1347 lines
168 KiB
XML
1347 lines
168 KiB
XML
<document id="9619FE3C9CF045D1393E08E7BAB82334" ID-DOI="10.11646/zootaxa.4040.2.1" ID-GBIF-Dataset="e672eb37-6da9-46e6-b7c2-1b281ebc02b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="235102" ID-ZooBank="BBCBBFFA-D62C-4278-BF3E-2E860C70182B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461334069115" checkinUser="plazi" docAuthor="Carleton, Michael D., Banasiak, Rebecca A. & Stanley, William T." docDate="2015" docId="776AA715FF8F62248785CF9EF8EF19F7" docLanguage="en" docName="zt04040p128.pdf" docOrigin="Zootaxa 4040 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hylomyscus heinrichorum Carleton, Banasiak & Stanley, 2015, new species" docType="treatment" docVersion="9" lastPageNumber="119" masterDocId="8B53DF6DFF8562368712CB35F80C1F6F" masterDocTitle="A new species of the rodent genus Hylomyscus from Angola, with a distributional summary of the H. anselli species group (Muridae: Murinae: Praomyini)" masterLastPageNumber="128" masterPageNumber="101" pageNumber="111" updateTime="1698618952974" updateUser="plazi">
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<mods:title id="5E1A10A1550DB56B5C7D65A24823FC12">A new species of the rodent genus Hylomyscus from Angola, with a distributional summary of the H. anselli species group (Muridae: Murinae: Praomyini)</mods:title>
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<mods:namePart id="022FF2191E64425D2A494A7A0D7A6F48">Carleton, Michael D.</mods:namePart>
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<mods:namePart id="523B4097DD6DE201F817C8C3245B8BAD">Banasiak, Rebecca A.</mods:namePart>
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<mods:roleTerm id="5BED1E84D62B09B0135602D788706BDA">Author</mods:roleTerm>
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<mods:namePart id="5750437243AC31C2F70DC94F45BD32DE">Stanley, William T.</mods:namePart>
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<mods:typeOfResource id="349E2F4C41E4F64C9384E98A38A4025D">text</mods:typeOfResource>
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<mods:title id="8C1E01CC2B54FD946BE941ECBD76EB7E">Zootaxa</mods:title>
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<mods:part id="5B650DFA32BA7E040844C0BDD6EF9BCC">
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<mods:date id="5CE3F4C98EA90E0CC32A3710088BE56D">2015</mods:date>
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<mods:number id="EE9A83F67A02E52AF150EAE37E986783">4040</mods:number>
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<treatment id="776AA715FF8F62248785CF9EF8EF19F7" ID-DOI="http://doi.org/10.5281/zenodo.6118774" ID-GBIF-Taxon="119644329" ID-Zenodo-Dep="6118774" LSID="urn:lsid:plazi:treatment:776AA715FF8F62248785CF9EF8EF19F7" httpUri="http://treatment.plazi.org/id/776AA715FF8F62248785CF9EF8EF19F7" lastPageId="18" lastPageNumber="119" pageId="10" pageNumber="111">
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<subSubSection id="B7D94588FF8F623C8785CF9EF9781B88" pageId="10" pageNumber="111" type="nomenclature">
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<paragraph id="FF7C1603FF8F623C8785CF9EFA7B1BA9" blockId="10.[151,631,1195,1255]" box="[151,631,1195,1222]" pageId="10" pageNumber="111">
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<heading id="A434A16FFF8F623C8785CF9EFA7B1BA9" bold="true" box="[151,631,1195,1222]" fontSize="11" level="1" pageId="10" pageNumber="111" reason="1">
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<emphasis id="CDB7CA11FF8F623C8785CF9EFA7B1BA9" bold="true" box="[151,631,1195,1222]" pageId="10" pageNumber="111">
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<taxonomicName id="38C36D80FF8F623C8785CF9EF9D51BAA" box="[151,473,1195,1221]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="heinrichorum" status="sp. nov.">
|
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<emphasis id="CDB7CA11FF8F623C8785CF9EF9D51BAA" bold="true" box="[151,473,1195,1221]" italics="true" pageId="10" pageNumber="111">Hylomyscus heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D684776AFF8F623C86F5CF99FA7B1BA9" box="[487,631,1196,1222]" pageId="10" pageNumber="111" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
<paragraph id="FF7C1603FF8F623C8785CFFBF9781B88" blockId="10.[151,631,1195,1255]" box="[151,372,1230,1255]" pageId="10" pageNumber="111">
|
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(
|
||
<figureCitation id="67F80A86FF8F623C878DCFFBF9071B88" box="[159,267,1230,1255]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="12.[151,250,1334,1356]" captionStartId-1="14.[151,250,1520,1542]" captionStartId-2="15.[151,250,1851,1873]" captionTargetBox-0="[171,1413,183,1311]" captionTargetBox-1="[265,1313,213,1481]" captionTargetBox-2="[359,1238,675,1815]" captionTargetId-0="figure@12.[171,1413,183,1312]" captionTargetId-1="figure@14.[256,1331,193,1499]" captionTargetId-2="figure@15.[340,1247,661,1829]" captionTargetPageId-0="12" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="FIGURE 5. Dorsal (left set) and ventral (right set) views of adult round skins (about 0.5 ×), illustrating typical pelage patterns of the three species of the Hylomyscus anselli group discussed herein: AA ′, H. arcimontensis (FMNH 151244), a male from 12.5 km NW Korogwe, Tanga Region, West Usambara Mts, 1300 m, Tanzania; BB ′, H. heinrichorum (FMNH 83796, holotype), new species, a male from “ High mountain region ” of Mt. Moco, Provincia Huambo, Angola; CC ′, H. anselli (FMNH 171353), a male from 0.5 km S and 3 km E Wipanga, Mbizi FR, Mbizi Mts, Rukwa Region, 2300 m, Tanzania. Total length of each skin, as measured in the field, = 245 mm, but lengths of the prepared skin as photographed appear different due to the circumstances of individual preparation and drying conditions." captionText-1="FIGURE 6. Dorsal (top row) and ventral (bottom row) views of adult crania (ca. 2.75 ×) representing members of the Hylomyscus anselli species group: A, H. arcimontensis (FMNH 147260; ONL = 24.9 mm), a male from Tanzania, Tanga Region, West Usambara Mts; B, H. heinrichorum, new species (FMNH 83796, holotype; ONL = 27.3 mm), a male from Angola, Provincia Huambo, Mt Moco; C, H. anselli (FMNH 171367; ONL = 28.1 mm), a male from Tanzania, Rukwa Region, Mbizi FR." captionText-2="FIGURE 7. Lateral views (ca. 4.25 ×) of adult crania of the same specimens portrayed in Fig. 6: A, H. arcimontensis; B, H. heinrichorum, new species; C, H. anselli." httpUri-0="https://zenodo.org/record/235107/files/figure.png" httpUri-1="https://zenodo.org/record/235108/files/figure.png" httpUri-2="https://zenodo.org/record/235109/files/figure.png" pageId="10" pageNumber="111">Figs. 5–7</figureCitation>
|
||
;
|
||
<tableCitation id="B24123B8FF8F623C8605CFFBF9671B88" box="[279,363,1230,1255]" captionStart="TABLE 6" captionStartId="13.[151,239,260,283]" captionTargetBox="[151,1436,372,1987]" captionTargetPageId="13" captionText="TABLE 6. Measurements of the holotype (FMNH 83796) of Hylomyscus heinrichorum, new species, and descriptive statistics for samples of the H. anselli group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations)." httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="10" pageNumber="111" tableUuid="ABBC468BFF88623B8785CA31FC0A1E34">Table 6</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7D94588FF8F623C8785CE23F9781AA9" pageId="10" pageNumber="111" type="reference_group">
|
||
<paragraph id="FF7C1603FF8F623C8785CE23FD131A43" blockId="10.[151,1436,1302,1478]" box="[151,1311,1302,1324]" pageId="10" pageNumber="111">
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||
<treatmentCitationGroup id="DFD3312DFF8F623C8785CE23FD131A43" box="[151,1311,1302,1324]" pageId="10" pageNumber="111">
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<taxonomicName id="38C36D80FF8F623C8785CE23FA511A43" ID-CoL="6MNQ9" authority="Hill & Carter, 1941: 97" authorityName="Hill & Carter" authorityPageNumber="97" authorityYear="1941" box="[151,605,1302,1324]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="carillus">
|
||
<emphasis id="CDB7CA11FF8F623C8785CE23F9681A43" box="[151,356,1302,1324]" italics="true" pageId="10" pageNumber="111">Hylomyscus carillus</emphasis>
|
||
, Hill & Carter, 1941: 97
|
||
</taxonomicName>
|
||
(part, faunal report);
|
||
<treatmentCitation id="7E623012FF8F623C842ACE23FC461A43" author="Crawford-Cabral" box="[824,1098,1302,1324]" page="79" pageId="10" pageNumber="111" year="1998">
|
||
<bibRefCitation id="9B526BF2FF8F623C842ACE23FC2B1A43" author="Crawford-Cabral" box="[824,1063,1302,1324]" pageId="10" pageNumber="111" refString="Crawford-Cabral, J. (1998) The Angolan rodents of the Superfamily Muroidea: An account of their distribution. Insituto de Investigacao Ceintifica Tropical, Lisboa (Estudos, Ensaios e Documentos, 161, 1 - 222." type="journal article" year="1998">Crawford-Cabral, 1998</bibRefCitation>
|
||
: 79
|
||
</treatmentCitation>
|
||
(part, faunal report).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF8F623C8785CE03FC171A24" blockId="10.[151,1436,1302,1478]" box="[151,1051,1333,1355]" pageId="10" pageNumber="111">
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||
<treatmentCitationGroup id="DFD3312DFF8F623C8785CE03FC171A24" box="[151,1051,1333,1355]" pageId="10" pageNumber="111">
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||
<taxonomicName id="38C36D80FF8F623C8785CE03FA7C1A24" authority="Crawford-Cabral, 1986: 163" authorityName="Crawford-Cabral" authorityPageNumber="163" authorityYear="1986" box="[151,624,1333,1355]" class="Mammalia" family="Muridae" genus="Praomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="carillus">
|
||
<emphasis id="CDB7CA11FF8F623C8785CE03F9481A24" box="[151,324,1333,1355]" italics="true" pageId="10" pageNumber="111">Praomys carillus</emphasis>
|
||
,
|
||
<treatmentCitation id="7E623012FF8F623C8643CE00FA7C1A24" author="Crawford-Cabral" box="[337,624,1333,1355]" page="163" pageId="10" pageNumber="111" year="1986">
|
||
<bibRefCitation id="9B526BF2FF8F623C8643CE00FA4C1A24" author="Crawford-Cabral" box="[337,576,1333,1355]" pageId="10" pageNumber="111" refString="Crawford-Cabral, J. (1986) A discussion of the taxa to be used in a zoogeographical analysis as illustrated in Angolan Muroidea. Cimbebasia, Serie A, 8 (19), 161 - 166." type="journal article" year="1986">Crawford-Cabral, 1986</bibRefCitation>
|
||
: 163
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
(part, name combination, zoogeography).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF8F623C8785CE61FBF01AE7" blockId="10.[151,1436,1302,1478]" pageId="10" pageNumber="111">
|
||
<treatmentCitationGroup id="DFD3312DFF8F623C8785CE61FBF01AE7" pageId="10" pageNumber="111">
|
||
<taxonomicName id="38C36D80FF8F623C8785CE61FB251A05" ID-CoL="6MNPW" authority="Musser & Carleton, 1993: 599" authorityName="Musser & Carleton" authorityPageNumber="599" authorityYear="1993" box="[151,809,1364,1386]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="denniae">
|
||
<emphasis id="CDB7CA11FF8F623C8785CE61F96B1A05" box="[151,359,1364,1386]" italics="true" pageId="10" pageNumber="111">Hylomyscus denniae</emphasis>
|
||
,
|
||
<treatmentCitation id="7E623012FF8F623C8666CE60FAA71A05" ID-DOI="http://doi.org/10.5281/zenodo.7284012" ID-Zenodo-Dep="7284012" author="Musser" box="[372,683,1364,1386]" httpUri="http://treatment.plazi.org/id/03D087AEFF94FFD9FF2F09C2FE7DFAC0" page="599" pageId="10" pageNumber="111" year="1993">
|
||
<bibRefCitation id="9B526BF2FF8F623C8666CE60FA751A05" author="Musser" box="[372,633,1364,1386]" pageId="10" pageNumber="111" refString="Musser, G. G. & Carleton, M. D. (1993) Family Muridae. In: Wilson, D. E. & Reeder, D. M. (Eds.), Mammal species of the world: A taxonomic and geographic reference, Second Edition. Smithsonian Institution Press, Washington, D. C., pp. 501 - 755." type="book chapter" year="1993">Musser & Carleton, 1993</bibRefCitation>
|
||
: 599
|
||
</treatmentCitation>
|
||
, 2005: 1336
|
||
</taxonomicName>
|
||
(part, systematic checklists, referenced as isolated population of indeterminate status).—
|
||
<treatmentCitation id="7E623012FF8F623C86C1CE47FAAA1AE7" author="Dieterlen" box="[467,678,1394,1416]" page="435" pageId="10" pageNumber="111" year="2013">
|
||
<bibRefCitation id="9B526BF2FF8F623C86C1CE47FA7A1AE7" author="Dieterlen" box="[467,630,1394,1416]" pageId="10" pageNumber="111" refString="Dieterlen, F. (2013) Hylomyscus denniae Montane Wood Mouse (Montane Hylomyscus). In: Happold, D. C. D. (Ed.), Mammals of Africa. Vol. III. Rodents, Hares and Rabbits. Bloomsbury Publishing, London, pp. 434 - 436." type="book chapter" year="2013">Dieterlen, 2013</bibRefCitation>
|
||
: 435
|
||
</treatmentCitation>
|
||
(part, faunal report, distribution).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF8F623C8785CEA7F9781AA9" blockId="10.[151,1436,1302,1478]" pageId="10" pageNumber="111">
|
||
<emphasis id="CDB7CA11FF8F623C8785CEA7F8A51AC8" box="[151,169,1426,1447]" italics="true" pageId="10" pageNumber="111">H</emphasis>
|
||
[
|
||
<emphasis id="CDB7CA11FF8F623C87A3CEA4F9021AC8" box="[177,270,1425,1447]" italics="true" pageId="10" pageNumber="111">lomyscus</emphasis>
|
||
].
|
||
<taxonomicName id="38C36D80FF8F623C8636CEA4F9861AC8" box="[292,394,1425,1447]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="anselli">
|
||
cf.
|
||
<emphasis id="CDB7CA11FF8F623C8654CEA4F9861AC8" box="[326,394,1425,1447]" italics="true" pageId="10" pageNumber="111">anselli</emphasis>
|
||
</taxonomicName>
|
||
group,
|
||
<bibRefCitation id="9B526BF2FF8F623C86C9CEA4FAEF1AC8" author="Carleton" box="[475,739,1425,1447]" pageId="10" pageNumber="111" refString="Carleton, M. D. & Stanley, W. T. (2005) Review of the Hylomyscus denniae complex in Tanzania, with description of a new species. Proceedings of the Biological Society of Washington, 118, 619 - 646." type="journal article" year="2005">Carleton & Stanley, 2005</bibRefCitation>
|
||
: 629 (taxonomic revision, definition of the
|
||
<taxonomicName id="38C36D80FF8F623C83B6CEA7FD051AC8" box="[1188,1289,1425,1447]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="111" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8F623C83B6CEA7FCBA1AC8" box="[1188,1206,1426,1447]" italics="true" pageId="10" pageNumber="111">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8F623C83D7CEA4FD051AC8" box="[1221,1289,1425,1447]" italics="true" pageId="10" pageNumber="111">anselli</emphasis>
|
||
</taxonomicName>
|
||
species group and its contents).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7D94588FF8F623D8785CEC7FCC81E9A" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111" type="materials_examined">
|
||
<paragraph id="FF7C1603FF8F623C8785CEC7FAD7198C" blockId="10.[151,1437,1522,2015]" pageId="10" pageNumber="111">
|
||
<emphasis id="CDB7CA11FF8F623C8785CEC7F9071964" bold="true" box="[151,267,1522,1547]" pageId="10" pageNumber="111">
|
||
<typeStatus id="2078A8A1FF8F623C8785CEC7F90A1964" box="[151,262,1522,1547]" pageId="10" pageNumber="111" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="99D28EC6FF8F623C8601CEC6F9681965" LSID="urn:lsid:biocol.org:col:34795" box="[275,356,1523,1546]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="111">FMNH</collectionCode>
|
||
83796, an adult male prepared as skin and skull, captured
|
||
<date id="8B7D30C3FF8F623C8312CEC6FCBC1964" box="[1024,1200,1522,1547]" pageId="10" pageNumber="111" value="1954-10-07">7 October 1954</date>
|
||
(skin tag reads “
|
||
<date id="8B7D30C3FF8F623C8263CEC6F8DE1940" pageId="10" pageNumber="111" value="1954-10-07">7 X 1954</date>
|
||
”) by Gerd H. Heinrich; the field number is recorded on the skin tag as 8778 and as GH 8778 on the now loose skull tag. Machine-printed on back of the tag is “Angolan Zoological Expedition, 1954.” Written in black pen in the collector’s script are the external measurements (“T.L.: 245, Tail: 145, H.F.: 22, Ear: 19”) and habitat (“Evergreen wood. – High mountain region”); the testes were noted as scrotal in position. See
|
||
<tableCitation id="B24123B8FF8F623C83D9CDB7FD1319F4" box="[1227,1311,1666,1691]" captionStart="TABLE 6" captionStartId="13.[151,239,260,283]" captionTargetBox="[151,1436,372,1987]" captionTargetPageId="13" captionText="TABLE 6. Measurements of the holotype (FMNH 83796) of Hylomyscus heinrichorum, new species, and descriptive statistics for samples of the H. anselli group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations)." httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="10" pageNumber="111" tableUuid="ABBC468BFF88623B8785CA31FC0A1E34">Table 6</tableCitation>
|
||
for cranial measurements of the
|
||
<typeStatus id="2078A8A1FF8F623C8696CD93F9E619D0" box="[388,490,1702,1727]" pageId="10" pageNumber="111" type="holotype">holotype</typeStatus>
|
||
. The skin is well prepared, in fine condition, and the skull is intact, in good condition with minor damage confined to the left orbital wall.
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF8F623C87D5CDDBFA071844" blockId="10.[151,1437,1522,2015]" pageId="10" pageNumber="111">
|
||
<emphasis id="CDB7CA11FF8F623C87D5CDDBF96A1868" bold="true" box="[199,358,1774,1799]" pageId="10" pageNumber="111">
|
||
<typeStatus id="2078A8A1FF8F623C87D5CDDBF90E1868" box="[199,258,1774,1799]" pageId="10" pageNumber="111">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="87D45693FF8F623C867FCDDBF9CE1868" box="[365,450,1774,1799]" name="Angola" pageId="10" pageNumber="111">Angola</collectingCountry>
|
||
, Provincia Huambo, Mount Moco, ca. 12°27.712ʹS, 15°10.600ʹE (per the
|
||
<collectingCountry id="87D45693FF8F623C8211CDDAFD381868" box="[1283,1332,1775,1799]" name="United States of America" pageId="10" pageNumber="111">U.S.</collectingCountry>
|
||
National Geospatial-Intelligence Agency).
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF8F623D87D5CC03F96D1F97" blockId="10.[151,1437,1522,2015]" lastBlockId="11.[151,1436,151,789]" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111">
|
||
The locality as recorded on the skin tag reads only “
|
||
<collectingCountry id="87D45693FF8F623C841ACC03FB531820" box="[776,863,1846,1871]" name="Angola" pageId="10" pageNumber="111">Angola</collectingCountry>
|
||
, Mount Moco.” Heinrich collected at two localities on Mt Moco, as evidenced by locality modifiers recorded in the field catalog and by an annotated, X-marked field map accessioned with that catalog (both documents maintained in the
|
||
<collectionCode id="99D28EC6FF8F623C84A6CC4AFC0918F9" LSID="urn:lsid:biocol.org:col:34795" box="[948,1029,1919,1942]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="10" pageNumber="111">FMNH</collectionCode>
|
||
Mammal Division). One X-marked site is centered near the peak of Mt Moco, and a second X is located nearby to the southeast. Specimens obtained at the latter place were labeled “foot” and collected
|
||
<date id="8B7D30C3FF8F623C85ADCCF2FB7418B0" box="[703,888,1990,2015]" pageId="10" pageNumber="111" value="1954-09-14" valueMax="1954-09-20" valueMin="1954-09-14">14–20 Sep 1954</date>
|
||
. Heinrch then relocated to a higher elevation and settled in a “high mountain region,” presumably corresponding to the former X, where he remained for over three weeks,
|
||
<date id="8B7D30C3FF8E623D87F8CB89F9DD1FBA" box="[234,465,188,213]" pageId="11" pageNumber="112" value="1954-09-22" valueMax="1954-10-16" valueMin="1954-09-22">22 Sep–16 Oct 1954</date>
|
||
. The
|
||
<typeStatus id="2078A8A1FF8E623D851CCB88FA321FBA" box="[526,574,189,213]" pageId="11" pageNumber="112">type</typeStatus>
|
||
specimen and referred specimens from Mt Moco all originated from the high mountain locality.
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF8E623D87D5CA31FBFB1EE7" blockId="11.[151,1436,151,789]" pageId="11" pageNumber="112">
|
||
<emphasis id="CDB7CA11FF8E623D87D5CA31F94A1E72" bold="true" box="[199,326,260,285]" pageId="11" pageNumber="112">
|
||
<typeStatus id="2078A8A1FF8E623D87D5CA31F94D1E72" box="[199,321,260,285]" pageId="11" pageNumber="112" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
All material here assigned to
|
||
<taxonomicName id="38C36D80FF8E623D8589CA30FB5A1E72" box="[667,854,260,285]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF8E623D8589CA30FAA31E73" box="[667,687,261,284]" italics="true" pageId="11" pageNumber="112">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8E623D85ACCA31FB5A1E72" box="[702,854,260,285]" italics="true" pageId="11" pageNumber="112">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
originates from two localities in
|
||
<collectingCountry id="87D45693FF8E623D83DDCA31FD281E72" box="[1231,1316,260,285]" name="Angola" pageId="11" pageNumber="112">Angola</collectingCountry>
|
||
, Provincia Huambo. These include 14 additional specimens from Mt Moco (
|
||
<collectionCode id="99D28EC6FF8E623D8486CA1DFBEB1E2F" LSID="urn:lsid:biocol.org:col:34795" box="[916,999,296,320]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="112">FMNH</collectionCode>
|
||
83793–83795, 83797–83799, 83801– 83807, 83895), collected from
|
||
<date id="8B7D30C3FF8E623D86E1CA79FA961E0A" box="[499,666,332,357]" pageId="11" pageNumber="112" value="1954-10-05" valueMax="1954-10-10" valueMin="1954-10-05">5–10 Oct 1954</date>
|
||
by G. Heinrich; and
|
||
<specimenCount id="E9C5DD8AFF8E623D8497CA79FC121E0A" box="[901,1054,332,357]" pageId="11" pageNumber="112" type="generic">10 specimens</specimenCount>
|
||
from Mount Soque,
|
||
<quantity id="383BBBE6FF8E623D821BCA79FD5D1E0A" box="[1289,1361,332,357]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="11" pageNumber="112" unit="km" value="42.0">42 km</quantity>
|
||
WSW Luimbale (
|
||
<collectionCode id="99D28EC6FF8E623D8602CA45F96F1EE7" LSID="urn:lsid:biocol.org:col:34795" box="[272,355,368,392]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="11" pageNumber="112">FMNH</collectionCode>
|
||
83783–83792), collected
|
||
<date id="8B7D30C3FF8E623D859ACA45FB481EE7" box="[648,836,367,392]" pageId="11" pageNumber="112" value="1954-08-24" valueMax="1954-08-28" valueMin="1954-08-24">24–28 Aug 1954</date>
|
||
by G. Heinrich.
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF8E623D87D5CAA1FCC81E9A" blockId="11.[151,1436,151,789]" pageId="11" pageNumber="112">
|
||
Heinrich’s Mount Soque (also as Serra Ussoque) is a ridge-shaped inselberg (12°17.450ʹS, 15°08. 633ʹE), located approximately
|
||
<quantity id="383BBBE6FF8E623D868DCA8DF9E61EBF" box="[415,490,439,464]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="112" unit="km" value="20.0">20 km</quantity>
|
||
NNW Mt Moco and
|
||
<quantity id="383BBBE6FF8E623D85FECA8DFB3A1EBF" box="[748,822,439,464]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="112" unit="km" value="20.0">20 km</quantity>
|
||
WSW Luimbale, its peak elevation ca.
|
||
<quantity id="383BBBE6FF8E623D821CCA82FD651EA0" box="[1294,1385,439,464]" metricMagnitude="3" metricUnit="m" metricValue="2.165" pageId="11" pageNumber="112" unit="m" value="2165.0">2165 m</quantity>
|
||
. We presume that Heinrich’s locality modifier, “
|
||
<quantity id="383BBBE6FF8E623D856ECAE9FAC91E9A" box="[636,709,476,501]" metricMagnitude="4" metricUnit="m" metricValue="4.2" pageId="11" pageNumber="112" unit="km" value="42.0">42 km</quantity>
|
||
WSW Luimbale,” was a by-road calculation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7D94588FF8E623D87D5CACAFA491D9F" pageId="11" pageNumber="112" type="diagnosis">
|
||
<paragraph id="FF7C1603FF8E623D87D5CACAFA491D9F" blockId="11.[151,1436,151,789]" pageId="11" pageNumber="112">
|
||
<emphasis id="CDB7CA11FF8E623D87D5CACAF94D1D77" bold="true" box="[199,321,511,536]" pageId="11" pageNumber="112">Diagnosis.</emphasis>
|
||
A species of the
|
||
<taxonomicName id="38C36D80FF8E623D8500CACAFAE21D77" box="[530,750,511,536]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8E623D8500CACAFAE21D77" box="[530,750,511,536]" italics="true" pageId="11" pageNumber="112">Hylomyscus anselli</emphasis>
|
||
</taxonomicName>
|
||
group characterized by larger size (ONL ≈ 26.0–
|
||
<quantity id="383BBBE6FF8E623D823CCACAFD941D78" box="[1326,1432,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="11" pageNumber="112" unit="mm" value="27.5">27.5 mm</quantity>
|
||
; CLM ≈
|
||
<quantity id="383BBBE6FF8E623D87E5C911F98E1D53" box="[247,386,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.3" metricValueMin="4.1" pageId="11" pageNumber="112" unit="mm" value="4.199999999999999" valueMax="4.3" valueMin="4.1">4.1–4.3 mm</quantity>
|
||
) compared with examples of
|
||
<taxonomicName id="38C36D80FF8E623D85CCC910FB931D53" box="[734,927,549,572]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF8E623D85CCC910FAFE1D53" box="[734,754,549,572]" italics="true" pageId="11" pageNumber="112">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8E623D8411C910FB931D53" box="[771,927,549,572]" italics="true" pageId="11" pageNumber="112">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF8E623D84CEC910FCB71D52" box="[988,1211,548,573]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="kerbispeterhansi">
|
||
<emphasis id="CDB7CA11FF8E623D84CEC910FBFC1D53" box="[988,1008,549,572]" italics="true" pageId="11" pageNumber="112">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8E623D8313C911FCB71D52" box="[1025,1211,548,573]" italics="true" pageId="11" pageNumber="112">kerbispeterhansi</emphasis>
|
||
</taxonomicName>
|
||
(ONL ≈
|
||
<quantity id="383BBBE6FF8E623D823EC911F8CA1D30" metricMagnitude="-2" metricUnit="m" metricValue="2.55" metricValueMax="2.65" metricValueMin="2.45" pageId="11" pageNumber="112" unit="mm" value="25.5" valueMax="26.5" valueMin="24.5">24.5–26.5 mm</quantity>
|
||
; CLM ≈ 3.5–4.0 mm), as reflected in most craniodental dimensions recorded (
|
||
<tableCitation id="B24123B8FF8E623D8361C972FCC31D0F" box="[1139,1231,583,608]" captionStart="TABLE 6" captionStartId="13.[151,239,260,283]" captionTargetBox="[151,1436,372,1987]" captionTargetPageId="13" captionText="TABLE 6. Measurements of the holotype (FMNH 83796) of Hylomyscus heinrichorum, new species, and descriptive statistics for samples of the H. anselli group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations)." httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="11" pageNumber="112" tableUuid="ABBC468BFF88623B8785CA31FC0A1E34">Table 6</tableCitation>
|
||
). Compared with examples of
|
||
<taxonomicName id="38C36D80FF8E623D8634C958F9981DEA" box="[294,404,620,645]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8E623D8634C958F9361DEB" box="[294,314,621,644]" italics="true" pageId="11" pageNumber="112">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8E623D865BC959F9981DEA" box="[329,404,620,645]" italics="true" pageId="11" pageNumber="112">anselli</emphasis>
|
||
</taxonomicName>
|
||
, size comparable, but bony palate notably shorter with absolutely and proportionally longer incisive foramina (LIF ≈ 75–80% of LD) that penetrate between the anterior roots of the first molars; molars slightly more robust and zygomatic plate broader; head-and-body and tail average shorter in length, and rostral dimensions (LR, LD) average smaller.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7D94588FF8E623D87D5C9C9FD201C7B" box="[199,1324,764,789]" pageId="11" pageNumber="112" type="distribution">
|
||
<paragraph id="FF7C1603FF8E623D87D5C9C9FD201C7B" blockId="11.[151,1436,151,789]" box="[199,1324,764,789]" pageId="11" pageNumber="112">
|
||
<emphasis id="CDB7CA11FF8E623D87D5C9C9F96C1C7A" bold="true" box="[199,352,764,789]" pageId="11" pageNumber="112">Distribution.</emphasis>
|
||
Restricted to high mountainous region of westcentral
|
||
<collectingCountry id="87D45693FF8E623D84D2C9C9FC1F1C7A" box="[960,1043,764,789]" name="Angola" pageId="11" pageNumber="112">Angola</collectingCountry>
|
||
as so far known (
|
||
<figureCitation id="67F80A86FF8E623D83CBC9C9FD131C7B" box="[1241,1311,764,789]" captionStart="FIGURE 4" captionStartId="11.[151,250,1866,1888]" captionTargetBox="[155,1431,846,1839]" captionTargetId="figure@11.[151,1436,841,1845]" captionTargetPageId="11" captionText="FIGURE 4. Distributions of species representing the Hylomyscus anselli and H. denniae groups and their relation to principal mountainous systems of central Africa. Localities of the H. anselli group are based on specimens documented herein and by Ansell (1957, 1978), Carleton & Stanley (2005), and Carleton et al. (2006); see Demos et al. (2014) for additional localities of H. kerbispeterhansi. Localities of the H. denniae group are based on specimens previously reported by Carleton et al. (2006)." httpUri="https://zenodo.org/record/235106/files/figure.png" pageId="11" pageNumber="112">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="ABBC468BFF8E623D8785CC7FFD8118D3" httpUri="https://zenodo.org/record/235106/files/figure.png" pageId="11" pageNumber="112" targetBox="[155,1431,846,1839]" targetPageId="11">
|
||
<paragraph id="FF7C1603FF8E623D8785CC7FFD8118D3" blockId="11.[151,1436,1866,1980]" pageId="11" pageNumber="112">
|
||
<emphasis id="CDB7CA11FF8E623D8785CC7FF901180F" bold="true" box="[151,269,1866,1888]" pageId="11" pageNumber="112">FIGURE 4</emphasis>
|
||
. Distributions of species representing the
|
||
<taxonomicName id="38C36D80FF8E623D85A8CC7FFB73180F" box="[698,895,1866,1888]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8E623D85A8CC7FFB73180F" box="[698,895,1866,1888]" italics="true" pageId="11" pageNumber="112">Hylomyscus anselli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF8E623D84A3CC7EFC2C180F" box="[945,1056,1866,1888]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="denniae">
|
||
<emphasis id="CDB7CA11FF8E623D84A3CC7EFBCF180F" box="[945,963,1867,1888]" italics="true" pageId="11" pageNumber="112">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8E623D84C2CC7FFC2C180F" box="[976,1056,1866,1888]" italics="true" pageId="11" pageNumber="112">denniae</emphasis>
|
||
</taxonomicName>
|
||
groups and their relation to principal mountainous systems of central Africa. Localities of the
|
||
<taxonomicName id="38C36D80FF8E623D85F0CC5FFB4A1810" box="[738,838,1897,1919]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8E623D85F0CC5FFAF81810" box="[738,756,1898,1919]" italics="true" pageId="11" pageNumber="112">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8E623D8410CC5CFB4A1810" box="[770,838,1897,1919]" italics="true" pageId="11" pageNumber="112">anselli</emphasis>
|
||
</taxonomicName>
|
||
group are based on specimens documented herein and by Ansell (1957, 1978), Carleton & Stanley (2005), and Carleton
|
||
<emphasis id="CDB7CA11FF8E623D8400CCBCFB4A18F2" box="[786,838,1927,1950]" italics="true" pageId="11" pageNumber="112">et al.</emphasis>
|
||
(2006); see Demos
|
||
<emphasis id="CDB7CA11FF8E623D8300CCBCFC4918F2" box="[1042,1093,1927,1950]" italics="true" pageId="11" pageNumber="112">et al.</emphasis>
|
||
(2014) for additional localities of
|
||
<taxonomicName id="38C36D80FF8E623D8785CC92F95218D3" box="[151,350,1958,1980]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="kerbispeterhansi">
|
||
<emphasis id="CDB7CA11FF8E623D8785CC92F8A518D3" box="[151,169,1959,1980]" italics="true" pageId="11" pageNumber="112">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8E623D87A4CC93F95218D3" box="[182,350,1958,1980]" italics="true" pageId="11" pageNumber="112">kerbispeterhansi</emphasis>
|
||
</taxonomicName>
|
||
. Localities of the
|
||
<taxonomicName id="38C36D80FF8E623D8507CC92FA8818D3" box="[533,644,1958,1980]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="112" phylum="Chordata" rank="species" species="denniae">
|
||
<emphasis id="CDB7CA11FF8E623D8507CC92FA2B18D3" box="[533,551,1959,1980]" italics="true" pageId="11" pageNumber="112">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8E623D8526CC93FA8818D3" box="[564,644,1958,1980]" italics="true" pageId="11" pageNumber="112">denniae</emphasis>
|
||
</taxonomicName>
|
||
group are based on specimens previously reported by Carleton
|
||
<emphasis id="CDB7CA11FF8E623D8218CC92FD3118D3" box="[1290,1341,1958,1980]" italics="true" pageId="11" pageNumber="112">et al.</emphasis>
|
||
(2006).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="ABBC468BFF89623A8785CE03FB1F196B" httpUri="https://zenodo.org/record/235107/files/figure.png" pageId="12" pageNumber="113" targetBox="[171,1413,183,1311]" targetPageId="12">
|
||
<paragraph id="FF7C1603FF89623A8785CE03FB1F196B" blockId="12.[151,1436,1334,1540]" pageId="12" pageNumber="113">
|
||
<emphasis id="CDB7CA11FF89623A8785CE03F9011A23" bold="true" box="[151,269,1334,1356]" pageId="12" pageNumber="113">FIGURE 5</emphasis>
|
||
. Dorsal (left set) and ventral (right set) views of adult round skins (about 0.5 ×), illustrating typical pelage patterns of the three species of the
|
||
<taxonomicName id="38C36D80FF89623A86B9CE60FA7C1A04" box="[427,624,1365,1387]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF89623A86B9CE60FA7C1A04" box="[427,624,1365,1387]" italics="true" pageId="12" pageNumber="113">Hylomyscus anselli</emphasis>
|
||
</taxonomicName>
|
||
group discussed herein: AA ′,
|
||
<taxonomicName id="38C36D80FF89623A84BECE63FC551A04" box="[940,1113,1366,1387]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF89623A84BECE63FC551A04" box="[940,1113,1366,1387]" italics="true" pageId="12" pageNumber="113">H. arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 151244), a male from 12.5 km NW Korogwe, Tanga Region, West Usambara Mts, 1300 m, Tanzania; BB ′,
|
||
<taxonomicName id="38C36D80FF89623A832CCE40FCE11AE5" box="[1086,1261,1396,1418]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF89623A832CCE40FCE11AE5" box="[1086,1261,1396,1418]" italics="true" pageId="12" pageNumber="113">H. heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 83796, holotype), new species, a male from “High mountain region” of Mt. Moco, Provincia Huambo, Angola; CC ′,
|
||
<taxonomicName id="38C36D80FF89623A83F1CEA6FD491AC7" box="[1251,1349,1426,1448]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF89623A83F1CEA6FD491AC7" box="[1251,1349,1426,1448]" italics="true" pageId="12" pageNumber="113">H. anselli</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 171353), a male from 0.5 km S and 3 km E Wipanga, Mbizi FR, Mbizi Mts, Rukwa Region, 2300 m, Tanzania. Total length of each skin, as measured in the field, = 245 mm, but lengths of the prepared skin as photographed appear different due to the circumstances of individual preparation and drying conditions.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="B7D94588FF89622487D5CD19FA681E2F" lastPageId="18" lastPageNumber="119" pageId="12" pageNumber="113" type="description">
|
||
<paragraph id="FF7C1603FF89623A87D5CD19FB60182F" blockId="12.[151,1436,1580,2037]" pageId="12" pageNumber="113">
|
||
<emphasis id="CDB7CA11FF89623A87D5CD19FA38192A" bold="true" box="[199,564,1580,1605]" pageId="12" pageNumber="113">Description and comparisons.</emphasis>
|
||
<taxonomicName id="38C36D80FF89623A852DCD19FB66192A" box="[575,874,1580,1605]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF89623A852DCD19FB66192A" box="[575,874,1580,1605]" italics="true" pageId="12" pageNumber="113">Hylomyscus heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
can be associated with the other species so far recognized in the
|
||
<taxonomicName id="38C36D80FF89623A8670CD64F9C31907" box="[354,463,1615,1640]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF89623A8670CD64F97A1907" box="[354,374,1617,1640]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A8697CD7AF9C31907" box="[389,463,1615,1640]" italics="true" pageId="12" pageNumber="113">anselli</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<taxonomicName id="38C36D80FF89623A8539CD64FA951907" box="[555,665,1615,1640]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF89623A8539CD64FA331907" box="[555,575,1617,1640]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A855CCD7AFA951907" box="[590,665,1615,1640]" italics="true" pageId="12" pageNumber="113">anselli</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF89623A85BACD64FB6B1907" box="[680,871,1616,1640]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF89623A85BACD64FAB01907" box="[680,700,1617,1640]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A85D9CD64FB6B1907" box="[715,871,1617,1640]" italics="true" pageId="12" pageNumber="113">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF89623A8464CD64FC581907" box="[886,1108,1615,1640]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="kerbispeterhansi">
|
||
<emphasis id="CDB7CA11FF89623A8464CD64FB861907" box="[886,906,1617,1640]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A8488CD7AFC581907" box="[922,1108,1615,1640]" italics="true" pageId="12" pageNumber="113">kerbispeterhansi</emphasis>
|
||
</taxonomicName>
|
||
) based on certain qualitative traits. Foremost among these are the lack of pectoral mammae; the large expanse of the subsquamosal fenestrum, which together with the postglenoid foramen defines a slender and long hamular process (see
|
||
<bibRefCitation id="9B526BF2FF89623A83ACCDA2F8D819BA" author="Carleton" pageId="12" pageNumber="113" refString="Carleton, M. D. & Stanley, W. T. (2005) Review of the Hylomyscus denniae complex in Tanzania, with description of a new species. Proceedings of the Biological Society of Washington, 118, 619 - 646." type="journal article" year="2005">Carleton & Stanley, 2005</bibRefCitation>
|
||
:
|
||
<figureCitation id="67F80A86FF89623A87CDCD89F92819BA" box="[223,292,1724,1749]" captionStart="FIGURE 6" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[265,1313,213,1481]" captionTargetId="figure@14.[256,1331,193,1499]" captionTargetPageId="14" captionText="FIGURE 6. Dorsal (top row) and ventral (bottom row) views of adult crania (ca. 2.75 ×) representing members of the Hylomyscus anselli species group: A, H. arcimontensis (FMNH 147260; ONL = 24.9 mm), a male from Tanzania, Tanga Region, West Usambara Mts; B, H. heinrichorum, new species (FMNH 83796, holotype; ONL = 27.3 mm), a male from Angola, Provincia Huambo, Mt Moco; C, H. anselli (FMNH 171367; ONL = 28.1 mm), a male from Tanzania, Rukwa Region, Mbizi FR." httpUri="https://zenodo.org/record/235108/files/figure.png" pageId="12" pageNumber="113">Fig. 6</figureCitation>
|
||
); and its relatively shorter incisive foramina (namely, shorter in comparison with members of the
|
||
<taxonomicName id="38C36D80FF89623A8293CD88F8FC1997" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="denniae">
|
||
<emphasis id="CDB7CA11FF89623A8293CD88FD9919BB" box="[1409,1429,1725,1748]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A8785CDEAF8FC1997" box="[151,240,1759,1784]" italics="true" pageId="12" pageNumber="113">denniae</emphasis>
|
||
</taxonomicName>
|
||
group). The number of mammary glands in
|
||
<taxonomicName id="38C36D80FF89623A85FCCDD4FBA71997" box="[750,939,1759,1784]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF89623A85FCCDD4FB0E1997" box="[750,770,1761,1784]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A8400CDEAFBA71997" box="[786,939,1759,1784]" italics="true" pageId="12" pageNumber="113">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
—six, distributed as one post-axial pair and two inguinal pairs—was verified on several prepared round skins (FMNH 83793, 83795, 83802–83804), all females that were apparently in late-term pregnancy or lactating.
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF89623B87D5CC79FB791FBA" blockId="12.[151,1436,1580,2037]" lastBlockId="13.[151,1436,151,213]" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113">
|
||
Although we excluded external variables in our multivariate analyses, mean differences were highly significant for TOTL, TL, and HBL in one-way ANOVAs that compared our tabulated samples of
|
||
<taxonomicName id="38C36D80FF89623A83F0CC44FD9018E7" box="[1250,1436,1903,1928]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF89623A83F0CC44FCFA18E7" box="[1250,1270,1905,1928]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A8216CC5AFD9018E7" box="[1284,1436,1903,1928]" italics="true" pageId="12" pageNumber="113">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF89623A87DACCA0F93B18C2" box="[200,311,1940,1965]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF89623A87DACCA0F8D018C3" box="[200,220,1941,1964]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A87FECCA1F93B18C2" box="[236,311,1940,1965]" italics="true" pageId="12" pageNumber="113">anselli</emphasis>
|
||
</taxonomicName>
|
||
(F values = 13.8–14.4, P <0.001, df 1, 35), but not for HFL (F = 0.02, P = 0.884, df 1, 35). The sample of
|
||
<taxonomicName id="38C36D80FF89623A8605CC8CF9DB18BF" box="[279,471,1975,2000]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF89623A8605CC8CF92718BF" box="[279,299,1977,2000]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A862DCC82F9DB18BF" box="[319,471,1975,2000]" italics="true" pageId="12" pageNumber="113">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
averaged smaller than that of
|
||
<taxonomicName id="38C36D80FF89623A845FCC8CFBCC18BF" box="[845,960,1975,2000]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="113" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF89623A845FCC8CFB6D18BF" box="[845,865,1977,2000]" italics="true" pageId="12" pageNumber="113">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF89623A8467CC82FBCC18BF" box="[885,960,1975,2000]" italics="true" pageId="12" pageNumber="113">anselli</emphasis>
|
||
</taxonomicName>
|
||
for all three significantly different skin variables (
|
||
<tableCitation id="B24123B8FF89623A861DCCE9F960189A" box="[271,364,2012,2037]" captionStart="TABLE 6" captionStartId="13.[151,239,260,283]" captionTargetBox="[151,1436,372,1987]" captionTargetPageId="13" captionText="TABLE 6. Measurements of the holotype (FMNH 83796) of Hylomyscus heinrichorum, new species, and descriptive statistics for samples of the H. anselli group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations)." httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="12" pageNumber="113" tableUuid="ABBC468BFF88623B8785CA31FC0A1E34">Table 6</tableCitation>
|
||
). Our samples of the two species were obtained by different collectors, and such external variables are notoriously variable depending upon investigator habit and field experience. Even so, the disparity in means seems suitably large to convey real taxonomic differences.
|
||
</paragraph>
|
||
<caption id="ABBC468BFF88623B8785CA31FC0A1E34" ID-Table-UUID="ABBC468BFF88623B8785CA31FC0A1E34" httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="13" pageNumber="114" targetBox="[151,1436,372,1987]" targetIsTable="true" targetPageId="13">
|
||
<paragraph id="FF7C1603FF88623B8785CA31FC0A1E34" blockId="13.[151,1436,259,347]" pageId="13" pageNumber="114">
|
||
<emphasis id="CDB7CA11FF88623B8785CA31F9011E74" bold="true" box="[151,269,260,283]" pageId="13" pageNumber="114">TABLE 6.</emphasis>
|
||
Measurements of the holotype (FMNH 83796) of
|
||
<taxonomicName id="38C36D80FF88623B8429CA36FC5A1E74" box="[827,1110,259,283]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="heinrichorum" status="sp. nov.">
|
||
<emphasis id="CDB7CA11FF88623B8429CA36FC5A1E74" box="[827,1110,259,283]" italics="true" pageId="13" pageNumber="114">Hylomyscus heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D684776AFF88623B8377CA31FCE61E74" box="[1125,1258,260,283]" pageId="13" pageNumber="114" rank="species">new species</taxonomicNameLabel>
|
||
, and descriptive statistics for samples of the
|
||
<taxonomicName id="38C36D80FF88623B86ADCA11FA2A1E54" box="[447,550,291,315]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF88623B86ADCA11F9DE1E54" box="[447,466,292,315]" italics="true" pageId="13" pageNumber="114">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF88623B86CDCA16FA2A1E54" box="[479,550,291,315]" italics="true" pageId="13" pageNumber="114">anselli</emphasis>
|
||
</taxonomicName>
|
||
group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF7C1603FF88623B8661CA43FD5118AC" pageId="13" pageNumber="114">
|
||
<table id="8DC3E4A3FF889DC98785CA41FD9018AC" box="[151,1436,372,1987]" gridcols="4" gridrows="24" pageId="13" pageNumber="114">
|
||
<tr id="41F31441FF889DC98785CA41FD901EE2" box="[151,1436,372,397]" gridrow="0" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CA41FA711EE2" box="[151,637,372,397]" gridcol="0" gridrow="0" pageId="13" pageNumber="114">
|
||
<taxonomicName id="38C36D80FF88623B8661CA43FA281EE3" box="[371,548,372,397]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF88623B8661CA43F98A1EE2" box="[371,390,374,397]" italics="true" pageId="13" pageNumber="114">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF88623B8681CA41FA281EE3" box="[403,548,372,396]" italics="true" pageId="13" pageNumber="114">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="02227D3DFF889DC985A7CA41FB471EE2" box="[693,843,372,397]" gridcol="1" gridrow="0" pageId="13" pageNumber="114">
|
||
<taxonomicName id="38C36D80FF88623B85DECA43FB381EE3" box="[716,820,372,397]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF88623B85DECA43FAD31EE2" box="[716,735,374,397]" italics="true" pageId="13" pageNumber="114">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF88623B85FFCA41FB381EE3" box="[749,820,372,396]" italics="true" pageId="13" pageNumber="114">anselli</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="02227D3DFF889DC984B9CA41FC6C1EE2" box="[939,1120,372,397]" gridcol="2" gridrow="0" pageId="13" pageNumber="114">
|
||
<taxonomicName id="38C36D80FF88623B84B9CA43FC6C1EE2" box="[939,1120,373,397]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF88623B84B9CA43FBB21EE2" box="[939,958,374,397]" italics="true" pageId="13" pageNumber="114">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF88623B84DECA43FC6C1EE2" box="[972,1120,374,397]" italics="true" pageId="13" pageNumber="114">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="02227D3DFF889DC983A1CA41FD901EE2" box="[1203,1436,372,397]" gridcol="3" gridrow="0" pageId="13" pageNumber="114">
|
||
<taxonomicName id="38C36D80FF88623B83A1CA43FD881EE3" box="[1203,1412,372,397]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="114" phylum="Chordata" rank="species" species="kerbispeterhansi">
|
||
<emphasis id="CDB7CA11FF88623B83A1CA43FCCA1EE2" box="[1203,1222,374,397]" italics="true" pageId="13" pageNumber="114">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF88623B83C1CA41FD881EE3" box="[1235,1412,372,396]" italics="true" pageId="13" pageNumber="114">kerbispeterhansi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CAABFD901EDA" box="[151,1436,414,437]" gridrow="1" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CAABFA711EDA" box="[151,637,414,437]" gridcol="0" gridrow="1" pageId="13" pageNumber="114">Variable Holotype Angolan Highlands</th>
|
||
<td id="02227D3DFF889DC985A7CAABFB471EDA" box="[693,843,414,437]" gridcol="1" gridrow="1" pageId="13" pageNumber="114">Mbizi Mts</td>
|
||
<td id="02227D3DFF889DC984B9CAABFC6C1EDA" box="[939,1120,414,437]" gridcol="2" gridrow="1" pageId="13" pageNumber="114">E Usambara Mts</td>
|
||
<td id="02227D3DFF889DC983A1CAABFD901EDA" box="[1203,1436,414,437]" gridcol="3" gridrow="1" pageId="13" pageNumber="114">Kenyan Mts</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CAFDFD901E93" box="[151,1436,456,508]" gridrow="2" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CAFDFA711E93" box="[151,637,456,508]" gridcol="0" gridrow="2" pageId="13" pageNumber="114">TOTL 245.0 231.4 ± 12.5 207–255 (25)</th>
|
||
<td id="02227D3DFF889DC985A7CAFDFB471E93" box="[693,843,456,508]" gridcol="1" gridrow="2" pageId="13" pageNumber="114">246.1 ± 7.4 238–264 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CAFDFC6C1E93" box="[939,1120,456,508]" gridcol="2" gridrow="2" pageId="13" pageNumber="114">224.4 ± 17.1 186–247 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CAFDFD901E93" box="[1203,1436,456,508]" gridcol="3" gridrow="2" pageId="13" pageNumber="114">219.3 ± 13.3 195–231 (7)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785C93BFD901D2C" box="[151,1436,526,579]" gridrow="3" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785C93BFA711D2C" box="[151,637,526,579]" gridcol="0" gridrow="3" pageId="13" pageNumber="114">HBL 100.0 94.0 ± 6.6 82–106 (25)</th>
|
||
<td id="02227D3DFF889DC985A7C93BFB471D2C" box="[693,843,526,579]" gridcol="1" gridrow="3" pageId="13" pageNumber="114">101.6 ± 3.8 95–109 (12)</td>
|
||
<td id="02227D3DFF889DC984B9C93BFC6C1D2C" box="[939,1120,526,579]" gridcol="2" gridrow="3" pageId="13" pageNumber="114">88.1 ± 8.3 72–110 (24)</td>
|
||
<td id="02227D3DFF889DC983A1C93BFD901D2C" box="[1203,1436,526,579]" gridcol="3" gridrow="3" pageId="13" pageNumber="114">90.3 ± 7.8 76–100 (7)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785C961FD901DE6" box="[151,1436,596,649]" gridrow="4" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785C961FA711DE6" box="[151,637,596,649]" gridcol="0" gridrow="4" pageId="13" pageNumber="114">TL 145.0 137.3 ± 7.4 121–150 (25)</th>
|
||
<td id="02227D3DFF889DC985A7C961FB471DE6" box="[693,843,596,649]" gridcol="1" gridrow="4" pageId="13" pageNumber="114">146.3 ± 5.0 141–159 (12)</td>
|
||
<td id="02227D3DFF889DC984B9C961FC6C1DE6" box="[939,1120,596,649]" gridcol="2" gridrow="4" pageId="13" pageNumber="114">136.5 ± 10.1 116–150 (24)</td>
|
||
<td id="02227D3DFF889DC983A1C961FD901DE6" box="[1203,1436,596,649]" gridcol="3" gridrow="4" pageId="13" pageNumber="114">129.0 ± 6.8 119–138 (7)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785C9AEFD901DBF" box="[151,1436,667,720]" gridrow="5" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785C9AEFA711DBF" box="[151,637,667,720]" gridcol="0" gridrow="5" pageId="13" pageNumber="114">HFL 22.0 21.1 ± 1.0 18.5–22 (25)</th>
|
||
<td id="02227D3DFF889DC985A7C9AEFB471DBF" box="[693,843,667,720]" gridcol="1" gridrow="5" pageId="13" pageNumber="114">21.2 ± 0.7 20–22 (12)</td>
|
||
<td id="02227D3DFF889DC984B9C9AEFC6C1DBF" box="[939,1120,667,720]" gridcol="2" gridrow="5" pageId="13" pageNumber="114">19.5 ± 1.3 17–22 (24)</td>
|
||
<td id="02227D3DFF889DC983A1C9AEFD901DBF" box="[1203,1436,667,720]" gridcol="3" gridrow="5" pageId="13" pageNumber="114">19.1 ± 1.2 17–21 (8)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785C9D4FD901C79" box="[151,1436,737,790]" gridrow="6" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785C9D4FA711C79" box="[151,637,737,790]" gridcol="0" gridrow="6" pageId="13" pageNumber="114">EAR 19.0 18.5 ± 0.9 17–20 (24)</th>
|
||
<td id="02227D3DFF889DC985A7C9D4FB471C79" box="[693,843,737,790]" gridcol="1" gridrow="6" pageId="13" pageNumber="114">20.1 ± 0.5 19–21 (12)</td>
|
||
<td id="02227D3DFF889DC984B9C9D4FC6C1C79" box="[939,1120,737,790]" gridcol="2" gridrow="6" pageId="13" pageNumber="114">17.8 ± 1.0 15–19 (24)</td>
|
||
<td id="02227D3DFF889DC983A1C9D4FD901C79" box="[1203,1436,737,790]" gridcol="3" gridrow="6" pageId="13" pageNumber="114">18.6 ± 0.8 17–19.5 (8)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785C81DFD901C33" box="[151,1436,808,860]" gridrow="7" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785C81DFA711C33" box="[151,637,808,860]" gridcol="0" gridrow="7" pageId="13" pageNumber="114">WT -- --</th>
|
||
<td id="02227D3DFF889DC985A7C81DFB471C33" box="[693,843,808,860]" gridcol="1" gridrow="7" pageId="13" pageNumber="114">28.5 ± 3.1 22.0–34.5 (12)</td>
|
||
<td id="02227D3DFF889DC984B9C81DFC6C1C33" box="[939,1120,808,860]" gridcol="2" gridrow="7" pageId="13" pageNumber="114">20.2 ± 4.4 12.0–29.0 (24)</td>
|
||
<td id="02227D3DFF889DC983A1C81DFD901C33" box="[1203,1436,808,860]" gridcol="3" gridrow="7" pageId="13" pageNumber="114">21.5 ± 3.3 18.0–26.0 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785C85BFD901CCC" box="[151,1436,878,931]" gridrow="8" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785C85BFA711CCC" box="[151,637,878,931]" gridcol="0" gridrow="8" pageId="13" pageNumber="114">ONL 27.4 26.7 ± 0.8 25.0–28.1 (14)</th>
|
||
<td id="02227D3DFF889DC985A7C85BFB471CCC" box="[693,843,878,931]" gridcol="1" gridrow="8" pageId="13" pageNumber="114">27.4 ± 0.7 26.3–28.6 (12)</td>
|
||
<td id="02227D3DFF889DC984B9C85BFC6C1CCC" box="[939,1120,878,931]" gridcol="2" gridrow="8" pageId="13" pageNumber="114">25.2 ± 1.0 23.0–27.0 (24)</td>
|
||
<td id="02227D3DFF889DC983A1C85BFD901CCC" box="[1203,1436,878,931]" gridcol="3" gridrow="8" pageId="13" pageNumber="114">25.5 ± 0.8 24.6–26.9 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785C881FD901C86" box="[151,1436,948,1001]" gridrow="9" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785C881FA711C86" box="[151,637,948,1001]" gridcol="0" gridrow="9" pageId="13" pageNumber="114">ZB 13.5 13.0 ± 0.4 12.4–13.8 (14)</th>
|
||
<td id="02227D3DFF889DC985A7C881FB471C86" box="[693,843,948,1001]" gridcol="1" gridrow="9" pageId="13" pageNumber="114">13.5 ± 0.2 12.9–13.7 (12)</td>
|
||
<td id="02227D3DFF889DC984B9C881FC6C1C86" box="[939,1120,948,1001]" gridcol="2" gridrow="9" pageId="13" pageNumber="114">12.5 ± 0.5 11.6–13.7 (24)</td>
|
||
<td id="02227D3DFF889DC983A1C881FD901C86" box="[1203,1436,948,1001]" gridcol="3" gridrow="9" pageId="13" pageNumber="114">12.7 ± 0.6 12.0–13.5 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785C8CEFD901B5F" box="[151,1436,1019,1072]" gridrow="10" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785C8CEFA711B5F" box="[151,637,1019,1072]" gridcol="0" gridrow="10" pageId="13" pageNumber="114">BBC 11.8 11.7 ± 0.2 11.3–12.1 (14)</th>
|
||
<td id="02227D3DFF889DC985A7C8CEFB471B5F" box="[693,843,1019,1072]" gridcol="1" gridrow="10" pageId="13" pageNumber="114">11.9 ± 0.2 11.5–12.3 (12)</td>
|
||
<td id="02227D3DFF889DC984B9C8CEFC6C1B5F" box="[939,1120,1019,1072]" gridcol="2" gridrow="10" pageId="13" pageNumber="114">11.2 ± 0.2 10.7–11.8 (24)</td>
|
||
<td id="02227D3DFF889DC983A1C8CEFD901B5F" box="[1203,1436,1019,1072]" gridcol="3" gridrow="10" pageId="13" pageNumber="114">11.3 ± 0.2 10.8–11.5 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CF74FD901B19" box="[151,1436,1089,1142]" gridrow="11" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CF74FA711B19" box="[151,637,1089,1142]" gridcol="0" gridrow="11" pageId="13" pageNumber="114">IOB 4.4 4.5 ± 0.1 4.4–4.7 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CF74FB471B19" box="[693,843,1089,1142]" gridcol="1" gridrow="11" pageId="13" pageNumber="114">4.6 ± 0.1 4.5–4.8 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CF74FC6C1B19" box="[939,1120,1089,1142]" gridcol="2" gridrow="11" pageId="13" pageNumber="114">4.3 ± 0.1 4.0–4.7 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CF74FD901B19" box="[1203,1436,1089,1142]" gridcol="3" gridrow="11" pageId="13" pageNumber="114">4.3 ± 0.1 4.2–4.6 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CFBDFD901BD3" box="[151,1436,1160,1212]" gridrow="12" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CFBDFA711BD3" box="[151,637,1160,1212]" gridcol="0" gridrow="12" pageId="13" pageNumber="114">BOC 6.5 6.2 ± 0.2 6.0–6.5 (13)</th>
|
||
