276 lines
23 KiB
XML
276 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1569" ID-GBIF-Dataset="226292c9-1f85-4f2f-8062-2276d90f43d1" ID-PMC="PMC3252762" ID-Pensoft-Pub="1313-2970-130-379" ID-PubMed="22259290" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-379" ModsDocOrigin="ZooKeys 130" ModsDocTitle="Lytopylus Förster (Hymenoptera, Braconidae, Agathidinae) species from Costa Rica, with an emphasis on specimens reared from caterpillars in Area de Conservación Guanacaste" checkinTime="1451249852228" checkinUser="pensoft" docAuthor="Sharkey, Michael J., Clutts, Stephanie, Tucker, Erika M., Janzen, Daniel, Hallwachs, Winnie, Dapkey, Tanya & Smith, M. Alex" docDate="2011" docId="793A557C08B0FA1BB82542E3DB90F44D" docLanguage="en" docName="ZooKeys 130: 379-419" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1569" docTitle="Lytopylus jessiehillae Sharkey, sp. n." docType="treatment" docVersion="4" lastPageNumber="392" masterDocId="FF89665BD33AF371227FFFCC1631614B" masterDocTitle="Lytopylus Foerster (Hymenoptera, Braconidae, Agathidinae) species from Costa Rica, with an emphasis on specimens reared from caterpillars in Area de Conservacion Guanacaste" masterLastPageNumber="419" masterPageNumber="379" pageNumber="389" updateTime="1668152276762" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Lytopylus Foerster (Hymenoptera, Braconidae, Agathidinae) species from Costa Rica, with an emphasis on specimens reared from caterpillars in Area de Conservacion Guanacaste</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Sharkey, Michael J.</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Clutts, Stephanie</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Tucker, Erika M.</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Janzen, Daniel</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Hallwachs, Winnie</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Dapkey, Tanya</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Smith, M. Alex</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2011</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>130</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>379</mods:start>
|
|
<mods:end>419</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.130.1569</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.130.1569</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-130-379</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="152030962" LSID="urn:lsid:zoobank.org:act:5ECF221A-A5D7-4923-AA1E-33BB0E1DCF34" httpUri="http://treatment.plazi.org/id/793A557C08B0FA1BB82542E3DB90F44D" lastPageId="13" lastPageNumber="392" pageId="10" pageNumber="389">
|
|
<subSubSection pageId="10" pageNumber="389" type="nomenclature">
|
|
<paragraph pageId="10" pageNumber="389">
|
|
<taxonomicName LSID="urn:lsid:zoobank.org:act:5ECF221A-A5D7-4923-AA1E-33BB0E1DCF34" authority="Sharkey" class="Insecta" family="Braconidae" genus="Lytopylus" higherTaxonomySource="GBIF" kingdom="Metazoa" lsidName="Lytopylus jessiehillae" order="Hymenoptera" pageId="10" pageNumber="389" phylum="Arthropoda" rank="species" species="jessiehillae">Lytopylus jessiehillae Sharkey</taxonomicName>
|
|
<taxonomicNameLabel pageId="10" pageNumber="389">sp. n.</taxonomicNameLabel>
|
|
Figs 1415
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="11" pageNumber="390" type="description">
|
|
<paragraph pageId="11" pageNumber="390">
|
|
<pageBreakToken pageId="11" pageNumber="390" start="start">Description</pageBreakToken>
|
|
.
|
|
</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">Body length 7.0 - 8.3 mm. Ovipositor length 7.2 - 7.7 mm. Gena rounded or with an obtuse angle posterolaterally. Longitudinal groove on interantennal prominence absent. Protuberances on occiput absent. Propodeum with carinae forming areolae, median areola rounded anteriorly. Notauli well-impressed, smooth without crenulae, or with one or two crenulae restricted to extreme anterior apex along border of mesoscutum. Posterior margin of syntergum 2+3 convex, covering most of terminal terga. Median syntergite 2 + 3 longitudinally striate except apical transverse lobe rugose. Forewing mostly or entirely infuscate. Color as in Figs 14, 15. Color variation: Most commonly the mesosoma varies from completely black, to black except for the propodeum and mesothorax which are yellow or orange in whole or in part. Rarely, two specimens, the mesosoma is almost entirely pale. The hind femur varies from entirely melanic to mostly pale. Pale parts of the body vary from yellow to orange.</paragraph>
|
|
<caption pageId="11" pageNumber="390">
|
|
<paragraph pageId="11" pageNumber="390">
|
|
Figure 14.
|
|
<taxonomicName class="Insecta" family="Braconidae" genus="Lytopylus" higherTaxonomySource="GBIF" kingdom="Metazoa" lsidName="Lytopylus jessiehillae" order="Hymenoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="species" species="jessiehillae">Lytopylus jessiehillae</taxonomicName>
|
|
sp. n. a lateral habitus b dorsal habitus c wings.
