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<mods:title id="48272DC3284530BCFBA4EC77FA65A7F1">A review of the coelidiine leafhopper genus Mahellus (Hemiptera: Cicadellidae: Coelidiinae) with description of two new species from the Oriental region</mods:title>
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Key to species of
<taxonomicName id="38D24D23FFA37A0BFEB2F9BCFE26AD3D" ID-CoL="8HLWS" box="[365,479,1551,1577]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="CDA6EAB2FFA37A0BFEB2F9BCFE26AD3D" bold="true" box="[365,479,1551,1577]" italics="true" pageId="1" pageNumber="272">Mahellus</emphasis>
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<paragraph id="FF6D36A0FFA37A0BFF48F9E6FA65AD78" blockId="1.[151,1436,1621,1908]" box="[151,1436,1621,1644]" pageId="1" pageNumber="272">
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1. Subgenital plate with posterior spine-like projection (
<figureCitation id="67E92A25FFA37A0BFD3FF9E6FCD0AD7F" box="[736,809,1621,1643]" captionStart="FIGURES 40 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="1" pageNumber="272">Fig. 44</figureCitation>
).................................... ..
<taxonomicName id="38D24D23FFA37A0BFB34F9E6FAB0AD7F" ID-CoL="8JWDT" box="[1259,1353,1621,1643]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="ungulatus" status="sp. nov.">
<emphasis id="CDA6EAB2FFA37A0BFB34F9E6FAB0AD7F" box="[1259,1353,1621,1643]" italics="true" pageId="1" pageNumber="272">ungulatus</emphasis>
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<emphasis id="CDA6EAB2FFA37A0BFA90F9E5FA65AD78" bold="true" box="[1359,1436,1622,1644]" pageId="1" pageNumber="272">sp. nov.</emphasis>
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</paragraph>
<paragraph id="FF6D36A0FFA37A0BFF48F9C7FA65AD9D" blockId="1.[151,1436,1621,1908]" box="[151,1436,1651,1673]" pageId="1" pageNumber="272">
<keyLead id="44239595FFA37A0BFF48F9C7FA65AD9D" box="[151,1436,1651,1673]" pageId="1" pageNumber="272">
- Subgenital plate without posterior spine-like projection (
<figureCitation id="67E92A25FFA37A0BFD20F9C0FCB0AD9D" box="[767,841,1651,1673]" captionStart="FIGURES 18 24" captionStartId="4.[151,264,1953,1976]" captionTargetBox="[205,1385,222,1919]" captionTargetId="figure@4.[178,1409,193,1932]" captionTargetPageId="4" captionText="FIGURES 18 24. Mahellus distanti, male genitalia. 18. Pygofer, lateral view; 19. Connective, dorsal view; 20. Style, dorsal view; 21, Subgenital plate, ventral view; 22. Aedeagus and dorsal connective, dorsal view; 23. Aedeagal shaft apex, dorsal view; 24. Aedeagus and dorsal connective, lateral view." httpUri="https://zenodo.org/record/569767/files/figure.png" pageId="1" pageNumber="272">Figs 21</figureCitation>
,
<figureCitation id="67E92A25FFA37A0BFC8CF9C0FC97AD9D" box="[851,878,1651,1673]" captionStart="FIGURES 32 39" captionStartId="6.[151,264,1923,1946]" captionTargetBox="[208,1351,202,1865]" captionTargetId="figure@6.[199,1368,193,1876]" captionTargetPageId="6" captionText="FIGURES 32 39. Mahellus cardoni sp. nov., male genitalia. 32. Abdominal segment VIII and genital capsule, lateral view; 33. Pygofer caudoventral process, lateral view; 34. Connective, dorsal view; 35. Style, dorsal view; 36. Subgenital plate, ventral view; 37. Aedeagus and dorsal connective, lateral view; 38, Aedeagus and dorsal connective, dorsal view; 39. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569769/files/figure.png" pageId="1" pageNumber="272">36</figureCitation>
;
<bibRefCitation id="9B434B51FFA37A0BFCA7F9C0FC05AD9D" author="Nielson" box="[888,1020,1651,1673]" pageId="1" pageNumber="272" refString="Nielson, M. W. (1982) A revision of the subfamily Coelidiinae (Homoptera: Cicadellidae) IV. Tribe Coelidiini. Pacific Insects Monograph, 38, 1 - 318." type="journal article" year="1982">Nielson 1982</bibRefCitation>
: Figs 260, 267).....................2
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<keyStep id="44262E05FFA37A0BFF48F923FA62ADEA" pageId="1" pageNumber="272">
<paragraph id="FF6D36A0FFA37A0BFF48F923FA6FADD7" blockId="1.[151,1436,1621,1908]" pageId="1" pageNumber="272">
<keyLead id="44239595FFA37A0BFF48F923FA6FADD7" pageId="1" pageNumber="272">
2. Pygofer ventral process short, glabrous (
<figureCitation id="67E92A25FFA37A0BFDAAF923FD46ADB2" box="[629,703,1680,1702]" captionStart="FIGURES 18 24" captionStartId="4.[151,264,1953,1976]" captionTargetBox="[205,1385,222,1919]" captionTargetId="figure@4.[178,1409,193,1932]" captionTargetPageId="4" captionText="FIGURES 18 24. Mahellus distanti, male genitalia. 18. Pygofer, lateral view; 19. Connective, dorsal view; 20. Style, dorsal view; 21, Subgenital plate, ventral view; 22. Aedeagus and dorsal connective, dorsal view; 23. Aedeagal shaft apex, dorsal view; 24. Aedeagus and dorsal connective, lateral view." httpUri="https://zenodo.org/record/569767/files/figure.png" pageId="1" pageNumber="272">Fig. 18</figureCitation>
); aedeagal shaft robust, tapered apically with lateral processes broad and spinose (
<figureCitation id="67E92A25FFA37A0BFEEAF91EFE5FADD7" box="[309,422,1709,1731]" captionStart="FIGURES 18 24" captionStartId="4.[151,264,1953,1976]" captionTargetBox="[205,1385,222,1919]" captionTargetId="figure@4.[178,1409,193,1932]" captionTargetPageId="4" captionText="FIGURES 18 24. Mahellus distanti, male genitalia. 18. Pygofer, lateral view; 19. Connective, dorsal view; 20. Style, dorsal view; 21, Subgenital plate, ventral view; 22. Aedeagus and dorsal connective, dorsal view; 23. Aedeagal shaft apex, dorsal view; 24. Aedeagus and dorsal connective, lateral view." httpUri="https://zenodo.org/record/569767/files/figure.png" pageId="1" pageNumber="272">Figs 2224</figureCitation>
).......................................................................
<taxonomicName id="38D24D23FFA37A0BFB26F91EFA6FADD7" ID-CoL="3XHX5" authority="Nielson" authorityName="Nielson" box="[1273,1430,1709,1731]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="distanti">
<emphasis id="CDA6EAB2FFA37A0BFB26F91EFABCADD7" box="[1273,1349,1709,1731]" italics="true" pageId="1" pageNumber="272">distanti</emphasis>
Nielson
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</paragraph>
<paragraph id="FF6D36A0FFA37A0BFF48F97FFA62ADEA" blockId="1.[151,1436,1621,1908]" pageId="1" pageNumber="272">
<keyLead id="44239595FFA37A0BFF48F97FFA62ADEA" pageId="1" pageNumber="272">
- Pygofer ventral process with dentate ventral margin (
<figureCitation id="67E92A25FFA37A0BFD01F978FCD0ADF5" box="[734,809,1739,1761]" captionStart="FIGURES 32 39" captionStartId="6.[151,264,1923,1946]" captionTargetBox="[208,1351,202,1865]" captionTargetId="figure@6.[199,1368,193,1876]" captionTargetPageId="6" captionText="FIGURES 32 39. Mahellus cardoni sp. nov., male genitalia. 32. Abdominal segment VIII and genital capsule, lateral view; 33. Pygofer caudoventral process, lateral view; 34. Connective, dorsal view; 35. Style, dorsal view; 36. Subgenital plate, ventral view; 37. Aedeagus and dorsal connective, lateral view; 38, Aedeagus and dorsal connective, dorsal view; 39. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569769/files/figure.png" pageId="1" pageNumber="272">Figs 33</figureCitation>
,
<figureCitation id="67E92A25FFA37A0BFCECF978FCB7ADF5" box="[819,846,1739,1761]" captionStart="FIGURES 40 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="1" pageNumber="272">41</figureCitation>
;
<bibRefCitation id="9B434B51FFA37A0BFC87F978FC25ADF5" author="Nielson" box="[856,988,1739,1761]" pageId="1" pageNumber="272" refString="Nielson, M. W. (1982) A revision of the subfamily Coelidiinae (Homoptera: Cicadellidae) IV. Tribe Coelidiini. Pacific Insects Monograph, 38, 1 - 318." type="journal article" year="1982">Nielson 1982</bibRefCitation>
: Fig. 263); aedeagal shaft narrow, with thinner processes (
<figureCitation id="67E92A25FFA37A0BFE96F95BFE6DADEA" box="[329,404,1768,1790]" captionStart="FIGURES 32 39" captionStartId="6.[151,264,1923,1946]" captionTargetBox="[208,1351,202,1865]" captionTargetId="figure@6.[199,1368,193,1876]" captionTargetPageId="6" captionText="FIGURES 32 39. Mahellus cardoni sp. nov., male genitalia. 32. Abdominal segment VIII and genital capsule, lateral view; 33. Pygofer caudoventral process, lateral view; 34. Connective, dorsal view; 35. Style, dorsal view; 36. Subgenital plate, ventral view; 37. Aedeagus and dorsal connective, lateral view; 38, Aedeagus and dorsal connective, dorsal view; 39. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569769/files/figure.png" pageId="1" pageNumber="272">Figs 37</figureCitation>
,
<figureCitation id="67E92A25FFA37A0BFE41F95BFE40ADEA" box="[414,441,1768,1790]" captionStart="FIGURES 40 47" captionStartId="7.[151,264,1953,1976]" captionTargetBox="[213,1386,201,1926]" captionTargetId="figure@7.[179,1408,193,1932]" captionTargetPageId="7" captionText="FIGURES 40 47. Mahellus ungulatus sp. nov., male genitalia. 40. Pygofer, lateral view; 41. Pygofer caudoventral process, lateral view; 42. Connective, dorsal view; 43. Style, dorsal view; 44. Subgenital plate, ventral view; 45. Aedeagus and dorsal connective, lateral view; 46. Aedeagus and dorsal connective, dorsal view; 47. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569770/files/figure.png" pageId="1" pageNumber="272">45</figureCitation>
;
<bibRefCitation id="9B434B51FFA37A0BFE1CF95BFDBEADEA" author="Nielson" box="[451,583,1768,1790]" pageId="1" pageNumber="272" refString="Nielson, M. W. (1982) A revision of the subfamily Coelidiinae (Homoptera: Cicadellidae) IV. Tribe Coelidiini. Pacific Insects Monograph, 38, 1 - 318." type="journal article" year="1982">Nielson 1982</bibRefCitation>
: Fig. 261)..............................................................3
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<paragraph id="FF6D36A0FFA37A0BFF48F8B6FA65AC2D" blockId="1.[151,1436,1621,1908]" pageId="1" pageNumber="272">
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3. Subgenital plate truncate apically (
<bibRefCitation id="9B434B51FFA37A0BFDF3F8B6FCF4AC0F" author="Nielson" box="[556,781,1797,1819]" pageId="1" pageNumber="272" refString="Nielson, M. W. (1982) A revision of the subfamily Coelidiinae (Homoptera: Cicadellidae) IV. Tribe Coelidiini. Pacific Insects Monograph, 38, 1 - 318." type="journal article" year="1982">Nielson 1982: Fig. 260</bibRefCitation>
); style with apophysis not produced into a thin process but narrowed (
<bibRefCitation id="9B434B51FFA37A0BFF38F890FE3EAC2D" author="Nielson" box="[231,455,1827,1849]" pageId="1" pageNumber="272" refString="Nielson, M. W. (1982) A revision of the subfamily Coelidiinae (Homoptera: Cicadellidae) IV. Tribe Coelidiini. Pacific Insects Monograph, 38, 1 - 318." type="journal article" year="1982">Nielson 1982: Fig. 259</bibRefCitation>
)...............................................................
<taxonomicName id="38D24D23FFA37A0BFB63F890FA65AC2D" authority="Distant" authorityName="Distant" box="[1212,1436,1827,1849]" class="Insecta" family="Cicadellidae" genus="Jassus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="determinatus">
<emphasis id="CDA6EAB2FFA37A0BFB63F890FAB8AC2D" box="[1212,1345,1827,1849]" italics="true" pageId="1" pageNumber="272">determinatus</emphasis>
(Distant)
</taxonomicName>
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</paragraph>
<paragraph id="FF6D36A0FFA37A0BFF48F8F2FA65AC60" blockId="1.[151,1436,1621,1908]" pageId="1" pageNumber="272">
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- Subgenital plate rounded apically (
<figureCitation id="67E92A25FFA37A0BFDF0F8F3FD8EAC42" box="[559,631,1856,1878]" captionStart="FIGURES 32 39" captionStartId="6.[151,264,1923,1946]" captionTargetBox="[208,1351,202,1865]" captionTargetId="figure@6.[199,1368,193,1876]" captionTargetPageId="6" captionText="FIGURES 32 39. Mahellus cardoni sp. nov., male genitalia. 32. Abdominal segment VIII and genital capsule, lateral view; 33. Pygofer caudoventral process, lateral view; 34. Connective, dorsal view; 35. Style, dorsal view; 36. Subgenital plate, ventral view; 37. Aedeagus and dorsal connective, lateral view; 38, Aedeagus and dorsal connective, dorsal view; 39. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569769/files/figure.png" pageId="1" pageNumber="272">Fig. 36</figureCitation>
); style with apophysis abruptly narrowed and produced into a thin process (
<figureCitation id="67E92A25FFA37A0BFA93F8F3FA6BAC42" box="[1356,1426,1856,1878]" captionStart="FIGURES 32 39" captionStartId="6.[151,264,1923,1946]" captionTargetBox="[208,1351,202,1865]" captionTargetId="figure@6.[199,1368,193,1876]" captionTargetPageId="6" captionText="FIGURES 32 39. Mahellus cardoni sp. nov., male genitalia. 32. Abdominal segment VIII and genital capsule, lateral view; 33. Pygofer caudoventral process, lateral view; 34. Connective, dorsal view; 35. Style, dorsal view; 36. Subgenital plate, ventral view; 37. Aedeagus and dorsal connective, lateral view; 38, Aedeagus and dorsal connective, dorsal view; 39. Apex of aedeagal shaft, dorsal view." httpUri="https://zenodo.org/record/569769/files/figure.png" pageId="1" pageNumber="272">Fig. 35</figureCitation>
)...................................................................................... ...
<taxonomicName id="38D24D23FFA37A0BFB20F8EEFAB0AC67" ID-CoL="8JWDS" box="[1279,1353,1885,1907]" class="Insecta" family="Cicadellidae" genus="Mahellus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="272" phylum="Arthropoda" rank="species" species="cardoni" status="sp. nov.">
<emphasis id="CDA6EAB2FFA37A0BFB20F8EEFAB0AC67" box="[1279,1353,1885,1907]" italics="true" pageId="1" pageNumber="272">cardoni</emphasis>
</taxonomicName>
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<emphasis id="CDA6EAB2FFA37A0BFA90F8EDFA65AC60" bold="true" box="[1359,1436,1886,1908]" pageId="1" pageNumber="272">sp. nov.</emphasis>
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</subSubSection>
</treatment>
</document>