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<mods:title id="DBB4DC6B8F4F2EE75CAC9994FBE95CD7">Taxonomy of the genus Homalota Mannerheim in Korea (Coleoptera, Staphylinidae, Aleocharinae)</mods:title>
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<mods:namePart id="72801F23AEF078221F514B23705D86F1">Kim, Yoon-Ho</mods:namePart>
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<mods:namePart id="1FE7D93A935C28C87E8A6FF3AA4E70D6">Ahn, Kee-Jeong</mods:namePart>
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<mods:identifier id="6741A6CA16C5E51F009A53887F9EA3AB" type="DOI">http://dx.doi.org/10.3897/zookeys.447.7728</mods:identifier>
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<treatment id="B97C5EFF79C876232624D65D14862C9F" ID-GBIF-Taxon="152055880" LSID="urn:lsid:plazi:treatment:B97C5EFF79C876232624D65D14862C9F" httpUri="http://treatment.plazi.org/id/B97C5EFF79C876232624D65D14862C9F" lastPageNumber="111" pageId="2" pageNumber="111">
<subSubSection id="8840FDB4FF6387DB5339F70B4B2053FB" pageId="2" pageNumber="111" type="multiple">
<paragraph id="B66B1945373065F263DA92ACC3C70C5B" pageId="2" pageNumber="111">Taxon classification Animalia Coleoptera Staphylinidae</paragraph>
</subSubSection>
<subSubSection id="DAE1E4AC1C9D7D084F2C1312EF57253A" pageId="2" pageNumber="111" type="nomenclature">
<paragraph id="4372926F73B2C7C596D11A74C6E8C18B" pageId="2" pageNumber="111">
<taxonomicName id="8BB66F2876600089B30E1FB3A99D2983" ID-CoL="6MB28" authority="Likovsky, 1984" authorityName="Likovsky" authorityYear="1984" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota mikado" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="mikado">
Homalota mikado
<normalizedToken id="7996E1EA2A140E2CA43ED16DAAD49DD5" originalValue="Likovský">Likovsky</normalizedToken>
, 1984
</taxonomicName>
Figs 2, 15- 20
</paragraph>
</subSubSection>
<subSubSection id="6DA12BCB5C8F8471AF9364EC1ABE8756" pageId="2" pageNumber="111" type="reference_group">
<paragraph id="45BE32AB214146E9221AC655D3015F77" pageId="2" pageNumber="111">
<taxonomicName id="00162FE02E33F9F5B308E16E650F4FD4" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota mikado" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="mikado">Homalota mikado</taxonomicName>
<normalizedToken id="EC70BE8C1F8186FCAF4C3FDAAE2059AA" originalValue="Likovský">Likovsky</normalizedToken>
, 1984: 6 [Replacement name];
<bibRefCitation id="47E3B0BC91BF0C994F915B93E384897B" author="Smetana, A" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="7" pageNumber="116" pagination="353 - 494" title="Family Staphylinidae: Aleocharinae." volumeTitle="Catalogue of Palaearctic Coleoptera Volume 2 Hydrophiloidea-Histeroidea-Staphylinoidea" year="2004">Smetana 2004</bibRefCitation>
: 448.
</paragraph>
<paragraph id="D47491A20C5CCBB270AC4DA4AD08B799" pageId="2" pageNumber="111">
<taxonomicName id="2A89B4D95556D295A6B99A71C6901198" class="Insecta" family="Staphylinidae" genus="Epipeda" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epipeda granigera" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="granigera">Epipeda granigera</taxonomicName>
Sharp, 1888: 375 [Homonym].
</paragraph>
<paragraph id="ADFAF5CADBB4540B641C650332471FB9" pageId="2" pageNumber="111">
<taxonomicName id="5ED7CA50C53A95546EA0C7C277DDA912" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota granigera" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="granigera">Homalota granigera</taxonomicName>
:
<bibRefCitation id="AFF3CBED00C49299A337D7A808B64A07" author="Fenyes, A" journalOrPublisher="Archiv fuer Naturgeschichte (A)" pageId="6" pageNumber="115" pagination="45 - 55" title="H. Sauter's Formosa-Ausbeute. Aleocharinae." volume="80" year="1914">Fenyes 1914</bibRefCitation>
: 45.