|
|
</paragraph>
|
|
</caption>
|
|
<caption pageId="11" pageNumber="390">
|
|
<paragraph pageId="11" pageNumber="390">
|
|
Figure 15.
|
|
<taxonomicName class="Insecta" family="Braconidae" genus="Lytopylus" higherTaxonomySource="GBIF" kingdom="Metazoa" lsidName="Lytopylus jessiehillae" order="Hymenoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="species" species="jessiehillae">Lytopylus jessiehillae</taxonomicName>
|
|
sp. n. a anterior head b dorsal propodeum c lateral head and mesosoma d dorsal head and mesosoma e dorsal metasoma.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="11" pageNumber="390" type="molecular data">
|
|
<paragraph pageId="11" pageNumber="390">Molecular data.</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">BOLD process ID/Janzen parasitoid voucher/GenBank accession:</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASAG139-07/DHJPAR0015453/JN034705</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYE376-08/DHJPAR0028139/JN034704</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASAG138-07/DHJPAR0015452/JN034704</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYF079-09/DHJPAR0028317/JN034702</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYE1603-09/DHJPAR0036692/JN034701</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYE1631-09/DHJPAR0036720/JN34700</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYF080-09/DHJPAR0028318/JN034699</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYF019-09/DHJPAR0028257/JN034698</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYF078-09/DHJPAR0028316/JN034697</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYF017-09/DHJPAR0028255/JN034696</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYF018-09/DHJPAR0028256/JN034695</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYF081-09/DHJPAR0028319/JN034694</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">ASHYF068-09/DHJPAR0028306/JN034693</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="11" pageNumber="390" type="distribution">
|
|
<paragraph pageId="11" pageNumber="390">Distribution.</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">Guanacaste Province and Heredia Province, Costa Rica. Click here for a distribution map.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="12" lastPageNumber="391" pageId="11" pageNumber="390" type="biology">
|
|
<paragraph pageId="11" pageNumber="390">Biology.</paragraph>
|
|
<paragraph pageId="11" pageNumber="390">
|
|
This wasp is the only species of
|
|
<taxonomicName class="Insecta" family="Braconidae" genus="Lytopylus" higherTaxonomySource="GBIF" kingdom="Metazoa" lsidName="Lytopylus" order="Hymenoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Lytopylus</taxonomicName>
|
|
found to date using
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
(
|
|
<taxonomicName family="Thyrididae" lsidName="" pageId="11" pageNumber="390" rank="family">Thyrididae</taxonomicName>
|
|
) as its host caterpillar (22 wasp rearings from 3,500+ rearings of about 20
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
species). These wasp rearings are almost entirely from three species of rain forest
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
(89
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
Janzen12 feeding exclusively on
|
|
<taxonomicName class="Magnoliopsida" family="Phyllanthaceae" genus="Hieronyma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hieronyma oblonga" order="Malpighiales" pageId="11" pageNumber="390" phylum="Tracheophyta" rank="species" species="oblonga">Hieronyma oblonga</taxonomicName>
|
|
; 1,241
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
Janzen06 feeding on many plant families; 182
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
Janzen35 feeding on
|
|
<taxonomicName genus="Annonaceae" lsidName="Annonaceae" pageId="11" pageNumber="390" rank="genus">Annonaceae</taxonomicName>
|
|
) in dry forest and dry-rain forest intergrades. Additionally, a single
|
|
<taxonomicName class="Insecta" family="Braconidae" genus="Lytopylus" higherTaxonomySource="GBIF" kingdom="Metazoa" lsidName="Lytopylus jessiehillae" order="Hymenoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="species" species="jessiehillae">Lytopylus jessiehillae</taxonomicName>
|
|
has been reared from
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Collinsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Collinsa ferreiceps" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="species" species="ferreiceps">Collinsa ferreiceps</taxonomicName>
|
|
(
|
|
<taxonomicName family="Thyrididae" lsidName="" pageId="11" pageNumber="390" rank="family">Thyrididae</taxonomicName>
|
|
) feeding on the rain forest tree
|
|
<taxonomicName class="Magnoliopsida" family="Elaeocarpaceae" genus="Sloanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sloanea faginea" order="Oxalidales" pageId="11" pageNumber="390" phylum="Tracheophyta" rank="species" species="faginea">Sloanea faginea</taxonomicName>
|
|
(
|
|
<taxonomicName genus="Elaeocarpaceae" lsidName="Elaeocarpaceae" pageId="11" pageNumber="390" rank="genus">Elaeocarpaceae</taxonomicName>
|
|
; 73 rearings); the adult of this moth is very similar in body size and wing shape to
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Zeuzerodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zeuzerodes caenosa" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="species" species="caenosa">Zeuzerodes caenosa</taxonomicName>
|
|
(and very different from
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
) but the caterpillar is very similar to that of
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
in body size and leaf roll type. Furthermore, it is the only agathidine reared from the 73 rearings of
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Collinsa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Collinsa" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Collinsa</taxonomicName>
|
|
on
|
|
<taxonomicName class="Magnoliopsida" family="Elaeocarpaceae" genus="Sloanea" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Sloanea" order="Oxalidales" pageId="11" pageNumber="390" phylum="Tracheophyta" rank="genus">Sloanea</taxonomicName>
|
|
, and I (DHJ) feel that it is very likely to be an
|
|
<normalizedToken originalValue="“abnormal”">"abnormal"</normalizedToken>
|
|
host record.
|
|
</paragraph>
|
|
<paragraph lastPageId="12" lastPageNumber="391" pageId="11" pageNumber="390">
|
|
Only one other agathidine has been reared from the 3,500+ rearings of
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
, a single specimen of
|
|
<taxonomicName class="Insecta" family="Braconidae" genus="Amputoearinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amputoearinus niger" order="Hymenoptera" pageId="11" pageNumber="390" phylum="Arthropoda" rank="species" species="niger">Amputoearinus niger</taxonomicName>
|
|
from the rain forest
|
|
<taxonomicName class="Magnoliopsida" family="Erythropalaceae" genus="Heisteria" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Heisteria concinna" order="Santalales" pageId="11" pageNumber="390" phylum="Tracheophyta" rank="species" species="concinna">Heisteria concinna</taxonomicName>
|
|
(
|
|
<taxonomicName genus="Olacaceae" lsidName="Olacaceae" pageId="12" pageNumber="391" rank="genus">
|
|
<pageBreakToken pageId="12" pageNumber="391" start="start">Olacaceae</pageBreakToken>
|
|
</taxonomicName>
|
|
) being fed on by
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="12" pageNumber="391" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
Janzen06.
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="12" pageNumber="391" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
in ACG is, however, also used by at least 7 species of
|
|
<taxonomicName family="Tachinidae" lsidName="" pageId="12" pageNumber="391" rank="family">Tachinidae</taxonomicName>
|
|
and
|
|
<taxonomicName lsidName="" pageId="12" pageNumber="391" rank="subFamily" subFamily="Microgastrinae">Microgastrinae</taxonomicName>
|
|
braconids.
|
|
</paragraph>
|
|
<paragraph pageId="12" pageNumber="391">
|
|
<taxonomicName class="Insecta" family="Thyrididae" genus="Dysodia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysodia" order="Lepidoptera" pageId="12" pageNumber="391" phylum="Arthropoda" rank="genus">Dysodia</taxonomicName>
|
|
caterpillars construct a close-fitting conical-tubular shelter by cutting a lengthy curved slit in the food plant leaf and rolling that partly detached segment up around its base. The caterpillar then eats the inner parts of the leaf roll. The ovipositing wasp must then penetrate 1-3 layers of leaf and the caterpillar can only move forward and backward to escape. The caterpillar remains in one leaf roll for many days, pushing or ejecting fecal pellets out the "front door". The distinctive leaf rolls are easily recognized by the researcher, and presumably by a searching wasp as well, but many are empty because the caterpillar has moved on or been taken by a predator. The prepupal caterpillar (Fig. 16) spins a tough silk cocoon between two overlapping leaves (or in the leaf roll). At this time the wasp larva feeds heavily and within 2-3 days has consumed the caterpillar innards and ruptures the caterpillar cuticle to emerge into the pupal chamber and spin a tough white cocoon with the caterpillar head capsule and remaining pelt scrunched onto one end (Fig. 17).
|
|
</paragraph>
|
|
<caption pageId="12" pageNumber="391">
|
|
<paragraph pageId="12" pageNumber="391">
|
|
Figure 16.
|
|
<taxonomicName class="Insecta" family="Braconidae" genus="Lytopylus" higherTaxonomySource="GBIF" kingdom="Metazoa" lsidName="Lytopylus jessiehillae" order="Hymenoptera" pageId="12" pageNumber="391" phylum="Arthropoda" rank="species" species="jessiehillae">Lytopylus jessiehillae</taxonomicName>
|
|
prepupal host caterpillar.
|
|
</paragraph>
|
|
</caption>
|
|
<caption pageId="12" pageNumber="391">
|
|
<paragraph pageId="12" pageNumber="391">
|
|
Figure 17.
|
|
<taxonomicName class="Insecta" family="Braconidae" genus="Lytopylus" higherTaxonomySource="GBIF" kingdom="Metazoa" lsidName="Lytopylus jessiehillae" order="Hymenoptera" pageId="12" pageNumber="391" phylum="Arthropoda" rank="species" species="jessiehillae">Lytopylus jessiehillae</taxonomicName>
|
|
cocoon.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="12" pageNumber="391" type="etymology">
|
|
<paragraph pageId="12" pageNumber="391">Etymology.</paragraph>
|
|
<paragraph pageId="12" pageNumber="391">Named in honor of Jessie Hill of Hawaii, who has enthusiastically supported the conservation and biodiversity inventory of the ACG forest occupied by this parasitoid wasp.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="13" lastPageNumber="392" pageId="12" pageNumber="391" type="material examined">
|
|
<paragraph pageId="12" pageNumber="391">Material examined.</paragraph>
|
|
<paragraph pageId="12" pageNumber="391">
|
|
Holotype: ♀, H7093 (DHJPAR0028316 - this is a short sequence) Costa Rica: Guanacaste: Area de
|
|
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
|
|
Guanacaste: Sector Pitilla: Loaiciga, 7.ix.2008,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="11.019">11.019N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.4134">85.4134W</geoCoordinate>
|
|
, 445m [AEI].
|
|
</paragraph>
|
|
<paragraph pageId="13" pageNumber="392">
|
|
<pageBreakToken pageId="13" pageNumber="392" start="start">Paratypes</pageBreakToken>
|
|
[AEI, HIC, INBio]: Costa Rica: Guanacaste: Area de
|
|
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
|
|
Guanacaste:Sector Pitilla: Loaiciga,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="11.019">11.019N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.4134">85.4134W</geoCoordinate>
|
|
, 445m: ♀ H7090 (DHJPAR0028305) 7.ix.2008, ♀ H7091 (DHJPAR0028257) 5.ix.2008, ♀ H7098 (DHJPAR0028319) 4.ix.2008, ♀ H7063 (DHJPAR0028306) 4.ix.2008, ♀ H7083 (DHJPAR0028309) 9.iv.2008. ♂ H7085 (DHJPAR0028313) 6.ix.2008, ♂ H7096 (DHJPAR0028255) 4.ix.2008, ♂ H7089 (DHJPAR0028256) 28.viii.2008. Sendero Cuestona,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="5" value="10.9945">10.9945N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.4146">85.4146W</geoCoordinate>
|
|
, 640m: ♀ H8398 (DHJPAR0015455) 9.iv.2004, ♂ H8399 (DHJPAR0015454) 7.iv.2004. Sector Cacao: Sendero Toma Agua,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.928">10.928N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.4668">85.4668W</geoCoordinate>
|
|
, 1140m: ♀ 4.iv.2001: H7088 (DHJPAR0015453), ♀ H7086 (DHJPAR0015452), ♂ H8396 (DHJPAR0015451). Sector Del Oro: Guacimos,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="11.014">11.014N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.4749">85.4749W</geoCoordinate>
|
|
, 380m: ♂ H7087 (DHJPAR0036692) 3.viii.2009. Sector Mundo Nuevo: Vado Ocotea,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="5" value="10.7638">10.7638N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.3784">85.3784W</geoCoordinate>
|
|
, 565m: ♀ H7095 (DHJPAR0028139) 30.vii.2008. Vado Zanja Tapada,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="5" value="10.7648">10.7648N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.3845">85.3845W</geoCoordinate>
|
|
, 550m: ♀ H7094 (DHJPAR0040216) 4.vii.2010, ♀ H7099 (DHJPAR0040224) 3.vii.2010. Porton Rivas,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="5" value="10.7586">10.7586N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.3727">85.3727W</geoCoordinate>
|
|
, 570m: ♀ H6410 (DHJPAR0028318) 9.vii.2008, ♂ H7092 (DHJPAR0028317) 6.vii.2008. Punta Plancha,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="5" value="10.7416">10.7416N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.4273">85.4273W</geoCoordinate>
|
|
, 420m: ♀ H7097 (DHJPAR0040226) 1.vii.2010. Area de
|
|
<normalizedToken originalValue="Conservación">Conservacion</normalizedToken>
|
|
Guanacaste: Sector Rincon Rain Forest: Camino Albergue Oscar,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="5" value="10.8774">10.8774N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-85.3236">85.3236W</geoCoordinate>
|
|
, 560m: ♀ H7084 (DHJPAR0036720) 1.ix.2009.
|
|
</paragraph>
|
|
<paragraph pageId="13" pageNumber="392">
|
|
Costa Rica: Heredia: Est. Biol. La Selva,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="55" value="10.433">10.433N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="55" value="-84.017">84.017W</geoCoordinate>
|
|
, 150m: ♀ H8397 vii.1993.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |