitish while alive, prostomium and pygidium rounded, no appendage organs; posterior hooks present, two pairs (5-7 hooks per bundle) arranged in three rows (Figs 3C, 4D).
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</paragraph>
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<caption pageId="8" pageNumber="61">
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<paragraph pageId="8" pageNumber="61">
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Figure 4. Scanning electron micrographs of
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax braziliensis" order="Sabellida" pageId="8" pageNumber="61" phylum="Annelida" rank="species" species="braziliensis">Osedax braziliensis</taxonomicName>
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sp. n. A Ventro-lateral view of bump (b)B lateral view of palps (pa) with pinnules (pi), and C dorsal view of palps (pa) and oviduct (od)D Dwarf male taken from the tube of a female.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="61" type="etymology">
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<paragraph pageId="8" pageNumber="61">Etymology.</paragraph>
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<paragraph pageId="8" pageNumber="61">This species is named after the type locality, Brazil. This name is an adjective used as a substantive in the genitive case.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="62" type="distribution">
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<paragraph pageId="9" pageNumber="62">
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<pageBreakToken pageId="9" pageNumber="62" start="start">Distribution</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="9" pageNumber="62">
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Only known from a whale carcass of the type locality.
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<normalizedToken originalValue="São">Sao</normalizedToken>
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Paulo Ridge, off Brazilian coast, 4,204 m depth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="62" type="phylogenetic analysis">
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<paragraph pageId="9" pageNumber="62">Phylogenetic analysis.</paragraph>
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<paragraph pageId="9" pageNumber="62">
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The final lengths of the aligned sequences were 1,004 bp (COI), 486 bp (16S), and 1,604 bp (18S). The phylogenetic position of
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<taxonomicName lsidName="O. braziliensis" pageId="9" pageNumber="62" rank="species" species="braziliensis">O. braziliensis</taxonomicName>
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sp. n. determined from our ML analysis recovered, with total support, a distinct species from that of all other
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
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species reported (Fig. 5). The six
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
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clades proposed by
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<bibRefCitation author="Rouse, GW" journalOrPublisher="Department of Zoology and Kewalo Marine Laboratory, University of Hawaii, Honolulu, Hawaii" pageId="11" pageNumber="64" url="https://doi.org/10.11646/zootaxa.4377.4.1" year="2018">Rouse et al. (2018)</bibRefCitation>
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were recovered. The phylogenetic analysis showed that
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<taxonomicName lsidName="O. braziliensis" pageId="9" pageNumber="62" rank="species" species="braziliensis">O. braziliensis</taxonomicName>
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sp. n. falls into Clade IV, and is a sister species of
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<taxonomicName lsidName="O. frankpressi" pageId="9" pageNumber="62" rank="species" species="frankpressi">O. frankpressi</taxonomicName>
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known from Monterey Bay at depths between 1,820 m and 2,898 m (Fig. 5).
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</paragraph>
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<caption pageId="9" pageNumber="62">
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<paragraph pageId="9" pageNumber="62">
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Figure 5. Phylogenetic placement of
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax braziliensis" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="braziliensis">Osedax braziliensis</taxonomicName>
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sp. n. based on nucleotide sequences on the concatenated COI, 16S, and 18S markers, using maximum likelihood. Scale bar represents 0.1 nucleotide substitutions per sequence position. Only bootstrap values greater than 50 are shown for each branch.