<td id="02227D3DFF889DC985A7CFBDFB471BD3" box="[693,843,1160,1212]" gridcol="1" gridrow="12" pageId="13" pageNumber="114">6.2 ± 0.1 5.9–6.5 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CFBDFC6C1BD3" box="[939,1120,1160,1212]" gridcol="2" gridrow="12" pageId="13" pageNumber="114">6.0 ± 0.2 5.5–6.5 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CFBDFD901BD3" box="[1203,1436,1160,1212]" gridcol="3" gridrow="12" pageId="13" pageNumber="114">6.0 ± 0.1 5.8–6.2 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CFFBFD901A6C" box="[151,1436,1230,1283]" gridrow="13" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CFFBFA711A6C" box="[151,637,1230,1283]" gridcol="0" gridrow="13" pageId="13" pageNumber="114">LR 8.2 8.2 ± 0.5 7.2–8.9 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CFFBFB471A6C" box="[693,843,1230,1283]" gridcol="1" gridrow="13" pageId="13" pageNumber="114">8.7 ± 0.3 8.3–9.1 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CFFBFC6C1A6C" box="[939,1120,1230,1283]" gridcol="2" gridrow="13" pageId="13" pageNumber="114">7.7 ± 0.4 7.1–8.4 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CFFBFD901A6C" box="[1203,1436,1230,1283]" gridcol="3" gridrow="13" pageId="13" pageNumber="114">7.9 ± 0.2 7.6–8.1 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CE21FD901A26" box="[151,1436,1300,1353]" gridrow="14" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CE21FA711A26" box="[151,637,1300,1353]" gridcol="0" gridrow="14" pageId="13" pageNumber="114">BR 4.7 4.4 ± 0.2 4.1–4.8 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CE21FB471A26" box="[693,843,1300,1353]" gridcol="1" gridrow="14" pageId="13" pageNumber="114">4.7 ± 0.2 4.5–5.0 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CE21FC6C1A26" box="[939,1120,1300,1353]" gridcol="2" gridrow="14" pageId="13" pageNumber="114">4.3 ± 0.2 3.9–4.7 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CE21FD901A26" box="[1203,1436,1300,1353]" gridcol="3" gridrow="14" pageId="13" pageNumber="114">4.5 ± 0.2 4.3–4.8 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CE6EFD901AFF" box="[151,1436,1371,1424]" gridrow="15" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CE6EFA711AFF" box="[151,637,1371,1424]" gridcol="0" gridrow="15" pageId="13" pageNumber="114">PPL 9.4 9.3 ± 0.5 8.6–10.1 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CE6EFB471AFF" box="[693,843,1371,1424]" gridcol="1" gridrow="15" pageId="13" pageNumber="114">9.5 ± 0.4 8.9–10.1 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CE6EFC6C1AFF" box="[939,1120,1371,1424]" gridcol="2" gridrow="15" pageId="13" pageNumber="114">8.7 ± 0.4 7.6–9.4 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CE6EFD901AFF" box="[1203,1436,1371,1424]" gridcol="3" gridrow="15" pageId="13" pageNumber="114">8.7 ± 0.6 8.0–9.6 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CE94FD901AB9" box="[151,1436,1441,1494]" gridrow="16" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CE94FA711AB9" box="[151,637,1441,1494]" gridcol="0" gridrow="16" pageId="13" pageNumber="114">HPL 4.7 4.4 ± 0.2 4.0–4.7 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CE94FB471AB9" box="[693,843,1441,1494]" gridcol="1" gridrow="16" pageId="13" pageNumber="114">4.8 ± 0.2 4.5–5.0 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CE94FC6C1AB9" box="[939,1120,1441,1494]" gridcol="2" gridrow="16" pageId="13" pageNumber="114">4.3 ± 0.2 4.0–4.7 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CE94FD901AB9" box="[1203,1436,1441,1494]" gridcol="3" gridrow="16" pageId="13" pageNumber="114">4.4 ± 0.2 4.2–4.8 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CEDDFD901973" box="[151,1436,1512,1564]" gridrow="17" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CEDDFA711973" box="[151,637,1512,1564]" gridcol="0" gridrow="17" pageId="13" pageNumber="114">LIF 5.9 5.7 ± 0.3 5.2–6.4 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CEDDFB471973" box="[693,843,1512,1564]" gridcol="1" gridrow="17" pageId="13" pageNumber="114">5.5 ± 0.1 5.3–5.8 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CEDDFC6C1973" box="[939,1120,1512,1564]" gridcol="2" gridrow="17" pageId="13" pageNumber="114">5.2 ± 0.2 4.6–5.6 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CEDDFD901973" box="[1203,1436,1512,1564]" gridcol="3" gridrow="17" pageId="13" pageNumber="114">5.3 ± 0.1 5.1–5.5 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CD1BFD90190C" box="[151,1436,1582,1635]" gridrow="18" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CD1BFA71190C" box="[151,637,1582,1635]" gridcol="0" gridrow="18" pageId="13" pageNumber="114">LD 7.4 7.4 ± 0.4 6.7–8.1 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CD1BFB47190C" box="[693,843,1582,1635]" gridcol="1" gridrow="18" pageId="13" pageNumber="114">7.7 ± 0.3 7.4–8.4 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CD1BFC6C190C" box="[939,1120,1582,1635]" gridcol="2" gridrow="18" pageId="13" pageNumber="114">7.0 ± 0.3 6.4–7.7 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CD1BFD90190C" box="[1203,1436,1582,1635]" gridcol="3" gridrow="18" pageId="13" pageNumber="114">7.5 ± 0.3 7.1–7.9 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CD41FD9019C6" box="[151,1436,1652,1705]" gridrow="19" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CD41FA7119C6" box="[151,637,1652,1705]" gridcol="0" gridrow="19" pageId="13" pageNumber="114">BBP 5.5 5.3 ± 0.1 5.1–5.5 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CD41FB4719C6" box="[693,843,1652,1705]" gridcol="1" gridrow="19" pageId="13" pageNumber="114">5.2 ± 0.1 5.1–5.4 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CD41FC6C19C6" box="[939,1120,1652,1705]" gridcol="2" gridrow="19" pageId="13" pageNumber="114">4.9 ± 0.1 4.6–5.1 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CD41FD9019C6" box="[1203,1436,1652,1705]" gridcol="3" gridrow="19" pageId="13" pageNumber="114">5.0 ± 0.1 4.8–5.2 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CD8EFD90199F" box="[151,1436,1723,1776]" gridrow="20" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CD8EFA71199F" box="[151,637,1723,1776]" gridcol="0" gridrow="20" pageId="13" pageNumber="114">BZP 2.8 2.5 ± 0.1 2.2–2.8 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CD8EFB47199F" box="[693,843,1723,1776]" gridcol="1" gridrow="20" pageId="13" pageNumber="114">2.4 ± 0.1 2.2–2.6 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CD8EFC6C199F" box="[939,1120,1723,1776]" gridcol="2" gridrow="20" pageId="13" pageNumber="114">2.3 ± 0.1 2.1–2.6 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CD8EFD90199F" box="[1203,1436,1723,1776]" gridcol="3" gridrow="20" pageId="13" pageNumber="114">2.3 ± 0.1 2.1–2.5 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CC34FD901859" box="[151,1436,1793,1846]" gridrow="21" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CC34FA711859" box="[151,637,1793,1846]" gridcol="0" gridrow="21" pageId="13" pageNumber="114">LAB 4.4 4.4 ± 0.1 4.3–4.6 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CC34FB471859" box="[693,843,1793,1846]" gridcol="1" gridrow="21" pageId="13" pageNumber="114">4.5 ± 0.1 4.5–4.7 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CC34FC6C1859" box="[939,1120,1793,1846]" gridcol="2" gridrow="21" pageId="13" pageNumber="114">4.1 ± 0.1 3.9–4.3 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CC34FD901859" box="[1203,1436,1793,1846]" gridcol="3" gridrow="21" pageId="13" pageNumber="114">4.4 ± 0.1 4.2–4.6 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CC7DFD901813" box="[151,1436,1864,1916]" gridrow="22" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CC7DFA711813" box="[151,637,1864,1916]" gridcol="0" gridrow="22" pageId="13" pageNumber="114">CLM 4.30 4.21 ± 0.12 4.02–4.39 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CC7DFB471813" box="[693,843,1864,1916]" gridcol="1" gridrow="22" pageId="13" pageNumber="114">4.16 ± 0.11 3.87–4.28 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CC7DFC6C1813" box="[939,1120,1864,1916]" gridcol="2" gridrow="22" pageId="13" pageNumber="114">3.70 ± 0.12 3.37–3.89 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CC7DFD901813" box="[1203,1436,1864,1916]" gridcol="3" gridrow="22" pageId="13" pageNumber="114">3.80 ± 0.04 3.76–3.89 (6)</td>
|
||
</tr>
|
||
<tr id="41F31441FF889DC98785CCBBFD9018AC" box="[151,1436,1934,1987]" gridrow="23" pageId="13" pageNumber="114">
|
||
<th id="02227D3DFF889DC98785CCBBFA7118AC" box="[151,637,1934,1987]" gridcol="0" gridrow="23" pageId="13" pageNumber="114">WM1 1.32 1.29 ± 0.04 1.20–1.34 (14)</th>
|
||
<td id="02227D3DFF889DC985A7CCBBFB4718AC" box="[693,843,1934,1987]" gridcol="1" gridrow="23" pageId="13" pageNumber="114">1.24 ± 0.04 1.17–1.29 (12)</td>
|
||
<td id="02227D3DFF889DC984B9CCBBFC6C18AC" box="[939,1120,1934,1987]" gridcol="2" gridrow="23" pageId="13" pageNumber="114">1.13 ± 0.03 1.06–1.18 (24)</td>
|
||
<td id="02227D3DFF889DC983A1CCBBFD9018AC" box="[1203,1436,1934,1987]" gridcol="3" gridrow="23" pageId="13" pageNumber="114">1.17 ± 0.01 1.16–1.19 (6)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="ABBC468BFF8B62388785CEC5F90C19EE" httpUri="https://zenodo.org/record/235108/files/figure.png" pageId="14" pageNumber="115" targetBox="[265,1313,213,1481]" targetPageId="14">
|
||
<paragraph id="FF7C1603FF8B62388785CEC5F90C19EE" blockId="14.[151,1436,1520,1665]" pageId="14" pageNumber="115">
|
||
<emphasis id="CDB7CA11FF8B62388785CEC5F91B1969" bold="true" box="[151,279,1520,1542]" pageId="14" pageNumber="115">FIGURE 6.</emphasis>
|
||
Dorsal (top row) and ventral (bottom row) views of adult crania (ca. 2.75 ×) representing members of the
|
||
<taxonomicName id="38C36D80FF8B62388785CD3AF953194A" box="[151,351,1551,1573]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8B62388785CD3AF953194A" box="[151,351,1551,1573]" italics="true" pageId="14" pageNumber="115">Hylomyscus anselli</emphasis>
|
||
</taxonomicName>
|
||
species group: A,
|
||
<taxonomicName id="38C36D80FF8B62388538CD25FAD5194A" box="[554,729,1552,1573]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF8B62388538CD25FA30194A" box="[554,572,1552,1573]" italics="true" pageId="14" pageNumber="115">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8B6238855ECD25FAD5194A" box="[588,729,1552,1573]" italics="true" pageId="14" pageNumber="115">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 147260; ONL = 24.9 mm), a male from Tanzania, Tanga Region, West Usambara Mts; B,
|
||
<taxonomicName id="38C36D80FF8B623886E4CD1BFAAE192C" box="[502,674,1581,1603]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="heinrichorum" status="sp. nov.">
|
||
<emphasis id="CDB7CA11FF8B623886E4CD1BFA04192C" box="[502,520,1582,1603]" italics="true" pageId="14" pageNumber="115">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8B6238850ACD18FAAE192C" box="[536,674,1581,1603]" italics="true" pageId="14" pageNumber="115">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D684776AFF8B623885A1CD1BFB3C192C" box="[691,816,1581,1603]" pageId="14" pageNumber="115" rank="species">new species</taxonomicNameLabel>
|
||
(FMNH 83796, holotype; ONL = 27.3 mm), a male from Angola, Provincia Huambo, Mt Moco; C,
|
||
<taxonomicName id="38C36D80FF8B62388553CD78FAA8190D" box="[577,676,1612,1634]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8B62388553CD78FA5F190D" box="[577,595,1613,1634]" italics="true" pageId="14" pageNumber="115">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8B62388572CD79FAA8190D" box="[608,676,1612,1634]" italics="true" pageId="14" pageNumber="115">anselli</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 171367; ONL = 28.1 mm), a male from Tanzania, Rukwa Region, Mbizi FR.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF7C1603FF8B623987D5CD9DFAC71D30" blockId="14.[151,1437,1704,2017]" lastBlockId="15.[151,1437,151,608]" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115">
|
||
The pelage of
|
||
<taxonomicName id="38C36D80FF8B6238867ECD9CFA2419AE" box="[364,552,1704,1729]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF8B6238867ECD9CF98C19AF" box="[364,384,1705,1728]" italics="true" pageId="14" pageNumber="115">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8B62388682CD9DFA2419AE" box="[400,552,1704,1729]" italics="true" pageId="14" pageNumber="115">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
resembles that observed in other members of the
|
||
<taxonomicName id="38C36D80FF8B62388376CD9CFCDE19AE" box="[1124,1234,1704,1729]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8B62388376CD9CFC7419AF" box="[1124,1144,1705,1728]" italics="true" pageId="14" pageNumber="115">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8B62388395CD9DFCDE19AE" box="[1159,1234,1704,1729]" italics="true" pageId="14" pageNumber="115">anselli</emphasis>
|
||
</taxonomicName>
|
||
group, being soft and fine in texture, short and closely adpressed to the body. The most notable pelage contrast of the Angolan form involves dorsal pelage color. Individual dorsal hairs can be characterized as medium plumbeous gray tipped with bright buff; the overall effect, however, is not so dark and somber a brown as in examples of
|
||
<taxonomicName id="38C36D80FF8B623883C4CC20FD4A1842" box="[1238,1350,1812,1837]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8B623883C4CC20FCE61843" box="[1238,1258,1813,1836]" italics="true" pageId="14" pageNumber="115">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8B623883E9CC21FD4A1842" box="[1275,1350,1812,1837]" italics="true" pageId="14" pageNumber="115">anselli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF8B62388293CC20F93F183F" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF8B62388293CC20FD991843" box="[1409,1429,1813,1836]" italics="true" pageId="14" pageNumber="115">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8B62388785CC0CF93F183F" box="[151,307,1849,1872]" italics="true" pageId="14" pageNumber="115">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="67F80A86FF8B62388657CC0DF980183E" box="[325,396,1848,1873]" captionStart="FIGURE 5" captionStartId="12.[151,250,1334,1356]" captionTargetBox="[171,1413,183,1311]" captionTargetId="figure@12.[171,1413,183,1312]" captionTargetPageId="12" captionText="FIGURE 5. Dorsal (left set) and ventral (right set) views of adult round skins (about 0.5 ×), illustrating typical pelage patterns of the three species of the Hylomyscus anselli group discussed herein: AA ′, H. arcimontensis (FMNH 151244), a male from 12.5 km NW Korogwe, Tanga Region, West Usambara Mts, 1300 m, Tanzania; BB ′, H. heinrichorum (FMNH 83796, holotype), new species, a male from “ High mountain region ” of Mt. Moco, Provincia Huambo, Angola; CC ′, H. anselli (FMNH 171353), a male from 0.5 km S and 3 km E Wipanga, Mbizi FR, Mbizi Mts, Rukwa Region, 2300 m, Tanzania. Total length of each skin, as measured in the field, = 245 mm, but lengths of the prepared skin as photographed appear different due to the circumstances of individual preparation and drying conditions." httpUri="https://zenodo.org/record/235107/files/figure.png" pageId="14" pageNumber="115">Fig. 5</figureCitation>
|
||
), but rather an even toned, dull yellowish brown (buckthorn brown), light to moderate in saturation. Guard hairs are brown with hyaline tips, only a little longer than the dorsal fur except over the rump. There is little tendency toward darker over the middle dorsum or conspicuously brighter flanks as is common in the other two species; color is generally evenly graded in expression over the entire dorsum, from the middle back to the flanks. A few individuals of
|
||
<taxonomicName id="38C36D80FF8B62388517CCFCFACC188E" box="[517,704,1992,2017]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="14" pageNumber="115" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF8B62388517CCFCFA15188F" box="[517,537,1993,2016]" italics="true" pageId="14" pageNumber="115">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8B6238853ACCFDFACC188E" box="[552,704,1992,2017]" italics="true" pageId="14" pageNumber="115">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
exhibit brighter ochraceous-tipped hairs along the upper leg and shoulder and side of the head; nevertheless, the dorsal pelage color is predominantly even toned in appearance. Dorsal-ventral pelage contrast is well marked but not accentuated by a brighter lateral line. In all three species, the ventral hairs are basally gray and tipped with white, imparting a grayish-white effect. The three can be crudely sorted as bright gray (
|
||
<taxonomicName id="38C36D80FF8A62398686CA30FA581E73" box="[404,596,261,284]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF8A62398686CA30F9A41E73" box="[404,424,261,284]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A623986AACA30FA581E73" box="[440,596,261,284]" italics="true" pageId="15" pageNumber="116">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
), medium gray (
|
||
<taxonomicName id="38C36D80FF8A62398404CA30FBDE1E72" box="[790,978,260,285]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF8A62398404CA30FB261E73" box="[790,810,261,284]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A62398428CA31FBDE1E72" box="[826,978,260,285]" italics="true" pageId="15" pageNumber="116">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
), and dark gray (
|
||
<taxonomicName id="38C36D80FF8A62398389CA30FD051E72" box="[1179,1289,260,285]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8A62398389CA30FCA31E73" box="[1179,1199,261,284]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A623983ADCA31FD051E72" box="[1215,1289,260,285]" italics="true" pageId="15" pageNumber="116">anselli</emphasis>
|
||
</taxonomicName>
|
||
), with much variation and overlap among them. The tail of
|
||
<taxonomicName id="38C36D80FF8A623985A1CA1CFB7C1E2F" box="[691,880,295,320]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF8A623985A1CA1CFACB1E2F" box="[691,711,297,320]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A623985CACA12FB7C1E2F" box="[728,880,295,320]" italics="true" pageId="15" pageNumber="116">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
, as in other members of the
|
||
<taxonomicName id="38C36D80FF8A623983ADCA1CFD231E2F" box="[1215,1327,295,320]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8A623983ADCA1CFCDF1E2F" box="[1215,1235,297,320]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A623983F6CA12FD231E2F" box="[1252,1327,295,320]" italics="true" pageId="15" pageNumber="116">anselli</emphasis>
|
||
</taxonomicName>
|
||
group, is notably longer than the head and body (TL ≈ 142–146% of HBL), its color dusky-brown and wholly dark around the circumference; caudal scales are finely textured and hairs short, imparting a naked appearance over most of its length, the fine caudal hairs becoming macroscopically visible toward the tip. The hind foot is short and narrow, as per the genus, with digit 5 nearly as long as digits 2–4; the naked plantar surface bears six well-formed, cushiony pads. Pale brown hairs of the limb usually continue onto the ankle and proximal metatarsum, forming a dusky medial metatarsal streak in some individuals, replaced by white hairs over the distal metatarsum and phalanges; glistening white ungual tufts are present. Pinnae are pale brown to dark tan in the Angolan species, compared with darker, more blackish brown ears in the other two.
|
||
</paragraph>
|
||
<caption id="ABBC468BFF8A62398785CC0EFA361800" httpUri="https://zenodo.org/record/235109/files/figure.png" pageId="15" pageNumber="116" targetBox="[359,1238,675,1815]" targetPageId="15">
|
||
<paragraph id="FF7C1603FF8A62398785CC0EFA361800" blockId="15.[151,1437,1851,1903]" pageId="15" pageNumber="116">
|
||
<emphasis id="CDB7CA11FF8A62398785CC0EF918183E" bold="true" box="[151,276,1851,1873]" pageId="15" pageNumber="116">FIGURE 7.</emphasis>
|
||
Lateral views (ca. 4.25 ×) of adult crania of the same specimens portrayed in Fig. 6: A,
|
||
<taxonomicName id="38C36D80FF8A623983BACC09FD5A183E" box="[1192,1366,1852,1873]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF8A623983BACC09FCB6183E" box="[1192,1210,1852,1873]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A623983DBCC09FD5A183E" box="[1225,1366,1852,1873]" italics="true" pageId="15" pageNumber="116">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
; B,
|
||
<taxonomicName id="38C36D80FF8A62398291CC09F92D1800" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="heinrichorum" status="sp. nov.">
|
||
<emphasis id="CDB7CA11FF8A62398291CC09F92D1800" italics="true" pageId="15" pageNumber="116">H. heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D684776AFF8A6239863FCC6FF9A61800" box="[301,426,1881,1903]" pageId="15" pageNumber="116" rank="species">new species</taxonomicNameLabel>
|
||
; C,
|
||
<taxonomicName id="38C36D80FF8A623986C0CC6FFA391800" box="[466,565,1881,1903]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8A623986C0CC6FF9E81800" box="[466,484,1882,1903]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A623986E3CC6CFA391800" box="[497,565,1881,1903]" italics="true" pageId="15" pageNumber="116">anselli</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF7C1603FF8A622687D5CCA8FB521B13" blockId="15.[151,1436,1948,2009]" lastBlockId="16.[151,1437,151,1977]" lastPageId="16" lastPageNumber="117" pageId="15" pageNumber="116">
|
||
<collectingCountry id="87D45693FF8A623987D5CCA8F8EA18DA" box="[199,230,1949,1973]" name="American Samoa" pageId="15" pageNumber="116">As</collectingCountry>
|
||
a gestalt observation, the skull of
|
||
<taxonomicName id="38C36D80FF8A62398573CCA8FB1718DA" box="[609,795,1948,1973]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF8A62398573CCA8FA7918DB" box="[609,629,1949,1972]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A62398591CCA9FB1718DA" box="[643,795,1948,1973]" italics="true" pageId="15" pageNumber="116">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
is approximately equal to or only slightly smaller than
|
||
<taxonomicName id="38C36D80FF8A62398293CCA8F8EE18B6" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF8A62398293CCA8FD9918DB" box="[1409,1429,1949,1972]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A62398785CCF5F8EE18B6" box="[151,226,1984,2009]" italics="true" pageId="15" pageNumber="116">anselli</emphasis>
|
||
</taxonomicName>
|
||
, albeit heavier in build, but both are substantially larger than
|
||
<taxonomicName id="38C36D80FF8A623984B3CCF4FC6C18B7" box="[929,1120,1985,2008]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="15" pageNumber="116" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF8A623984B3CCF4FBB918B7" box="[929,949,1985,2008]" italics="true" pageId="15" pageNumber="116">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF8A623984D6CCF4FC6C18B7" box="[964,1120,1985,2008]" italics="true" pageId="15" pageNumber="116">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="67F80A86FF8A62398363CCF5FCCE18B6" box="[1137,1218,1984,2009]" captionStart="FIGURE 6" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[265,1313,213,1481]" captionTargetId="figure@14.[256,1331,193,1499]" captionTargetPageId="14" captionText="FIGURE 6. Dorsal (top row) and ventral (bottom row) views of adult crania (ca. 2.75 ×) representing members of the Hylomyscus anselli species group: A, H. arcimontensis (FMNH 147260; ONL = 24.9 mm), a male from Tanzania, Tanga Region, West Usambara Mts; B, H. heinrichorum, new species (FMNH 83796, holotype; ONL = 27.3 mm), a male from Angola, Provincia Huambo, Mt Moco; C, H. anselli (FMNH 171367; ONL = 28.1 mm), a male from Tanzania, Rukwa Region, Mbizi FR." httpUri="https://zenodo.org/record/235108/files/figure.png" pageId="15" pageNumber="116">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="67F80A86FF8A623983DDCCF4FCD318B6" box="[1231,1247,1985,2009]" captionStart="FIGURE 7" captionStartId="15.[151,250,1851,1873]" captionTargetBox="[359,1238,675,1815]" captionTargetId="figure@15.[340,1247,661,1829]" captionTargetPageId="15" captionText="FIGURE 7. Lateral views (ca. 4.25 ×) of adult crania of the same specimens portrayed in Fig. 6: A, H. arcimontensis; B, H. heinrichorum, new species; C, H. anselli." httpUri="https://zenodo.org/record/235109/files/figure.png" pageId="15" pageNumber="116">7</figureCitation>
|
||
). Thus, standard external measurements and those we recorded for the skull and molar rows easily serve to distinguish examples of
|
||
<taxonomicName id="38C36D80FF9562268785CB88F9591FBA" box="[151,341,188,213]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF9562268785CB88F8A71FBB" box="[151,171,189,212]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622687AFCB89F9591FBA" box="[189,341,188,213]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
from those of the smaller
|
||
<taxonomicName id="38C36D80FF9562268584CB88FB551FBB" box="[662,857,189,212]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF9562268584CB88FAA61FBB" box="[662,682,189,212]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622685AFCB88FB551FBB" box="[701,857,189,212]" italics="true" pageId="16" pageNumber="117">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF956226848ACB88FC741FBA" box="[920,1144,188,213]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="kerbispeterhansi">
|
||
<emphasis id="CDB7CA11FF956226848ACB88FBA01FBB" box="[920,940,189,212]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622684ACCB89FC741FBA" box="[958,1144,188,213]" italics="true" pageId="16" pageNumber="117">kerbispeterhansi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="B24123B8FF9562268399CB89FCE91FBA" box="[1163,1253,188,213]" captionStart="TABLE 6" captionStartId="13.[151,239,260,283]" captionTargetBox="[151,1436,372,1987]" captionTargetPageId="13" captionText="TABLE 6. Measurements of the holotype (FMNH 83796) of Hylomyscus heinrichorum, new species, and descriptive statistics for samples of the H. anselli group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations)." httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="16" pageNumber="117" tableUuid="ABBC468BFF88623B8785CA31FC0A1E34">Table 6</tableCitation>
|
||
); size routinely emerged as the preeminent factor accounting for their separation in multivariate space (
|
||
<figureCitation id="67F80A86FF95622683B1CBEAFCF91F97" box="[1187,1269,223,248]" captionStart="FIGURE 1" captionStartId="5.[151,250,1756,1778]" captionTargetBox="[273,1302,202,1731]" captionTargetId="figure@5.[267,1303,193,1738]" captionTargetPageId="5" captionText="FIGURE 1. Results of discriminant function analysis performed on 16 log-transformed craniodental variables, as measured on 175 intact adult specimens representing 10 OTUs of the Hylomyscus anselli species group. A) Projection of OTU centroids onto first two canonical variates (CV) extracted (see Table 1 for variable loadings). Polygons enclose maximal dispersion of individual specimen scores around each group centroid; half-filled circles indicate the scores of the three Zambian specimens from the type series of anselli Bishop (1979), which were entered as unknowns. B) Cluster diagram (UPGMA) based on average Mahalanobis distances between the 10 OTU centroids as derived from discriminant function analysis (coefficient of cophenetic correlation = 0.934). Bootstrapping values are indicated for all nodes (1000 iterations); mountain system and OTU identifier are indicated for each terminal OTU, along with species recognized herein at those stems that subtend major geographic associations." httpUri="https://zenodo.org/record/235103/files/figure.png" pageId="16" pageNumber="117">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="67F80A86FF956226821ACBD5FD141F97" box="[1288,1304,224,248]" captionStart="FIGURE 3" captionStartId="7.[151,250,1756,1778]" captionTargetBox="[316,1246,200,1721]" captionTargetId="figure@7.[311,1247,193,1724]" captionTargetPageId="7" captionText="FIGURE 3. Morphometric comparisons of the Angolan sample with Hylomyscus kerbispeterhansi based on analyses of 16 logtransformed craniodental variables; selected representatives of related species were included to provide multivariate context (see Material and Methods for variable abbreviations and Table 5 for variable loadings). A) Scatterplot of OTU centroids for the first two canonical variates (CV) extracted from six-group discriminant function analysis (H. anselli, N = 12; H. arcimontensis, N = 55 — OTUs 3, 6, and 8; H. kerbispeterhansi, N = 6; Angolan sample, N = 13). Irregular polygons circumscribe maximal dispersion of specimen scores around an OTU centroid. B) Scatterplot of individual scores for the first two principal components (PC) extracted from ordination of specimens representing H. arcimontensis (OTU 3, N = 23), H. kerbispeterhansi (N = 6), and the Angolan sample (N = 13). Major axes of the species constellations and 1 SD confidence ellipses around the sample centroid are indicated (see text for discussion)." httpUri="https://zenodo.org/record/235105/files/figure.png" pageId="16" pageNumber="117">3</figureCitation>
|
||
). Mensural differences between
|
||
<taxonomicName id="38C36D80FF9562268690CA30FA321E72" box="[386,574,260,285]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF9562268690CA30F99A1E73" box="[386,406,261,284]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622686B4CA31FA321E72" box="[422,574,260,285]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF9562268568CA30FAE51E72" box="[634,745,260,285]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF9562268568CA30FA821E73" box="[634,654,261,284]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF956226858CCA31FAE51E72" box="[670,745,260,285]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
, on the other hand, are not readily grasped when eyeballing series of skulls, yet specimens of each did conform to distinct morphometric footprints based on the cranial measurements we obtained (see
|
||
<figureCitation id="67F80A86FF956226850CCA79FA7D1E0B" box="[542,625,332,357]" captionStart="FIGURE 1" captionStartId="5.[151,250,1756,1778]" captionTargetBox="[273,1302,202,1731]" captionTargetId="figure@5.[267,1303,193,1738]" captionTargetPageId="5" captionText="FIGURE 1. Results of discriminant function analysis performed on 16 log-transformed craniodental variables, as measured on 175 intact adult specimens representing 10 OTUs of the Hylomyscus anselli species group. A) Projection of OTU centroids onto first two canonical variates (CV) extracted (see Table 1 for variable loadings). Polygons enclose maximal dispersion of individual specimen scores around each group centroid; half-filled circles indicate the scores of the three Zambian specimens from the type series of anselli Bishop (1979), which were entered as unknowns. B) Cluster diagram (UPGMA) based on average Mahalanobis distances between the 10 OTU centroids as derived from discriminant function analysis (coefficient of cophenetic correlation = 0.934). Bootstrapping values are indicated for all nodes (1000 iterations); mountain system and OTU identifier are indicated for each terminal OTU, along with species recognized herein at those stems that subtend major geographic associations." httpUri="https://zenodo.org/record/235103/files/figure.png" pageId="16" pageNumber="117">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="67F80A86FF9562268594CA79FA991E0B" box="[646,661,332,356]" captionStart="FIGURE 2" captionStartId="6.[151,250,1708,1730]" captionTargetBox="[157,1434,202,1685]" captionTargetId="figure@6.[153,1435,193,1686]" captionTargetPageId="6" captionText="FIGURE 2. Three scatterplots depicting results of principal component (PC) analyses of 16 log-transformed craniodental variables as measured on 175 intact crania representing the Hylomyscus anselli species group. A) Projection of individual scores onto components I and II extracted from ordination of specimens representing H. anselli (N = 15), H. arcimontensis (N = 147), and the Angolan sample (N = 13). B) Projection of specimen scores onto components I and III extracted from the same ordination as performed in plot A. C) Projection of individual scores onto the first two components extracted from ordination of specimens representing H. anselli (N = 15) and the Angolan sample (N = 13); major axes of the species constellations and regression statistics are indicated; the Y intercepts differ significantly (F = 8.23, P = 0.009), but not their slopes (F = 0.38, P = 0.546) (see text for discussion). For scatterplots A and B, see Table 2 for variable correlations and variance explained; for scatterplot C, see Table 4; variable abbreviations are given in the Materials and Methods. The half-filled circles in all three plots indicate the scores of the three Zambian specimens from the type series of anselli Bishop (1979)." httpUri="https://zenodo.org/record/235104/files/figure.png" pageId="16" pageNumber="117">2</figureCitation>
|
||
C). However, fewer variables contributed to their discrimination, rendering their cranial separation correspondingly more difficult, based on subtle distinctions. Configuration of the hard palate relative to the length of the incisive foramina offers particularly important contrasts, those variables (BBP, HPL, LIF) loading strongly on that principal component which captured their separation in multivariate space (
|
||
<figureCitation id="67F80A86FF95622687FBCAE9F93D1E9B" box="[233,305,476,501]" captionStart="FIGURE 2" captionStartId="6.[151,250,1708,1730]" captionTargetBox="[157,1434,202,1685]" captionTargetId="figure@6.[153,1435,193,1686]" captionTargetPageId="6" captionText="FIGURE 2. Three scatterplots depicting results of principal component (PC) analyses of 16 log-transformed craniodental variables as measured on 175 intact crania representing the Hylomyscus anselli species group. A) Projection of individual scores onto components I and II extracted from ordination of specimens representing H. anselli (N = 15), H. arcimontensis (N = 147), and the Angolan sample (N = 13). B) Projection of specimen scores onto components I and III extracted from the same ordination as performed in plot A. C) Projection of individual scores onto the first two components extracted from ordination of specimens representing H. anselli (N = 15) and the Angolan sample (N = 13); major axes of the species constellations and regression statistics are indicated; the Y intercepts differ significantly (F = 8.23, P = 0.009), but not their slopes (F = 0.38, P = 0.546) (see text for discussion). For scatterplots A and B, see Table 2 for variable correlations and variance explained; for scatterplot C, see Table 4; variable abbreviations are given in the Materials and Methods. The half-filled circles in all three plots indicate the scores of the three Zambian specimens from the type series of anselli Bishop (1979)." httpUri="https://zenodo.org/record/235104/files/figure.png" pageId="16" pageNumber="117">Fig. 2</figureCitation>
|
||
C,
|
||
<tableCitation id="B24123B8FF9562268644CAE9F9BC1E9B" box="[342,432,476,501]" captionStart="TABLE 4" captionStartId="8.[151,239,978,1001]" captionTargetBox="[159,1321,1125,1976]" captionTargetPageId="8" captionText="TABLE 4. Results of principal component analysis performed on 16 log-transformed measurements of Hylomyscus anselli (N = 12) and the Angolan sample (N = 13), complemented with one-way ANOVAs of input variables with OTU as categorical effect (df = 1 / 26). See Fig. 2 C for a scatterplot of principal component scores and Materials and Methods for variable abbreviations." httpUri="http://table.plazi.org/id/ABBC468BFF8D623E8785C8E7F9BC1B26" pageId="16" pageNumber="117" tableUuid="ABBC468BFF8D623E8785C8E7F9BC1B26">Table 4</tableCitation>
|
||
).
|
||
<collectingCountry id="87D45693FF95622686DBCAE9F9E41E9B" box="[457,488,476,500]" name="American Samoa" pageId="16" pageNumber="117">As</collectingCountry>
|
||
practical anatomical landmarks when viewing skulls, these loadings translate as absolutely shorter incisive foramina (LIF ca. 69–73% of LD) whose posterior end falls just short of or about level with the anterior border of the M1 anterior root in
|
||
<taxonomicName id="38C36D80FF95622685C6C910FB4F1D52" box="[724,835,548,573]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF95622685C6C910FAE41D53" box="[724,744,549,572]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622685EAC911FB4F1D52" box="[760,835,548,573]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
, versus longer incisive foramina (LIF ca. 75–80% of LD) that extend beyond the anterior roots about to the rim of the first lamina of M
|
||
<quantity id="383BBBE6FF9562268323C97DFC5B1D0F" box="[1073,1111,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="16" pageNumber="117" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="38C36D80FF956226834CC97CFD141D0F" box="[1118,1304,583,608]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF956226834CC97CFC7E1D0F" box="[1118,1138,585,608]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF9562268392C972FD141D0F" box="[1152,1304,583,608]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, the incisive foramina in
|
||
<taxonomicName id="38C36D80FF95622686B9C958FA6A1DEA" box="[427,614,620,645]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF95622686B9C958F9B31DEB" box="[427,447,621,644]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622686DCC959FA6A1DEA" box="[462,614,620,645]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
are proportionately the longest within the
|
||
<taxonomicName id="38C36D80FF9562268359C958FCB51DEA" box="[1099,1209,620,645]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF9562268359C958FC531DEB" box="[1099,1119,621,644]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF956226837CC959FCB51DEA" box="[1134,1209,620,645]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
group, albeit not so “long” as members of the
|
||
<taxonomicName id="38C36D80FF95622686ABC9A4FA3F1DC7" box="[441,563,655,680]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="denniae">
|
||
<emphasis id="CDB7CA11FF95622686ABC9A4F9C11DC7" box="[441,461,657,680]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622686C8C9BAFA3F1DC7" box="[474,563,655,680]" italics="true" pageId="16" pageNumber="117">denniae</emphasis>
|
||
</taxonomicName>
|
||
group. The absolutely longer bony palate documented in
|
||
<taxonomicName id="38C36D80FF95622683A6C9A4FD2C1DC7" box="[1204,1312,655,680]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF95622683A6C9A4FCC41DC7" box="[1204,1224,657,680]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622683C7C9BAFD2C1DC7" box="[1237,1312,655,680]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
, compared with a shorter palate in
|
||
<taxonomicName id="38C36D80FF95622686A3C980FA621DA2" box="[433,622,692,717]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF95622686A3C980F9C91DA3" box="[433,453,693,716]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622686C4C981FA621DA2" box="[470,622,692,717]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
, is a reciprocal geometric corollary of the different foraminal lengths (
|
||
<tableCitation id="B24123B8FF956226878CC9E2F8F61D9F" box="[158,250,727,752]" captionStart="TABLE 6" captionStartId="13.[151,239,260,283]" captionTargetBox="[151,1436,372,1987]" captionTargetPageId="13" captionText="TABLE 6. Measurements of the holotype (FMNH 83796) of Hylomyscus heinrichorum, new species, and descriptive statistics for samples of the H. anselli group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations)." httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="16" pageNumber="117" tableUuid="ABBC468BFF88623B8785CA31FC0A1E34">Table 6</tableCitation>
|
||
). The shape of the incisive foramina, however, is similar in both: broader over the anterior portion, narrowing slightly over the posterior half. The dorsal notch formed between the upper zygomatic plate and rostrum appears slightly deeper and wider in specimens of
|
||
<taxonomicName id="38C36D80FF95622685F5C814FBA81C57" box="[743,932,799,824]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF95622685F5C814FAF71C57" box="[743,763,801,824]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF956226841EC82AFBA81C57" box="[780,932,799,824]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
compared with crania of
|
||
<taxonomicName id="38C36D80FF95622683C6C814FD481C57" box="[1236,1348,799,824]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF95622683C6C814FCE41C57" box="[1236,1256,801,824]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622683EBC82AFD481C57" box="[1273,1348,799,824]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF9562268293C814F93F1C33" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF9562268293C814FD991C57" box="[1409,1429,801,824]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF9562268785C870F93F1C33" box="[151,307,837,860]" italics="true" pageId="16" pageNumber="117">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="67F80A86FF9562268657C871F9801C32" box="[325,396,836,861]" captionStart="FIGURE 6" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[265,1313,213,1481]" captionTargetId="figure@14.[256,1331,193,1499]" captionTargetPageId="14" captionText="FIGURE 6. Dorsal (top row) and ventral (bottom row) views of adult crania (ca. 2.75 ×) representing members of the Hylomyscus anselli species group: A, H. arcimontensis (FMNH 147260; ONL = 24.9 mm), a male from Tanzania, Tanga Region, West Usambara Mts; B, H. heinrichorum, new species (FMNH 83796, holotype; ONL = 27.3 mm), a male from Angola, Provincia Huambo, Mt Moco; C, H. anselli (FMNH 171367; ONL = 28.1 mm), a male from Tanzania, Rukwa Region, Mbizi FR." httpUri="https://zenodo.org/record/235108/files/figure.png" pageId="16" pageNumber="117">Fig. 6</figureCitation>
|
||
), a visual impression consistent with the actually broader zygomatic plate recorded in the former (
|
||
<tableCitation id="B24123B8FF95622687E0C852F9451CEF" box="[242,329,871,896]" captionStart="TABLE 6" captionStartId="13.[151,239,260,283]" captionTargetBox="[151,1436,372,1987]" captionTargetPageId="13" captionText="TABLE 6. Measurements of the holotype (FMNH 83796) of Hylomyscus heinrichorum, new species, and descriptive statistics for samples of the H. anselli group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations)." httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="16" pageNumber="117" tableUuid="ABBC468BFF88623B8785CA31FC0A1E34">Table 6</tableCitation>
|
||
) and the important contribution of this variable (BZP) to between-group discrimination (
|
||
<tableCitation id="B24123B8FF9562268224C852FD831CEF" box="[1334,1423,871,896]" captionStart="TABLE 4" captionStartId="8.[151,239,978,1001]" captionTargetBox="[159,1321,1125,1976]" captionTargetPageId="8" captionText="TABLE 4. Results of principal component analysis performed on 16 log-transformed measurements of Hylomyscus anselli (N = 12) and the Angolan sample (N = 13), complemented with one-way ANOVAs of input variables with OTU as categorical effect (df = 1 / 26). See Fig. 2 C for a scatterplot of principal component scores and Materials and Methods for variable abbreviations." httpUri="http://table.plazi.org/id/ABBC468BFF8D623E8785C8E7F9BC1B26" pageId="16" pageNumber="117" tableUuid="ABBC468BFF8D623E8785C8E7F9BC1B26">Table 4</tableCitation>
|
||
). The interorbital region of
|
||
<taxonomicName id="38C36D80FF95622686D2C8B8FA701CCA" box="[448,636,908,933]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF95622686D2C8B8F9D81CCB" box="[448,468,909,932]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622686F6C8B9FA701CCA" box="[484,636,908,933]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
conforms to that described for the
|
||
<taxonomicName id="38C36D80FF9562268303C8B8FC731CCA" box="[1041,1151,908,933]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF9562268303C8B8FC291CCB" box="[1041,1061,909,932]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF9562268327C8B9FC731CCA" box="[1077,1151,908,933]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
group: relatively narrow and amphoral in shape over its anterior portion, with post-orbital shelving weakly expressed in full and old adults, its edges lacking supraorbital ridging or beading. When examining series of skulls arrayed side-by-side (
|
||
<figureCitation id="67F80A86FF9562268231C8E1FD7E1C82" box="[1315,1394,980,1005]" captionStart="FIGURE 6" captionStartId="14.[151,250,1520,1542]" captionTargetBox="[265,1313,213,1481]" captionTargetId="figure@14.[256,1331,193,1499]" captionTargetPageId="14" captionText="FIGURE 6. Dorsal (top row) and ventral (bottom row) views of adult crania (ca. 2.75 ×) representing members of the Hylomyscus anselli species group: A, H. arcimontensis (FMNH 147260; ONL = 24.9 mm), a male from Tanzania, Tanga Region, West Usambara Mts; B, H. heinrichorum, new species (FMNH 83796, holotype; ONL = 27.3 mm), a male from Angola, Provincia Huambo, Mt Moco; C, H. anselli (FMNH 171367; ONL = 28.1 mm), a male from Tanzania, Rukwa Region, Mbizi FR." httpUri="https://zenodo.org/record/235108/files/figure.png" pageId="16" pageNumber="117">Figs. 6</figureCitation>
|
||
,
|
||
<figureCitation id="67F80A86FF956226826CC8E1FD821C83" box="[1406,1422,980,1004]" captionStart="FIGURE 7" captionStartId="15.[151,250,1851,1873]" captionTargetBox="[359,1238,675,1815]" captionTargetId="figure@15.[340,1247,661,1829]" captionTargetPageId="15" captionText="FIGURE 7. Lateral views (ca. 4.25 ×) of adult crania of the same specimens portrayed in Fig. 6: A, H. arcimontensis; B, H. heinrichorum, new species; C, H. anselli." httpUri="https://zenodo.org/record/235109/files/figure.png" pageId="16" pageNumber="117">7</figureCitation>
|
||
), examples of
|
||
<taxonomicName id="38C36D80FF9562268634C8CCF99E1B7F" box="[294,402,1015,1040]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF9562268634C8CCF9361B7F" box="[294,314,1017,1040]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF956226865AC8C2F99E1B7F" box="[328,402,1015,1040]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
exhibit the longest, more attenuate rostrum, those of
|
||
<taxonomicName id="38C36D80FF95622684FBC8CCFCA41B7F" box="[1001,1192,1016,1040]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF95622684FBC8CCFBF11B7F" box="[1001,1021,1017,1040]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF956226831EC8CCFCA41B7F" box="[1036,1192,1017,1040]" italics="true" pageId="16" pageNumber="117">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
the shortest and most truncate, and those of
|
||
<taxonomicName id="38C36D80FF95622686BBCF28FA651B5A" box="[425,617,1052,1077]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF95622686BBCF28F9B11B5B" box="[425,445,1053,1076]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622686C3CF29FA651B5A" box="[465,617,1052,1077]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
appear intermediate in rostral length and shape. These macroscopic impressions find some statistical precision in the sample statistics (
|
||
<tableCitation id="B24123B8FF9562268485CF0AFBFC1B37" box="[919,1008,1087,1112]" captionStart="TABLE 6" captionStartId="13.[151,239,260,283]" captionTargetBox="[151,1436,372,1987]" captionTargetPageId="13" captionText="TABLE 6. Measurements of the holotype (FMNH 83796) of Hylomyscus heinrichorum, new species, and descriptive statistics for samples of the H. anselli group (Statistics include the sample mean, standard deviation, observed range, and sample size in parentheses; see Materials and Methods for variable abbreviations)." httpUri="http://table.plazi.org/id/ABBC468BFF88623B8785CA31FC0A1E34" pageId="16" pageNumber="117" tableUuid="ABBC468BFF88623B8785CA31FC0A1E34">Table 6</tableCitation>
|
||
), variable loadings of the ordinations performed (
|
||
<tableCitation id="B24123B8FF9562268608CF51F9761B13" box="[282,378,1124,1149]" captionStart="TABLE 1" captionStartId="3.[151,235,764,786]" captionTargetBox="[159,1321,874,1724]" captionTargetPageId="3" captionText="TABLE 1. Results of 10 - group discriminant function analysis performed on 16 log-transformed craniodental measurements of Hylomyscus anselli (N = 12), H. arcimontensis (N = 147), and the Angolan sample (N = 13). See Fig. 1 for a scatterplot of the first two canonical variates extracted and Materials and Methods for variable abbreviations." httpUri="http://table.plazi.org/id/ABBC468BFF8662358785C9C9FC341C20" pageId="16" pageNumber="117" tableUuid="ABBC468BFF8662358785C9C9FC341C20">Tables 1</tableCitation>
|
||
,
|
||
<tableCitation id="B24123B8FF9562268695CF51F99B1B13" box="[391,407,1124,1148]" captionStart="TABLE 2" captionStartId="4.[151,239,944,967]" captionTargetBox="[159,1307,1058,1909]" captionTargetPageId="4" captionText="TABLE 2. Results of principal component analysis performed on 16 log-transformed craniodental measurements of intact specimens representing Hylomyscus anselli (N = 12), H. arcimontensis (N = 147), and the Angolan sample (N = 13). See Fig. 2 A, B for scatterplots of principal component scores and Materials and Methods for variable abbreviations." httpUri="http://table.plazi.org/id/ABBC468BFF8162328785C885FD901B68" pageId="16" pageNumber="117" tableUuid="ABBC468BFF8162328785C885FD901B68">2</tableCitation>
|
||
,
|
||
<tableCitation id="B24123B8FF95622686B6CF51F9BF1B12" box="[420,435,1124,1149]" captionStart="TABLE 5" captionStartId="9.[151,239,664,687]" captionTargetBox="[159,1372,811,1662]" captionTargetPageId="9" captionText="TABLE 5. Results of ordinations performed on 16 log-transformed craniodental measurements of Hylomyscus anselli (N = 12), H. arcimontensis (N = 55), H. kerbispeterhansi (N = 6), and the Angolan sample (N = 13). See Fig. 3 A for a scatterplot of the first two canonical variates (CV) extracted and Fig. 3 B for a scatterplot of the first two principal components (PC) extracted; variable abbreviations are given in the Materials and Methods." httpUri="http://table.plazi.org/id/ABBC468BFF8C623F8785C9ADFC6E1C60" pageId="16" pageNumber="117" tableUuid="ABBC468BFF8C623F8785C9ADFC6E1C60">5</tableCitation>
|
||
), and/or one-way ANOVAs (
|
||
<tableCitation id="B24123B8FF95622685EBCF51FB5D1B13" box="[761,849,1124,1149]" captionStart="TABLE 4" captionStartId="8.[151,239,978,1001]" captionTargetBox="[159,1321,1125,1976]" captionTargetPageId="8" captionText="TABLE 4. Results of principal component analysis performed on 16 log-transformed measurements of Hylomyscus anselli (N = 12) and the Angolan sample (N = 13), complemented with one-way ANOVAs of input variables with OTU as categorical effect (df = 1 / 26). See Fig. 2 C for a scatterplot of principal component scores and Materials and Methods for variable abbreviations." httpUri="http://table.plazi.org/id/ABBC468BFF8D623E8785C8E7F9BC1B26" pageId="16" pageNumber="117" tableUuid="ABBC468BFF8D623E8785C8E7F9BC1B26">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF95622687D5CFB2FD951A62" blockId="16.[151,1437,151,1977]" pageId="16" pageNumber="117">
|
||
Specimens of
|
||
<taxonomicName id="38C36D80FF9562268663CFBCFA3E1BCF" box="[369,562,1159,1184]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF9562268663CFBCF9891BCF" box="[369,389,1161,1184]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF9562268688CFB2FA3E1BCF" box="[410,562,1159,1184]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
dentally resemble those of
|
||
<taxonomicName id="38C36D80FF9562268497CFBCFBF41BCF" box="[901,1016,1159,1184]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF9562268497CFBCFB951BCF" box="[901,921,1161,1184]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622684BFCFB2FBF41BCF" box="[941,1016,1159,1184]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF9562268328CFBCFCF31BCF" box="[1082,1279,1160,1184]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF9562268328CFBCFC421BCF" box="[1082,1102,1161,1184]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF9562268371CFBCFCF31BCF" box="[1123,1279,1161,1184]" italics="true" pageId="16" pageNumber="117">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
in the slight curvature of the upper incisors (nearly orthodont or weakly opisthodont), molar proportions, and cusp development. Pigmentation of the enamel face of the upper incisors in
|
||
<taxonomicName id="38C36D80FF95622684F5CFE4FCAA1B87" box="[999,1190,1231,1256]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF95622684F5CFE4FBF71B87" box="[999,1019,1233,1256]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF956226831CCFFAFCAA1B87" box="[1038,1190,1231,1256]" italics="true" pageId="16" pageNumber="117">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
is a more saturated, medium orange compared with pale yellow-orange in
|
||
<taxonomicName id="38C36D80FF95622685E4CFC0FB6E1A62" box="[758,866,1268,1293]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF95622685E4CFC0FB061A63" box="[758,778,1269,1292]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF956226840ACFC1FB6E1A62" box="[792,866,1268,1293]" italics="true" pageId="16" pageNumber="117">anselli</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="38C36D80FF956226848BCFC0FC541A63" box="[921,1112,1269,1292]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="arcimontensis">
|
||
<emphasis id="CDB7CA11FF956226848BCFC0FBA11A63" box="[921,941,1269,1292]" italics="true" pageId="16" pageNumber="117">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF95622684AECFC0FC541A63" box="[956,1112,1269,1292]" italics="true" pageId="16" pageNumber="117">arcimontensis</emphasis>
|
||
</taxonomicName>
|
||
; the tone is drab in all three.