</paragraph>
</subSubSection>
<subSubSection id="486EE2A99B32B674B97E331D874394B4" pageId="2" pageNumber="111" type="specimens examined">
<paragraph id="8F7DE1586B2ADC2B424911674E28987B" pageId="2" pageNumber="111">Specimens examined.</paragraph>
<paragraph id="412A3FB50CC92C7EF2434CA698194E26" pageId="2" pageNumber="111">
KOREA: Gangwon Prov., Pyeongchang-gun, Pyeongchang-eup, Noron-ri, Mt. Sambangsan, 13
<normalizedToken id="C9583EBD3572F4D89A181B6CFA12A78B" originalValue="VII">VII-</normalizedToken>
15 VIII 2001, KJ Ahn, SJ Park, CW Shin, ex FIT (2 exx., 1♀ on slide); Mt. Bokjusan, Seo-myeon, Cheolwon-gun, N
<geoCoordinate id="F08437225EA0C200EFAAE830E3E21BF1" direction="east" orientation="longitude" precision="1" value="38.143948">38°08'38.2&quot;E</geoCoordinate>
127°28'26.7&quot; 138 m, 25 IX 2005, YB Cho (2 exx.); Jeonnam Prov., Jangseong-gun, Mt. Naejangsan, Baekyangsa Area, 25 VI 2000, HJ Kim, ex sifting (3 exx., 1♂ on slide); Chungbuk Prov., Mt. Sokrisan, Beobjusa Temple, Naesokri-myeon, Boeun-gun, ex FIT, 1-31 V 2007, YB Cho,
<geoCoordinate id="C6A474FF1417033B63CBB50753F2AE95" direction="north" orientation="latitude" precision="1" value="36.539307">36°32'21.5&quot;N</geoCoordinate>
,
<geoCoordinate id="7F3EE6862358B9E86F0007BD8FDAAF37" direction="east" orientation="longitude" precision="1" value="127.83623">127°50'10.4&quot;E</geoCoordinate>
(1 ex.).
</paragraph>
</subSubSection>
<subSubSection id="205168345244BBBFF3B53ACBCDF2773E" pageId="2" pageNumber="111" type="description">
<paragraph id="579C56D36941768C19E4A5537212A9A3" pageId="2" pageNumber="111">Description.</paragraph>
<paragraph id="2C787AC404331CA4F23B2F003F951A5F" pageId="2" pageNumber="111">
Body (Fig. 2) length about 1.8-2.1 mm. Body dorsoventrally slightly flattened, widest at posterior margin of elytra; surface punctuate, subglossy, slightly pubescent; light brown to brown, abdominal tergite VI dark brown. Head. Subquadrate, slightly narrower than pronotum; eyes moderate in size, as long as tempora; head narrowed from behind of eyes to apical half of tempora but rarely narrowed to occipital construction; antennomere (Fig. 15) 4 slightly transverse, 5-10 transverse, slightly incrassate toward. Mouthparts. Labrum transverse, 8 pairs of macrosetae present, sensilla of antero-medial sensory area distinct, shallowly and width moderately emarginated;
<normalizedToken id="8EB47187812F896D2B7C1B05653C0FDD" originalValue="α-sensillum">α-sensillum</normalizedToken>
with a setose process,
<normalizedToken id="6BE146DFE2E3EC877D18AE407EFA4184" originalValue="β">β</normalizedToken>
and
<normalizedToken id="6BB1EEC60B99A31D09ADE42B51569BF8" originalValue="γ">γ</normalizedToken>
minute and conical,
<normalizedToken id="3E7E60E169DDC47E2AB68D84C8521902" originalValue="ε">ε</normalizedToken>
with a minute setose process, distinctly shorter than
<normalizedToken id="386E983C4F316F4E88AFA6605E8889A8" originalValue="α">α</normalizedToken>
, two lateral sensilla present on lateral margins of epipharynx, transverse row of sensory pores absent on basal region of epipharynx; right mandible with small median tooth, prostheca well developed, divided into 3 distinct area; maxillary palpomeres 2-3 dilated distally, 4 without small spine at apex; labium with ligula slender and elongated, bifid at apical third, almost as long as labial palpomere 1, labial palpus with two palpomeres, 1 longer than 2, two medial setae contiguous on prementum, side by side, median pseudopore field of prementum narrow and with pseudopores, mentum strongly emarginated in anterior margin. Thorax. Pronotum transverse, about 1.3 times wider than long, widest at apical third, surface pubescent, without distinct macrosetae; hypomeron broadly visible in lateral aspect; prosternum with a median knob; elytra (Fig. 16) wider than pronotum, postero-laterally sinuate; wings fully developed; mesoventrite (Fig. 17) without longitudinal carina, mesoventral process truncate at apex; metaventral process round at apex, longer than metaventral process; isthmus present; mesocoxae moderately separated; tarsomere 1 of front leg as long as 2, 1 slightly longer than 2 in middle and hind legs, without empodial setae between tarsal claws. Abdomen. Tergites