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax braziliensis" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="braziliensis">Osedax braziliensis</taxonomicName>
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sp. n. is boxed.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="62" type="remarks">
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<paragraph pageId="9" pageNumber="62">Remarks.</paragraph>
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<paragraph pageId="9" pageNumber="62">
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This species resembles
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax frankpressi" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="frankpressi">Osedax frankpressi</taxonomicName>
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in the pinnules distributed only at the inner margin of palps, lobulated root structure without branching, gelatinous hemispherical tube, and dwarf males (
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<bibRefCitation author="Rouse, GW" journalOrPublisher="Science" pageId="11" pageNumber="64" pagination="668 - 671" title="Osedax: bone-eating marine worms with dwarf males." url="https://doi.org/10.1126/science.1098650" volume="305" year="2004">Rouse et al. 2004</bibRefCitation>
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). However, it can be discriminated from
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<taxonomicName lsidName="O. frankpressi" pageId="9" pageNumber="62" rank="species" species="frankpressi">O. frankpressi</taxonomicName>
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by the presence of the yellow bump or patch behind the prostomium, trunk length, and genetic data. In
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<taxonomicName lsidName="O. braziliensis" pageId="9" pageNumber="62" rank="species" species="braziliensis">O. braziliensis</taxonomicName>
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sp. n., the yellow bump or patch was present in some specimens including holotype, and the trunk length is long (6-22 mm), whereas in
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<taxonomicName lsidName="O. frankpressi" pageId="9" pageNumber="62" rank="species" species="frankpressi">O. frankpressi</taxonomicName>
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, the bump or patch is absent in all specimens, and the trunk length is shorter (4.5 mm). COI genetic distances between
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<taxonomicName lsidName="O. braziliensis" pageId="9" pageNumber="62" rank="species" species="braziliensis">O. braziliensis</taxonomicName>
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sp. n. and
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<taxonomicName lsidName="O. frankpressi" pageId="9" pageNumber="62" rank="species" species="frankpressi">O. frankpressi</taxonomicName>
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are 0.111-0.117, which are greater than intraspecific values in
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<taxonomicName lsidName="O. braziliensis" pageId="9" pageNumber="62" rank="species" species="braziliensis">O. braziliensis</taxonomicName>
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sp. n. (0.001-0.006). Genetic distances between
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<taxonomicName lsidName="O. braziliensis" pageId="9" pageNumber="62" rank="species" species="braziliensis">O. braziliensis</taxonomicName>
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and the rest of the
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
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taxa for the COI ranged from 0.117 to 0.236 (Table 3).
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</paragraph>
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<caption ID-Table-UUID="90031946BEEB8B9151E2B1DE96BC07A4" httpUri="http://table.plazi.org/id/90031946BEEB8B9151E2B1DE96BC07A4" pageId="9" pageNumber="62">
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<paragraph pageId="9" pageNumber="62">
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Table 3. COI divergence values (Kimura 2 parameters) between
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
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species and OTUs.
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</paragraph>
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</caption>
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<paragraph pageId="9" pageNumber="62">
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<table pageId="9" pageNumber="62">
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<tr pageId="9" pageNumber="62">
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<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax frankpressi" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="frankpressi">Osedax frankpressi</taxonomicName>
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</td>
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</tr>
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<tr pageId="9" pageNumber="62">
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<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
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</td>
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</tr>
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<tr pageId="9" pageNumber="62">
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<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax tiburon" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="tiburon">Osedax tiburon</taxonomicName>
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</td>
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</tr>
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<tr pageId="9" pageNumber="62">
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<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
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<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
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</td>
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</tr>
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<tr pageId="9" pageNumber="62">
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<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax knutei" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="knutei">Osedax knutei</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax mucofloris" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="mucofloris">Osedax mucofloris</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax antarcticus" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="antarcticus">Osedax antarcticus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax talkovici" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="talkovici">Osedax talkovici</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax randyi" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="randyi">Osedax randyi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax crouchi" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="crouchi">Osedax crouchi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax sigridae" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="sigridae">Osedax sigridae</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax lehmani" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="lehmani">Osedax lehmani</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax ventana" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="ventana">Osedax ventana</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax roseus" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="roseus">Osedax roseus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax lonnyi" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="lonnyi">Osedax lonnyi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax braziliensis" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="braziliensis">Osedax braziliensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax rubiplumus" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="rubiplumus">Osedax rubiplumus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax rogersi" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="rogersi">Osedax rogersi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax packardorum" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="packardorum">Osedax packardorum</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax bryani" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="bryani">Osedax bryani</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax nordenskjoeldi" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="nordenskjoeldi">Osedax nordenskjoeldi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax westernflyer" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="westernflyer">Osedax westernflyer</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax ryderi" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="ryderi">Osedax ryderi</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax jabba" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="jabba">Osedax jabba</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax docricketts" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="docricketts">Osedax docricketts</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax priapus" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="priapus">Osedax priapus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="genus">Osedax</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax japonicus" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="japonicus">Osedax japonicus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="9" pageNumber="62">
|
||
<td colspan="1" pageId="9" pageNumber="62" rowspan="1">
|
||
<taxonomicName class="Polychaeta" family="Siboglinidae" genus="Osedax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Osedax deceptionensis" order="Sabellida" pageId="9" pageNumber="62" phylum="Annelida" rank="species" species="deceptionensis">Osedax deceptionensis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |