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<document ID-DOI="http://doi.org/10.5281/zenodo.4620130" ID-GBIF-Dataset="4d3d06a2-a773-4732-9a38-385951e4c647" ID-GBIF-Taxon="119316671" ID-ISSN="1175­5326" ID-Zenodo-Dep="4620130" approvalRequired="2" approvalRequired_for_document="1" approvalRequired_for_matCits="1" checkinTime="1459092556756" checkinUser="plazi" docAuthor="Greathead, D. J. &amp; Evenhuis, N. L." docDate="2001" docId="B526FE1FFFC8F128733CFDBE1A1CDAF9" docLanguage="en" docName="zt00014.pdf.imf" docOrigin="Zootaxa 14" docStyle="DocumentStyle:83E4DCE73C651764D97D3683740715A6.1:zzz_test.0000.journal_article" docStyleId="83E4DCE73C651764D97D3683740715A6" docStyleName="zzz_test.0000.journal_article" docStyleVersion="1" docTitle="Empidideicus socotrae Greathead &amp; Evenhuis 2001, sp.nov." docType="treatment" docVersion="9" lastPageId="3" lastPageNumber="4" masterDocId="491F8667FFCAF12B7234FF8A1E0CD867" masterDocTitle="Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae) from the island of Sokotra" masterLastPageNumber="11" masterPageNumber="1" pageId="2" pageNumber="3" updateTime="1643593816001" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae) from the island of Sokotra</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Greathead, D. J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Evenhuis, N. L.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2001</mods:date>
<mods:detail type="pubDate">
<mods:number>2001-11-26</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>14</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>11</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4620128</mods:identifier>
<mods:identifier type="GBIF-Dataset">4d3d06a2-a773-4732-9a38-385951e4c647</mods:identifier>
<mods:identifier type="ISSN">1175­5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">4620128</mods:identifier>
<mods:identifier type="ZooBank">39A75FD8-52B0-45A6-B39E-A0E2759F1B48</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4620130" ID-GBIF-Taxon="119316671" ID-Zenodo-Dep="4620130" LSID="urn:lsid:plazi:treatment:B526FE1FFFC8F128733CFDBE1A1CDAF9" httpUri="http://treatment.plazi.org/id/B526FE1FFFC8F128733CFDBE1A1CDAF9" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection box="[264,630,563,590]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph blockId="2.[264,630,563,590]" box="[264,630,563,590]" pageId="2" pageNumber="3">
<heading bold="true" box="[264,630,563,590]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis bold="true" box="[264,630,563,590]" pageId="2" pageNumber="3">
<taxonomicName authority="Greathead &amp; Evenhuis, 2001" authorityName="Greathead &amp; Evenhuis" authorityYear="2001" box="[264,537,564,590]" class="Insecta" family="Bombyliidae" genus="Empidideicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="socotrae" status="sp.nov.">
<emphasis bold="true" box="[264,537,564,590]" italics="true" pageId="2" pageNumber="3">Empidideicus socotrae</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[544,630,563,589]" pageId="2" pageNumber="3" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="diagnosis">
<paragraph blockId="2.[264,1325,639,1881]" pageId="2" pageNumber="3">
This new species belongs in the group of species (mostly undescribed) that is distinguished by having the mesonotum without contrasting pale and dark longitudinal vittae. Undescribed species of
<taxonomicName authorityName="Becker" authorityYear="1907" box="[548,708,719,744]" class="Insecta" family="Bombyliidae" genus="Empidideicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis box="[548,708,719,744]" italics="true" pageId="2" pageNumber="3">Empidideicus</emphasis>
</taxonomicName>
from
<collectingCountry box="[782,864,719,745]" name="Yemen" pageId="2" pageNumber="3">Yemen</collectingCountry>
and
<collectingCountry box="[923,995,719,745]" name="Oman" pageId="2" pageNumber="3">Oman</collectingCountry>
fit in this group and are similar to
<taxonomicName authorityName="Greathead &amp; Evenhuis" authorityYear="2001" box="[363,496,759,785]" class="Insecta" family="Bombyliidae" genus="Empidideicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="socotrae">
<emphasis box="[363,381,759,784]" italics="true" pageId="2" pageNumber="3">E</emphasis>
.
<emphasis box="[395,496,759,784]" italics="true" pageId="2" pageNumber="3">socotrae</emphasis>
</taxonomicName>
, but have the colouration of the mesonotum solid black (as opposed to the mostly brown colour in
<taxonomicName authorityName="Greathead &amp; Evenhuis" authorityYear="2001" box="[621,722,799,824]" class="Insecta" family="Bombyliidae" genus="Empidideicus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="socotrae">
<emphasis box="[621,722,799,824]" italics="true" pageId="2" pageNumber="3">socotrae</emphasis>
</taxonomicName>
with black restricted to the lateral areas of the mesonotum).
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph blockId="2.[264,1325,639,1881]" pageId="2" pageNumber="3">
<emphasis box="[312,400,879,904]" italics="true" pageId="2" pageNumber="3">Female</emphasis>
. Length:
<quantity box="[514,684,879,905]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" metricValueMax="1.08" metricValueMin="0.96" pageId="2" pageNumber="3" unit="mm" value="1.02" valueMax="1.08" valueMin="0.96">0.96­1.08 mm</quantity>
. Head longer than high, occiput well developed posteriorly; eyes dichoptic, separated at vertex by
<number box="[797,834,919,945]" pageId="2" pageNumber="3" value="1.5">1.5</number>
x distance between lateral ocelli; occiput and postgena black; front slightly depressed medially, white; face and tip of oral margin white; antennae (
<figureCitation box="[466,534,999,1025]" captionStart="FIGURES 1 ­ 4" captionStartId="3.[264,382,1574,1598]" captionTargetBox="[450,1120,740,1548]" captionTargetId="figure@3.[445,1125,732,1554]" captionTargetPageId="3" captionText="FIGURES 1 ­ 4. Empidideicus socotrae sp. nov. 1. Antenna; 2. Thorax and anterior part of abdomen, dorsal view; 3. Thorax and anterior part of abdomen, lateral view; 4. wing." httpUri="https://zenodo.org/record/155485/files/figure.png" pageId="2" pageNumber="3">
Fig.
<number box="[521,534,999,1025]" pageId="2" pageNumber="3" value="1.0">1</number>
</figureCitation>
) black; scape short, subtrapezoidal; pedicel subconical, wider than long; first flagellomere ovoid­triangular, length slightly longer than width; second flagellomere slightly shorter than first flagellomere, with small transparent apical style; mentum black; proboscis black, length slightly longer than head length; labrum sclerotised, stiff, pointed apically; palpus not evident.
</paragraph>
<paragraph blockId="2.[264,1325,639,1881]" pageId="2" pageNumber="3">
<emphasis box="[313,398,1199,1224]" italics="true" pageId="2" pageNumber="3">Thorax</emphasis>
(
<figureCitation box="[420,534,1199,1225]" captionStart="FIGURES 1 ­ 4" captionStartId="3.[264,382,1574,1598]" captionTargetBox="[450,1120,740,1548]" captionTargetId="figure@3.[445,1125,732,1554]" captionTargetPageId="3" captionText="FIGURES 1 ­ 4. Empidideicus socotrae sp. nov. 1. Antenna; 2. Thorax and anterior part of abdomen, dorsal view; 3. Thorax and anterior part of abdomen, lateral view; 4. wing." httpUri="https://zenodo.org/record/155485/files/figure.png" pageId="2" pageNumber="3">
Figs.
<number box="[492,534,1199,1225]" pageId="2" pageNumber="3" value="2.5" valueMax="3.0" valueMin="2.0">2­3</number>
</figureCitation>
). Mesonotum pale yellowish, uniformly brown dorsally, darker brown only above postalar calli; scutellum brownish with paler brown colour apically; pleura yellow except for brown on lower anepisternum, anterior portion of anepimeron, most of katepisternum, and lower portion of meron; coxae and legs yellow; halter stem and knob white.
</paragraph>
<paragraph blockId="2.[264,1325,639,1881]" pageId="2" pageNumber="3">
<emphasis box="[312,370,1399,1424]" italics="true" pageId="2" pageNumber="3">Wing</emphasis>
(
<figureCitation box="[389,464,1398,1424]" captionStart="FIGURES 1 ­ 4" captionStartId="3.[264,382,1574,1598]" captionTargetBox="[450,1120,740,1548]" captionTargetId="figure@3.[445,1125,732,1554]" captionTargetPageId="3" captionText="FIGURES 1 ­ 4. Empidideicus socotrae sp. nov. 1. Antenna; 2. Thorax and anterior part of abdomen, dorsal view; 3. Thorax and anterior part of abdomen, lateral view; 4. wing." httpUri="https://zenodo.org/record/155485/files/figure.png" pageId="2" pageNumber="3">
Fig.