<normalizedToken id="69BBB5E54BCF5D20AD9C83C6769FB4A2" originalValue="IIIVI">III-VI</normalizedToken>
transversely impressed; tergite X with medial setal patch subquadrate, with 3 macrosetae on each side. Genitalia. Spermatheca (Fig. 18) simple and elongate at base, duct short; median lobe (Fig. 19) bulbous at base, apical process slender and elongate, longer than basal bulb, flagellum well sclerotized and short; paramere (Fig. 20) with apical lobe of paramerite with four setae, two setae longer than others, condylite subequal in length to apex of paramerite. Secondary sexual characteristics. Absent.
</paragraph>
<caption id="F8EB490C4C758C6AD4D3498024EE374C" pageId="2" pageNumber="111">
<paragraph id="E240DFC1DE9306746823FF4E1A0C98ED" pageId="2" pageNumber="111">
Figures 15-20.
<taxonomicName id="F649DBD151C6A517E1074BC0FC3B96D7" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota mikado" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="mikado">Homalota mikado</taxonomicName>
: 15 antenna 16 elytron, dorsal aspect 17 meso- and metaventrites, ventral aspect 18 spermatheca 19 median lobe, lateral aspect 20 paramere, lateral aspect. Scale bars 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7798DCE5E65889DDCF40AA0AB3A431BE" pageId="2" pageNumber="111" type="distribution">
<paragraph id="CFC3B5889B49E454BE47ECAE6DBDB9F5" pageId="2" pageNumber="111">Distribution.</paragraph>
<paragraph id="4DF791588C862BAF3767412A75B5345F" pageId="2" pageNumber="111">Korea (South), Japan, Taiwan.</paragraph>
</subSubSection>
<subSubSection id="2F9454359A050CFBA75571ED68F3369A" pageId="2" pageNumber="111" type="remarks">
<paragraph id="4B2D7C83C18B99D672E91BF961CB009B" pageId="2" pageNumber="111">Remarks.</paragraph>
<paragraph id="8F72439981C514962CBF059482F359B7" pageId="2" pageNumber="111">
The taxonomic position of
<taxonomicName id="4CCDE9F87F595188B80EC74B8F6F114D" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota mikado" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="mikado">Homalota mikado</taxonomicName>
is unclear, considering the numerous characters that distinguish it from other
<taxonomicName id="A2D869ADC57F498F5D4817D502A9869D" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">Homalota</taxonomicName>
species, including the type species
<taxonomicName id="A37022EDBB79B8841D9831CAC5EB3E29" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota plana" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="plana">Homalota plana</taxonomicName>
: body not subparallel-sided; surface with numerous large punctures; labrum with 8 pairs of macrosetae; ligula slender and elongate, mentum deeply emarginated in anterior margin; mesocoxae moderately separated. However, we are not sure what other aleocharine genus it may belong to. Therefore, the position of
<taxonomicName id="AABF2CCCBCAB4AAF0027205B5C292323" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota mikado" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="species" species="mikado">Homalota mikado</taxonomicName>
in
<taxonomicName id="BCBAA4396AC2CCB83C2F353E70936625" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Homalota" order="Coleoptera" pageId="2" pageNumber="111" phylum="Arthropoda" rank="genus">Homalota</taxonomicName>
is tentatively maintained here, pending further comprehensive research of this species.
</paragraph>
</subSubSection>
</treatment>
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