<number box="[446,464,1398,1424]" pageId="2" pageNumber="3" value="4.0">4</number>
</figureCitation>
). Subhyaline; veins brown, well sclerotised except vein separating basal cells, M
<number box="[361,372,1450,1469]" pageId="2" pageNumber="3" value="1.0">1</number>
, M
<number box="[415,426,1450,1469]" pageId="2" pageNumber="3" value="2.0">2</number>
, M
<number box="[469,477,1450,1469]" pageId="2" pageNumber="3" value="1.0">1</number>
+
<number box="[490,502,1450,1469]" pageId="2" pageNumber="3" value="2.0">2</number>
, and CuA
<number box="[627,638,1450,1469]" pageId="2" pageNumber="3" value="1.0">1</number>
; costa ends slightly beyond end of R
<number box="[1089,1100,1450,1469]" pageId="2" pageNumber="3" value="4.0">4</number>
+
<number box="[1112,1122,1450,1469]" pageId="2" pageNumber="3" value="5.0">5</number>
; vein Sc incomplete; Rs evanescent at connection with R
<number box="[759,770,1495,1514]" pageId="2" pageNumber="3" value="1.0">1</number>
; R
<number box="[806,817,1495,1514]" pageId="2" pageNumber="3" value="4.0">4</number>
+
<number box="[829,839,1495,1514]" pageId="2" pageNumber="3" value="5.0">5</number>
slightly curved to wing margin; vein M
<number box="[1312,1323,1495,1514]" pageId="2" pageNumber="3" value="1.0">1</number>
curved toward wing margin; M
<number box="[644,655,1538,1557]" pageId="2" pageNumber="3" value="2.0">2</number>
slightly sinuous to wing margin; length of M
<number box="[1213,1224,1538,1557]" pageId="2" pageNumber="3" value="2.0">2</number>
slightly longer than M
<number box="[438,446,1583,1602]" pageId="2" pageNumber="3" value="1.0">1</number>
+
<number box="[459,471,1583,1602]" pageId="2" pageNumber="3" value="2.0">2</number>
; crossvein dm­cu closing cell dm absent; A
<number box="[1009,1020,1583,1602]" pageId="2" pageNumber="3" value="1.0">1</number>
slightly sinuous to wing margin; fringe of hair on posterior margin of wing well developed.
</paragraph>
<paragraph blockId="2.[264,1325,639,1881]" pageId="2" pageNumber="3">
<emphasis box="[312,422,1655,1680]" italics="true" pageId="2" pageNumber="3">Abdomen</emphasis>
. Dorsum brown, concolorous with central mesonotal colour, posterior margins of tergites pale yellow to white, dorsal brown colour of tergites gradating to pale yellow laterally; tergites with sparsely scattered minute hairs; venter yellowish white.
</paragraph>
<paragraph blockId="2.[264,1325,639,1881]" lastBlockId="3.[264,1324,284,670]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis box="[312,422,1775,1800]" italics="true" pageId="2" pageNumber="3">Genitalia</emphasis>
(
<figureCitation box="[438,524,1775,1801]" captionStart="FIGURES 11 ­ 12" captionStartId="5.[264,383,1371,1395]" captionTargetBox="[609,1011,524,1337]" captionTargetId="figure@5.[596,1012,523,1339]" captionTargetPageId="5" captionText="FIGURES 11 ­ 12. Empidideicus spermatheca and ducts. 11. E. socotrae sp. nov.; 12. E. variegatus sp. nov." httpUri="https://zenodo.org/record/155487/files/figure.png" pageId="2" pageNumber="3">
Fig.
<number box="[493,524,1775,1801]" pageId="2" pageNumber="3" value="11.0">11</number>
</figureCitation>
). Spermatheca spheroid­ellipsoid with large indentation, length subequal to height of spermatheca; spermatheca with basal conical area leading to apical duct; apical spermathecal duct ca.
<number box="[612,649,1855,1881]" pageId="2" pageNumber="3" value="1.5">1.5</number>
x length of spermatheca, thin, transparent; sperm pump subequal in length to apical duct, transparent, not sclerotised apically or basally; basal duct slightly sclerotised brownish, thin, length slightly less than apical duct.
</paragraph>
<paragraph blockId="3.[264,1324,284,670]" box="[312,510,364,390]" pageId="3" pageNumber="4">
<emphasis box="[312,372,364,389]" italics="true" pageId="3" pageNumber="4">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="discussion">
<paragraph blockId="3.[264,1324,284,670]" pageId="3" pageNumber="4">
<emphasis box="[312,414,404,429]" italics="true" pageId="3" pageNumber="4">Remarks</emphasis>
. Upon wetting, a mesonotal pattern becomes apparent in which there is a short longitudinal median dark stripe from the midpoint of the mesonotum to the prescutellar area and a dark spot of brown lateral to the median stripe leading toward the postalar callus; otherwise the rest of the mesonotum is a pale brownish yellow colour.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="materials_examined">
<paragraph blockId="3.[264,1324,284,670]" pageId="3" pageNumber="4">
<emphasis box="[312,376,564,589]" italics="true" pageId="3" pageNumber="4">Types</emphasis>
.
<materialsCitation ID-GBIF-Occurrence="3056542302" collectingDate="2000-02-27" collectingDateMax="2000-03-01" collectingDateMin="2000-02-27" collectionCode="NMSA, NLE" collectorName="K. Stastny &amp; V. Bejcek" latitude="13.0" location="SOKOTRA" longLatPrecision="41492" longitude="54.0" pageId="3" pageNumber="4" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
<typeStatus box="[393,502,564,590]" pageId="3" pageNumber="4">Holotype</typeStatus>
<specimenCount box="[511,591,564,590]" pageId="3" pageNumber="4" type="female">female</specimenCount>
and
<specimenCount box="[650,752,564,590]" pageId="3" pageNumber="4" type="female">
<number box="[650,665,564,590]" pageId="3" pageNumber="4" value="1.0">1</number>
female
</specimenCount>
<typeStatus box="[760,861,564,590]" pageId="3" pageNumber="4">paratype</typeStatus>
from
<taxonomicName authority=": Noghed" authorityName="Noghed" box="[935,1194,563,590]" class="Insecta" family="Diptera (awaiting allocation)" genus="Sokotra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[935,1082,563,589]" pageId="3" pageNumber="4">
<location LSID="urn:lsid:plazi:treatment:B526FE1FFFC8F128733CFDBE1A1CDAF9:385019D2FFC9F1287193FDB91A36DA2A" box="[935,1082,563,589]" latitude="13.0" longLatPrecision="41492" longitude="54.0" name="SOKOTRA" pageId="3" pageNumber="4">SOKOTRA</location>
</emphasis>
:
<location LSID="urn:lsid:plazi:treatment:B526FE1FFFC8F128733CFDBE1A1CDAF9:385019D2FFC9F128767EFDBE1AA6DA29" box="[1098,1194,564,590]" latitude="13.0" longLatPrecision="41492" longitude="54.0" name="Noghed" pageId="3" pageNumber="4">Noghed</location>
</taxonomicName>
, cca.
<geoCoordinate box="[1261,1323,564,590]" degrees="13" direction="north" orientation="latitude" pageId="3" pageNumber="4" precision="55555" value="13.0">
<number box="[1261,1289,564,590]" pageId="3" pageNumber="4" value="13.0">13</number>
°N
</geoCoordinate>
<geoCoordinate box="[264,326,604,630]" degrees="54" direction="east" orientation="longitude" pageId="3" pageNumber="4" precision="55555" value="54.0">
<number box="[264,296,604,630]" pageId="3" pageNumber="4" value="54.0">54</number>
°E
</geoCoordinate>
,
<date box="[338,520,604,630]" pageId="3" pageNumber="4" value="2000-02-27" valueMax="2000-03-01" valueMin="2000-02-27">
<collectingDate box="[338,520,604,630]" pageId="3" pageNumber="4" value="2000-02-27" valueMax="2000-03-01" valueMin="2000-02-27">
<number box="[338,370,604,630]" pageId="3" pageNumber="4" value="27.0">27</number>
.ii–
<number box="[408,418,604,630]" pageId="3" pageNumber="4" value="1.0">1</number>
.iii.
<number box="[453,520,604,630]" pageId="3" pageNumber="4" value="2000.0">2000</number>
</collectingDate>
</date>
,
<collectorName box="[533,656,604,630]" pageId="3" pageNumber="4">K. Stástny</collectorName>
&amp;
<collectorName box="[693,811,604,630]" pageId="3" pageNumber="4">V. Bejcek</collectorName>
. Holotype in
<collectionCode box="[968,1057,604,630]" pageId="3" pageNumber="4">NMSA</collectionCode>
. Paratype in
<collectionCode box="[1208,1269,604,630]" pageId="3" pageNumber="4">NLE</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[312,1040,644,670]" pageId="3" pageNumber="4" type="etymology">
<paragraph blockId="3.[264,1324,284,670]" box="[312,1040,644,670]" pageId="3" pageNumber="4">
<emphasis box="[312,435,644,669]" italics="true" pageId="3" pageNumber="4">Etymology</emphasis>
. The specific epithet derives from the
<typeStatus box="[888,939,644,670]" pageId="3" pageNumber="4">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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