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF95622687D5CE22FB781967" blockId="16.[151,1437,151,1977]" pageId="16" pageNumber="117">
|
||
<emphasis id="CDB7CA11FF95622687D5CE22F99E1A5F" bold="true" box="[199,402,1303,1328]" pageId="16" pageNumber="117">Ecological notes.</emphasis>
|
||
Other species of the
|
||
<taxonomicName id="38C36D80FF9562268581CE22FB7C1A5F" box="[659,880,1303,1328]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="anselli">
|
||
<emphasis id="CDB7CA11FF9562268581CE22FB7C1A5F" box="[659,880,1303,1328]" italics="true" pageId="16" pageNumber="117">Hylomyscus anselli</emphasis>
|
||
</taxonomicName>
|
||
group are known to be closely associated with Afromontane forest, typically found within an elevational belt of
|
||
<quantity id="383BBBE6FF95622684BFCE09FC431A3B" box="[941,1103,1340,1365]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="16" pageNumber="117" unit="m" value="1750.0" valueMax="2500.0" valueMin="1000.0">1000–2500 m</quantity>
|
||
(
|
||
<bibRefCitation id="9B526BF2FF9562268371CE09FD9B1A3A" author="Carleton" box="[1123,1431,1340,1365]" pageId="16" pageNumber="117" refString="Carleton, M. D. & Stanley, W. T. (2005) Review of the Hylomyscus denniae complex in Tanzania, with description of a new species. Proceedings of the Biological Society of Washington, 118, 619 - 646." type="journal article" year="2005">Carleton & Stanley, 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B526BF2FF9562268785CE6AF98A1A17" author="Carleton" box="[151,390,1375,1400]" pageId="16" pageNumber="117" refString="Carleton, M. D., Kerbis Peterhans, J. & Stanley, W. T. (2006) Review of the Hylomyscus denniae group (Rodentia: Muridae) in eastern Africa, with comments on the generic allocation of Epimys endorobae Heller. Proceedings of the Biological Society of Washington, 119, 293 - 325. http: // dx. doi. org / 10.2988 / 0006 - 324 X (2006) 119 [293: ROTHDG] 2.0. CO; 2" type="journal article" year="2006">
|
||
Carleton
|
||
<emphasis id="CDB7CA11FF9562268612CE54F9371A17" box="[256,315,1375,1400]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
|
||
, 2006
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B526BF2FF9562268686CE55FA8F1A17" author="Demos" box="[404,643,1375,1400]" pageId="16" pageNumber="117" refString="Demos, T. C., Agwanda, B. & Hickerson, M. J. (2014 b) Integrative taxonomy within the Hylomyscus denniae complex (Rodentia: Muridae) and a new species from Kenya. Journal of Mammalogy, 95 (1), E 1 - E 15. http: // dx. doi. org / 10.1644 / 13 - MAMM-A- 268" type="journal article" year="2014" yearSuffix="b">
|
||
Demos
|
||
<emphasis id="CDB7CA11FF95622686FFCE54FA2C1A17" box="[493,544,1375,1400]" italics="true" pageId="16" pageNumber="117">et al</emphasis>
|
||
., 2014b
|
||
</bibRefCitation>
|
||
). The habitat information recorded on the skin tag of all
|
||
<taxonomicName id="38C36D80FF9562268207CE6AFD911A17" box="[1301,1437,1375,1400]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="16" pageNumber="117" phylum="Chordata" rank="genus">
|
||
<emphasis id="CDB7CA11FF9562268207CE6AFD911A17" box="[1301,1437,1375,1400]" italics="true" pageId="16" pageNumber="117">Hylomyscus</emphasis>
|
||
</taxonomicName>
|
||
specimens collected at Mt Moco—“Evergreen wood. – High mountain region”—fits this ecological and topographical setting. Also, a photograph, taken by some member of Heinrich’s field team and labeled only Mt Moco, captures a substantial stand of closed-canopy, high forest in the vicinity of a campsite that we believe to represent Heinrich’s “High mountain” collecting locality (
|
||
<figureCitation id="67F80A86FF9562268430CEDAFB6A1967" box="[802,870,1519,1544]" captionStart="FIGURE 8" captionStartId="17.[151,250,1214,1236]" captionTargetBox="[151,1436,157,1196]" captionTargetId="figure@17.[151,1436,157,1196]" captionTargetPageId="17" captionText="FIGURE 8. Camp site labeled as Mount Moco, which we believe represents the “ High mountain region ” visited by Gerd and Hildegarde Heinrich in Sep – Oct 1954. Their camp is located amidst a variety of habitats, including an appreciable stand of tall, multi-strata forest pictured in the background. This forest may correspond to the “ Evergreen wood ” where the Heinrichs obtained the sample of small, long-tailed murine that we here describe as Hylomyscus heinrichorum, new species. Photograph by Hildegarde Heinrich; reproduced with permission of Bernd Heinrich." httpUri="https://zenodo.org/record/235110/files/figure.png" pageId="16" pageNumber="117">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF95622687D5CD21FC3F18D6" blockId="16.[151,1437,151,1977]" pageId="16" pageNumber="117">
|
||
<bibRefCitation id="9B526BF2FF95622687D5CD21F9911943" box="[199,413,1556,1581]" pageId="16" pageNumber="117" refString="Mills, M. S. L., Olmos, F., Melo, M. & Dean, W. R. J. (2011) Mount Moco: its importance to the conservation of Swierstra's Francolin Pternistis swierstrai and the Afromontane avifauna of Angola. Bird Conversation International, 21, 119 - 133. http: // dx. doi. org / 10.1017 / S 0959270910000493" type="journal article">
|
||
Mills
|
||
<emphasis id="CDB7CA11FF956226861ECD20F9441942" box="[268,328,1556,1581]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
documented the condition of Afromontane forest on Mt Moco as part of their ongoing studies of the endangered Swierstra’s Francolin (
|
||
<taxonomicName id="38C36D80FF95622685ADCD02FC2E193F" authority="Roberts" authorityName="Roberts" box="[703,1058,1591,1616]" class="Aves" family="Phasianidae" genus="Francolinus" kingdom="Animalia" order="Galliformes" pageId="16" pageNumber="117" phylum="Chordata" rank="species" species="swierstrai">
|
||
<emphasis id="CDB7CA11FF95622685ADCD02FBB3193F" box="[703,959,1591,1616]" italics="true" pageId="16" pageNumber="117">Francolinus swierstrai</emphasis>
|
||
Roberts
|
||
</taxonomicName>
|
||
), a partridge-like bird endemic to
|
||
<collectingCountry id="87D45693FF9562268785CD69F8E1191A" box="[151,237,1628,1653]" name="Angola" pageId="16" pageNumber="117">Angola</collectingCountry>
|
||
. High forest persists as small patches, the largest about 25 hectares, restricted to narrow ravines and steep valleys in the remotest parts of the mountain; these isolated fragments are distributed over an elevation of
|
||
<quantity id="383BBBE6FF9562268255CDB5F8FA19D3" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="16" pageNumber="117" unit="m" value="2200.0" valueMax="2400.0" valueMin="2000.0">
|
||
<date id="8B7D30C3FF9562268255CDB5F8C319D3" pageId="16" pageNumber="117" value="2000" valueMax="2400">2000 to 2400</date>
|
||
m
|
||
</quantity>
|
||
, a range that falls within the
|
||
<quantity id="383BBBE6FF9562268560CD91FAE919D3" box="[626,741,1700,1724]" metricMagnitude="0" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="117" unit="mm" value="1400.0">1400 mm</quantity>
|
||
annual rainfall isohyet. Characteristic genera include the Gondwanan conifer
|
||
<taxonomicName id="38C36D80FF9562268696CDF2FA07198F" box="[388,523,1735,1760]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF9562268696CDF2FA07198F" box="[388,523,1735,1760]" italics="true" pageId="16" pageNumber="117">Podocarpus</emphasis>
|
||
</taxonomicName>
|
||
along with other evergreen, flowering trees and shrubs (species of
|
||
<taxonomicName id="38C36D80FF956226823CCDF2FD98198F" box="[1326,1428,1735,1760]" class="Magnoliopsida" family="Icacinaceae" genus="Apodytes" kingdom="Plantae" order="Icacinales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF956226823CCDF2FD98198F" box="[1326,1428,1735,1760]" italics="true" pageId="16" pageNumber="117">Apodytes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9562268785CDD8F8D9186B" box="[151,213,1773,1796]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="16" pageNumber="117" phylum="Mollusca" rank="genus">
|
||
<emphasis id="CDB7CA11FF9562268785CDD8F8D9186B" box="[151,213,1773,1796]" italics="true" pageId="16" pageNumber="117">Ficus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF95622687F7CDD9F948186A" box="[229,324,1772,1797]" class="Magnoliopsida" family="Stilbaceae" genus="Halleria" kingdom="Plantae" order="Lamiales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF95622687F7CDD9F948186A" box="[229,324,1772,1797]" italics="true" pageId="16" pageNumber="117">Halleria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9562268646CDD9F972186A" box="[340,382,1772,1797]" class="Magnoliopsida" family="Aquifoliaceae" genus="Ilex" kingdom="Plantae" order="Aquifoliales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF9562268646CDD9F972186A" box="[340,382,1772,1797]" italics="true" pageId="16" pageNumber="117">Ilex</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF956226869DCDD9F9C9186A" box="[399,453,1772,1797]" class="Gastropoda" family="Limapontiidae" genus="Olea" kingdom="Animalia" order="Sacoglossa" pageId="16" pageNumber="117" phylum="Mollusca" rank="genus">
|
||
<emphasis id="CDB7CA11FF956226869DCDD9F9C9186A" box="[399,453,1772,1797]" italics="true" pageId="16" pageNumber="117">Olea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF95622686C7CDD8FA6C186B" box="[469,608,1773,1796]" class="Magnoliopsida" family="Pittosporaceae" genus="Pittosporum" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF95622686C7CDD8FA6C186B" box="[469,608,1773,1796]" italics="true" pageId="16" pageNumber="117">Pittosporum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9562268562CDD9FAD7186A" box="[624,731,1772,1797]" class="Magnoliopsida" family="Araliaceae" genus="Polyscias" kingdom="Plantae" order="Apiales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF9562268562CDD9FAD7186A" box="[624,731,1772,1797]" italics="true" pageId="16" pageNumber="117">Polyscias</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="38C36D80FF956226840CCDD8FB84186B" box="[798,904,1773,1796]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Myrtales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF956226840CCDD8FB84186B" box="[798,904,1773,1796]" italics="true" pageId="16" pageNumber="117">Syzygium</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="9B526BF2FF95622684B6CDD9FCB4186B" author="Huntley" box="[932,1208,1772,1797]" pageId="16" pageNumber="117" refString="Huntley, B. J. & Matos, E. M. (1994) Botantical diversity and its conservation in Angola. Stelitzia, 7, 53 - 74." type="journal article" year="1994">Huntley & Matos, 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B526BF2FF95622683D7CDD9FD9C186B" author="Mills" box="[1221,1424,1772,1797]" pageId="16" pageNumber="117" refString="Mills, M. S. L., Olmos, F., Melo, M. & Dean, W. R. J. (2011) Mount Moco: its importance to the conservation of Swierstra's Francolin Pternistis swierstrai and the Afromontane avifauna of Angola. Bird Conversation International, 21, 119 - 133. http: // dx. doi. org / 10.1017 / S 0959270910000493" type="journal article" year="2011">
|
||
Mills
|
||
<emphasis id="CDB7CA11FF9562268218CDD8FD48186A" box="[1290,1348,1772,1797]" italics="true" pageId="16" pageNumber="117">et al.</emphasis>
|
||
, 2011
|
||
</bibRefCitation>
|
||
). Canopy height tends to be irregular, conforming to the steep slopes and rugged valleys that retain forest coverage. Montane grasslands, both natural and anthropogenic, and Miombo woodlands, dominated by
|
||
<taxonomicName id="38C36D80FF9562268213CC01FD991822" box="[1281,1429,1844,1869]" class="Magnoliopsida" family="Fabaceae" genus="Brachystegia" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF9562268213CC01FD991822" box="[1281,1429,1844,1869]" italics="true" pageId="16" pageNumber="117">Brachystegia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9562268785CC62F91E181F" box="[151,274,1879,1904]" class="Magnoliopsida" family="Fabaceae" genus="Isoberlinia" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF9562268785CC62F91E181F" box="[151,274,1879,1904]" italics="true" pageId="16" pageNumber="117">Isoberlinia</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="38C36D80FF9562268640CC62F9EF181F" box="[338,483,1879,1904]" class="Magnoliopsida" family="Fabaceae" genus="Julbernardia" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="CDB7CA11FF9562268640CC62F9EF181F" box="[338,483,1879,1904]" italics="true" pageId="16" pageNumber="117">Julbernardia</emphasis>
|
||
</taxonomicName>
|
||
, sprawl in between the high-forest patches; the latter woodlands, a relatively mesic savanna association (Zambezian Woodland Biotic Zone), cover most of the mountain. Presumably, tracts of Afromontane forest were more extensive when Heinrich visited Mt Moco in 1954.
|
||
</paragraph>
|
||
<caption id="ABBC468BFF9462278785CF8BFB781A21" httpUri="https://zenodo.org/record/235110/files/figure.png" pageId="17" pageNumber="118" targetBox="[151,1436,157,1196]" targetPageId="17">
|
||
<paragraph id="FF7C1603FF9462278785CF8BFB781A21" blockId="17.[151,1436,1214,1358]" pageId="17" pageNumber="118">
|
||
<emphasis id="CDB7CA11FF9462278785CF8BF91F1BBB" bold="true" box="[151,275,1214,1236]" pageId="17" pageNumber="118">FIGURE 8.</emphasis>
|
||
Camp site labeled as Mount Moco, which we believe represents the “High mountain region” visited by Gerd and Hildegarde Heinrich in Sep–Oct 1954. Their camp is located amidst a variety of habitats, including an appreciable stand of tall, multi-strata forest pictured in the background. This forest may correspond to the “Evergreen wood” where the Heinrichs obtained the sample of small, long-tailed murine that we here describe as
|
||
<taxonomicName id="38C36D80FF9462278495CE2FFC9E1A5F" box="[903,1170,1306,1328]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="heinrichorum" status="sp. nov.">
|
||
<emphasis id="CDB7CA11FF9462278495CE2FFC9E1A5F" box="[903,1170,1306,1328]" italics="true" pageId="17" pageNumber="118">Hylomyscus heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D684776AFF94622783B2CE2EFD111A5F" box="[1184,1309,1306,1328]" pageId="17" pageNumber="118" rank="species">new species</taxonomicNameLabel>
|
||
. Photograph by Hildegarde Heinrich; reproduced with permission of Bernd Heinrich.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF7C1603FF94622787D5CE42F965192D" blockId="17.[151,1437,1397,2035]" pageId="17" pageNumber="118">
|
||
Most skin tags of
|
||
<taxonomicName id="38C36D80FF9462278686CE42FA431AE1" box="[404,591,1397,1422]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF9462278686CE42F9A41AE1" box="[404,424,1399,1422]" italics="true" pageId="17" pageNumber="118">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF94622786A5CE40FA431AE1" box="[439,591,1397,1422]" italics="true" pageId="17" pageNumber="118">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
from Mt Soque indicate their capture “Along mountain brook through tall grass. – Below mt. top.” Whether remnant Afromontane forest existed in proximity to Heinrich’s trap line is indeterminate from this scant description. When Mills canvassed Mt Soque for Swierstra’s Francolin in 2005, he (
|
||
<bibRefCitation id="9B526BF2FF946227878DCED7F97C1A95" author="Mills" box="[159,368,1506,1531]" pageId="17" pageNumber="118" refString="Mills, M. S. L., Olmos, F., Melo, M. & Dean, W. R. J. (2011) Mount Moco: its importance to the conservation of Swierstra's Francolin Pternistis swierstrai and the Afromontane avifauna of Angola. Bird Conversation International, 21, 119 - 133. http: // dx. doi. org / 10.1017 / S 0959270910000493" type="journal article" year="2011">
|
||
Mills
|
||
<emphasis id="CDB7CA11FF94622787F4CED6F92F1A94" box="[230,291,1506,1531]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
|
||
, 2011
|
||
</bibRefCitation>
|
||
: 6) recorded that Afromontane forest no longer existed, but a few pairs of the endangered francolin were discovered in “dense herbaceous and shrubby growth in gullies and around the mountain summit [peak at
|
||
<quantity id="383BBBE6FF94622787E5CD1FF95D192D" box="[247,337,1578,1603]" metricMagnitude="3" metricUnit="m" metricValue="2.165" pageId="17" pageNumber="118" unit="m" value="2165.0">2165 m</quantity>
|
||
].”
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF94622787D5CD78FD1C189D" blockId="17.[151,1437,1397,2035]" pageId="17" pageNumber="118">
|
||
Other rodents collected with
|
||
<taxonomicName id="38C36D80FF946227851FCD78FB3F1909" box="[525,819,1613,1638]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF946227851FCD78FB3F1909" box="[525,819,1613,1638]" italics="true" pageId="17" pageNumber="118">Hylomyscus heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
over the same range of dates at both Mt Soque (
|
||
<date id="8B7D30C3FF9462278246CD7BF90619E4" pageId="17" pageNumber="118" value="1954-08-24" valueMax="1954-08-28" valueMin="1954-08-24">24–28 Aug 1954</date>
|
||
) and Mt Moco (
|
||
<date id="8B7D30C3FF94622786DECD47FA7519E4" box="[460,633,1650,1675]" pageId="17" pageNumber="118" value="1954-10-05" valueMax="1954-10-10" valueMin="1954-10-05">5–10 Oct 1954</date>
|
||
) include
|
||
<taxonomicName id="38C36D80FF94622785F5CD46FC1319E4" authority="Kuhl" authorityName="Kuhl" box="[743,1055,1650,1675]" class="Mammalia" family="Sciuridae" genus="Funisciurus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="congicus">
|
||
<emphasis id="CDB7CA11FF94622785F5CD46FBD719E5" box="[743,987,1651,1674]" italics="true" pageId="17" pageNumber="118">Funisciurus congicus</emphasis>
|
||
Kuhl
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF946227833FCD47FD9B19E5" authority="Desmarest" authorityName="Desmarest" box="[1069,1431,1650,1675]" class="Mammalia" family="Gliridae" genus="Graphiurus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="murinus">
|
||
<emphasis id="CDB7CA11FF946227833FCD47FD1919E5" box="[1069,1301,1650,1675]" italics="true" pageId="17" pageNumber="118">Graphiurus murinus</emphasis>
|
||
Desmarest
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9462278785CDA0F9F619C1" authority="Wroughton" authorityName="Wroughton" box="[151,506,1685,1710]" class="Mammalia" family="Muridae" genus="Dendromus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="nyikae">
|
||
<emphasis id="CDB7CA11FF9462278785CDA0F96119C1" box="[151,365,1685,1710]" italics="true" pageId="17" pageNumber="118">Dendromus nyikae</emphasis>
|
||
Wroughton
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9462278519CDA3FB7E19C1" authority="Smuts" authorityName="Smuts" box="[523,882,1685,1710]" class="Mammalia" family="Muridae" genus="Grammomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="dolichurus">
|
||
<emphasis id="CDB7CA11FF9462278519CDA3FB1119C1" box="[523,797,1685,1710]" italics="true" pageId="17" pageNumber="118">Grammomys dolichurus</emphasis>
|
||
Smuts
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9462278490CDA0FD7A19C1" authority="Verheyen, Dierckx" authorityName="Verheyen, Dierckx" box="[898,1398,1685,1710]" class="Mammalia" family="Muridae" genus="Lophuromys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis id="CDB7CA11FF9462278490CDA0FC9D19C1" box="[898,1169,1685,1710]" italics="true" pageId="17" pageNumber="118">Lophuromys angolensis</emphasis>
|
||
Verheyen, Dierckx
|
||
</taxonomicName>
|
||
, & Huselmans,
|
||
<taxonomicName id="38C36D80FF946227863BCD8EFA9B19BC" authority="Bocage" authorityName="Bocage" box="[297,663,1722,1747]" class="Mammalia" family="Muridae" genus="Myomyscus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis id="CDB7CA11FF946227863BCD8EFA3E19BC" box="[297,562,1722,1747]" italics="true" pageId="17" pageNumber="118">Myomyscus angolensis</emphasis>
|
||
Bocage
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF94622785BBCD8EFC2B19BC" authority="Pucheran" authorityName="Pucheran" box="[681,1063,1722,1747]" class="Mammalia" family="Muridae" genus="Oenomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="hypoxanthus">
|
||
<emphasis id="CDB7CA11FF94622785BBCD8EFBA119BC" box="[681,941,1722,1747]" italics="true" pageId="17" pageNumber="118">Oenomys hypoxanthus</emphasis>
|
||
Pucheran
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="38C36D80FF9462278362CD8FF9091999" authority="Huet. Examples" authorityName="Huet. Examples" class="Mammalia" family="Muridae" genus="Pelomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="campanae">
|
||
<emphasis id="CDB7CA11FF9462278362CD8FFD5C19BD" box="[1136,1360,1722,1747]" italics="true" pageId="17" pageNumber="118">Pelomys campanae</emphasis>
|
||
Huet. Examples
|
||
</taxonomicName>
|
||
of
|
||
<taxonomicName id="38C36D80FF9462278624CDE8FAB01999" authority="A. Smith" authorityName="A. Smith" box="[310,700,1757,1782]" class="Mammalia" family="Muridae" genus="Aethomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="namaquensis">
|
||
<emphasis id="CDB7CA11FF9462278624CDE8FA4D1999" box="[310,577,1757,1782]" italics="true" pageId="17" pageNumber="118">Aethomys namaquensis</emphasis>
|
||
A. Smith
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF94622785DFCDE8FCAE1999" authority="Thomas & Hinton" authorityName="Thomas & Hinton" box="[717,1186,1757,1782]" class="Mammalia" family="Gliridae" genus="Graphiurus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="rupicola">
|
||
<emphasis id="CDB7CA11FF94622785DFCDE8FBB71999" box="[717,955,1757,1782]" italics="true" pageId="17" pageNumber="118">Graphiurus rupicola</emphasis>
|
||
Thomas & Hinton
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF94622783A1CDEAFD9B1999" authority="Thomas" authorityName="Thomas" box="[1203,1431,1757,1782]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="triton">
|
||
<emphasis id="CDB7CA11FF94622783A1CDEAFD221999" box="[1203,1326,1759,1782]" italics="true" pageId="17" pageNumber="118">Mus triton</emphasis>
|
||
Thomas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9462278785CC36FA061875" authority="Hill & Carter" authorityName="Hill & Carter" box="[151,522,1794,1819]" class="Mammalia" family="Muridae" genus="Otomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="cuanzensis">
|
||
<emphasis id="CDB7CA11FF9462278785CC36F9601875" box="[151,364,1795,1818]" italics="true" pageId="17" pageNumber="118">Otomys cuanzensis</emphasis>
|
||
Hill & Carter
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="38C36D80FF9462278556CC36FB7E1875" authority="Peters" authorityName="Peters" box="[580,882,1794,1819]" class="Mammalia" family="Bathyergidae" genus="Cryptomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="mechowi">
|
||
<emphasis id="CDB7CA11FF9462278556CC36FB2B1874" box="[580,807,1794,1819]" italics="true" pageId="17" pageNumber="118">Cryptomys mechowi</emphasis>
|
||
Peters
|
||
</taxonomicName>
|
||
were captured in sympatry with
|
||
<taxonomicName id="38C36D80FF94622783F0CC36FD901874" box="[1250,1436,1794,1819]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF94622783F0CC36FCFA1875" box="[1250,1270,1795,1818]" italics="true" pageId="17" pageNumber="118">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF9462278216CC37FD901874" box="[1284,1436,1794,1819]" italics="true" pageId="17" pageNumber="118">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
only at Mt Soque, but not Mt Moco; specimens of
|
||
<taxonomicName id="38C36D80FF94622785E0CC12FBF51851" authority="Smith" authorityName="Smith" box="[754,1017,1829,1854]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="minutoides">
|
||
<emphasis id="CDB7CA11FF94622785E0CC12FBA41851" box="[754,936,1829,1854]" italics="true" pageId="17" pageNumber="118">Mus minutoides</emphasis>
|
||
Smith
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9462278315CC13FD691851" authority="Lesson" authorityName="Lesson" box="[1031,1381,1829,1854]" class="Mammalia" family="Bathyergidae" genus="Cryptomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="hottentotus">
|
||
<emphasis id="CDB7CA11FF9462278315CC13FD041851" box="[1031,1288,1829,1854]" italics="true" pageId="17" pageNumber="118">Cryptomys hottentotus</emphasis>
|
||
Lesson
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="38C36D80FF9462278785CC7FFA1E180C" authority="Thomas" authorityName="Thomas" box="[151,530,1866,1891]" class="Mammalia" family="Thryonomyidae" genus="Thryonomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="species" species="gregorianus">
|
||
<emphasis id="CDB7CA11FF9462278785CC7FF9BC180D" box="[151,432,1866,1891]" italics="true" pageId="17" pageNumber="118">Thryonomys gregorianus</emphasis>
|
||
Thomas
|
||
</taxonomicName>
|
||
were also obtained at Mt Moco, but not at Mt Soque. Certain species in some of these genera have been documented in moist forest at high elevations (e.g.,
|
||
<taxonomicName id="38C36D80FF94622784F7CC58FC6418E9" box="[997,1128,1901,1926]" class="Mammalia" family="Gliridae" genus="Graphiurus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="genus">
|
||
<emphasis id="CDB7CA11FF94622784F7CC58FC6418E9" box="[997,1128,1901,1926]" italics="true" pageId="17" pageNumber="118">Graphiurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9462278364CC58FCFB18E9" box="[1142,1271,1901,1926]" class="Mammalia" family="Muridae" genus="Dendromus" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="genus">
|
||
<emphasis id="CDB7CA11FF9462278364CC58FCFB18E9" box="[1142,1271,1901,1926]" italics="true" pageId="17" pageNumber="118">Dendromus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9462278214CC5BFD9918EA" box="[1286,1429,1902,1925]" class="Mammalia" family="Muridae" genus="Grammomys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="genus">
|
||
<emphasis id="CDB7CA11FF9462278214CC5BFD9918EA" box="[1286,1429,1902,1925]" italics="true" pageId="17" pageNumber="118">Grammomys</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="38C36D80FF9462278785CCA7F92E18C4" box="[151,290,1938,1963]" class="Mammalia" family="Muridae" genus="Lophuromys" kingdom="Animalia" order="Rodentia" pageId="17" pageNumber="118" phylum="Chordata" rank="genus">
|
||
<emphasis id="CDB7CA11FF9462278785CCA7F92E18C4" box="[151,290,1938,1963]" italics="true" pageId="17" pageNumber="118">Lophuromys</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="9B526BF2FF946227862CCCA7FA2218C5" author="Stanley" box="[318,558,1938,1963]" pageId="17" pageNumber="118" refString="Stanley, W. T., Kihaule, P. M., Howell, K. M. & Hutterer, R. (1998) Small mammals of the Eastern Arc Mountains, Tanzania. Journal of East African Natural History, 87, 91 - 100." type="journal article" year="1998">
|
||
Stanley
|
||
<emphasis id="CDB7CA11FF946227868DCCA6F9D118C4" box="[415,477,1938,1963]" italics="true" pageId="17" pageNumber="118">et al.</emphasis>
|
||
, 1998
|
||
</bibRefCitation>
|
||
); still more of them are commonly associated with non-forest habitatsgrasslands, open woodlands, marshes and riverine vegetation, thickets and forest edges—environments that are also represented on Mt Moco and were around Heinrich’s campsite at the time of their survey (
|
||
<figureCitation id="67F80A86FF94622783ACCCEFFD0E189D" box="[1214,1282,2010,2035]" captionStart="FIGURE 8" captionStartId="17.[151,250,1214,1236]" captionTargetBox="[151,1436,157,1196]" captionTargetId="figure@17.[151,1436,157,1196]" captionTargetPageId="17" captionText="FIGURE 8. Camp site labeled as Mount Moco, which we believe represents the “ High mountain region ” visited by Gerd and Hildegarde Heinrich in Sep – Oct 1954. Their camp is located amidst a variety of habitats, including an appreciable stand of tall, multi-strata forest pictured in the background. This forest may correspond to the “ Evergreen wood ” where the Heinrichs obtained the sample of small, long-tailed murine that we here describe as Hylomyscus heinrichorum, new species. Photograph by Hildegarde Heinrich; reproduced with permission of Bernd Heinrich." httpUri="https://zenodo.org/record/235110/files/figure.png" pageId="17" pageNumber="118">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF97622487D5CBADFA681E2F" blockId="18.[151,1437,151,1688]" pageId="18" pageNumber="119">
|
||
At this stage of understanding,
|
||
<taxonomicName id="38C36D80FF9762248529CBA2FB6A1FDF" box="[571,870,151,176]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF9762248529CBA2FB6A1FDF" box="[571,870,151,176]" italics="true" pageId="18" pageNumber="119">Hylomyscus heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
may be reasonably characterized as endemic to Afromontane forest based on the habitat recorded for the Mt Moco series and on the known affinity of its speciesgroup relatives for highland forest. However, without knowing the exact placement of Heinrich’s trap line and traps, its strict ecological reliance upon this biome must remain inferential pending renewed field surveys. Fresh autecological study is obviously in order.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7D94588FF97622487D5CA79FA621DC7" pageId="18" pageNumber="119" type="discussion">
|
||
<paragraph id="FF7C1603FF97622487D5CA79FA621DC7" blockId="18.[151,1437,151,1688]" pageId="18" pageNumber="119">
|
||
<emphasis id="CDB7CA11FF97622487D5CA79F9371E0A" bold="true" box="[199,315,332,357]" pageId="18" pageNumber="119">Remarks.</emphasis>
|
||
To our knowledge, specimens of
|
||
<taxonomicName id="38C36D80FF97622485ABCA79FBEC1E0A" box="[697,992,332,357]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF97622485ABCA79FBEC1E0A" box="[697,992,332,357]" italics="true" pageId="18" pageNumber="119">Hylomyscus heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
have not been collected since the 1954 Heinrich expedition. Until new field assessments are conducted, the conservation status of
|
||
<taxonomicName id="38C36D80FF97622483D2CA44FD721EE7" box="[1216,1406,367,392]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF97622483D2CA44FCD81EE7" box="[1216,1236,369,392]" italics="true" pageId="18" pageNumber="119">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF97622483F4CA5AFD721EE7" box="[1254,1406,367,392]" italics="true" pageId="18" pageNumber="119">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
is plausibly considered Near Threatened, if not Vulnerable, in view of the continuing decline in extent and quality of Afromontane forest in
|
||
<collectingCountry id="87D45693FF9762248687CA82F9E41EBF" box="[405,488,439,464]" name="Angola" pageId="18" pageNumber="119">Angola</collectingCountry>
|
||
(
|
||
<bibRefCitation id="9B526BF2FF97622486E4CA82FB0F1EBF" author="Huntley" box="[502,771,439,464]" pageId="18" pageNumber="119" refString="Huntley, B. J. & Matos, E. M. (1994) Botantical diversity and its conservation in Angola. Stelitzia, 7, 53 - 74." type="journal article" year="1994">Huntley & Matos, 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9B526BF2FF976224841CCA82FBDD1EBF" author="Mills" box="[782,977,439,464]" pageId="18" pageNumber="119" refString="Mills, M. S. L., Olmos, F., Melo, M. & Dean, W. R. J. (2011) Mount Moco: its importance to the conservation of Swierstra's Francolin Pternistis swierstrai and the Afromontane avifauna of Angola. Bird Conversation International, 21, 119 - 133. http: // dx. doi. org / 10.1017 / S 0959270910000493" type="journal article" year="2011">
|
||
Mills
|
||
<emphasis id="CDB7CA11FF9762248442CA8CFB8D1EBF" box="[848,897,439,464]" italics="true" pageId="18" pageNumber="119">et al</emphasis>
|
||
., 2011
|
||
</bibRefCitation>
|
||
). Unlike the Endangered classification of Swiestra’s Francolin (IUCN Red List, accessed
|
||
<date id="8B7D30C3FF97622485AACAE9FB111E9B" box="[696,797,476,501]" pageId="18" pageNumber="119" value="2014-07">Jul 2014</date>
|
||
), a comestible, partridge-sized bird subjected to hunting pressure compounded by disappearance of its preferred habitat, populations of a small murid rodent like
|
||
<taxonomicName id="38C36D80FF9762248293C934F9231D52" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF9762248293C934FD991D77" box="[1409,1429,513,536]" italics="true" pageId="18" pageNumber="119">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF9762248785C911F9231D52" box="[151,303,548,573]" italics="true" pageId="18" pageNumber="119">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
should remain resilient to habitat loss so long as even minimal forest cover, primary or secondary, persists. The recent documentation of substantial tracts of Afromontane forest in the Namba Mts (
|
||
<bibRefCitation id="9B526BF2FF9762248200C972F8DF1DEB" author="Mills" pageId="18" pageNumber="119" refString="Mills, M. S. L., Melo, M. & Vaz, A. (2013) The Namba mountains: new hope for Afromontane forest birds in Angola. Bird Conservation International, 23, 159 - 167. http: // dx. doi. org / 10.1017 / S 095927091200024 X" type="journal article" year="2013">
|
||
Mills
|
||
<emphasis id="CDB7CA11FF976224824BC97CFD821D0F" box="[1369,1422,583,608]" italics="true" pageId="18" pageNumber="119">et al</emphasis>
|
||
., 2013
|
||
</bibRefCitation>
|
||
), situated in southern Cuanza Sul Province about
|
||
<quantity id="383BBBE6FF976224842CC959FB811DEA" box="[830,909,620,645]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="119" unit="km" value="80.0">80 km</quantity>
|
||
northwest of Mt Moco, augurs that ample populations of
|
||
<taxonomicName id="38C36D80FF976224862DC9A4F9F51DC7" box="[319,505,655,680]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF976224862DC9A4F95F1DC7" box="[319,339,657,680]" italics="true" pageId="18" pageNumber="119">H</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF9762248673C9BAF9F51DC7" box="[353,505,655,680]" italics="true" pageId="18" pageNumber="119">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
still exist.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B7D94588FF97622487D5C981F8EF19F7" pageId="18" pageNumber="119" type="etymology">
|
||
<paragraph id="FF7C1603FF97622487D5C981F8E51BAB" blockId="18.[151,1437,151,1688]" pageId="18" pageNumber="119">
|
||
<emphasis id="CDB7CA11FF97622487D5C981F9421DA2" bold="true" box="[199,334,692,717]" pageId="18" pageNumber="119">Etymology.</emphasis>
|
||
Our specific epithet honors both Gerd Hermann Heinrich (
|
||
<date id="8B7D30C3FF9762248318C981FC871DA2" box="[1034,1163,692,717]" pageId="18" pageNumber="119" value="1896" valueMax="1984">1896–1984</date>
|
||
) and Hildegarde Maria Buruvna (
|
||
<date id="8B7D30C3FF976224861AC9EDF9801D9F" box="[264,396,728,752]" pageId="18" pageNumber="119" value="1917" valueMax="2012">1917–2012</date>
|
||
), husband and wife (
|
||
<figureCitation id="67F80A86FF976224856EC9E2FACE1D9F" box="[636,706,727,752]" captionStart="FIGURE 9" captionStartId="19.[151,250,1658,1680]" captionTargetBox="[151,1436,301,1637]" captionTargetId="figure@19.[151,1436,301,1637]" captionTargetPageId="19" captionText="FIGURE 9. Gerd and Hildegarde (Hilde) Heinrich pictured in a camp setting labeled as Mount Moco. After emigrating to the U. S. following World War II, the couple undertook several prolonged collecting trips within North America and Africa, including the 1953 – 1955 expedition to Angola, during which the sample of Hylomyscus heinrichorum, new species, was procured. Hilde was cradling an orphaned monkey, while Gerd attended the final steps of specimen preparation. Photograph by anonymous; reproduced with permission of Bernd Heinrich." httpUri="https://zenodo.org/record/235111/files/figure.png" pageId="18" pageNumber="119">Fig. 9</figureCitation>
|
||
), to belatedly acknowledge their important field work in Africa conducted in the middle 1900s. Notwithstanding the enduring scientific relevance of their bird and mammal collections, these museum expeditions—to
|
||
<collectingCountry id="87D45693FF9762248599C82AFAD21C57" box="[651,734,799,824]" name="Angola" pageId="18" pageNumber="119">Angola</collectingCountry>
|
||
(
|
||
<date id="8B7D30C3FF97622485FDC82AFB7D1C57" box="[751,881,799,824]" pageId="18" pageNumber="119" value="1953" valueMax="1955">1953–1955</date>
|
||
,
|
||
<date id="8B7D30C3FF9762248492C82AFC0D1C57" box="[896,1025,799,824]" pageId="18" pageNumber="119" value="1957" valueMax="1958">1957–1958</date>
|
||
),
|
||
<collectingCountry id="87D45693FF976224830AC82AFC711C57" box="[1048,1149,799,824]" name="Tanzania" pageId="18" pageNumber="119">Tanzania</collectingCountry>
|
||
(
|
||
<date id="8B7D30C3FF976224839DC82AFD1E1C57" box="[1167,1298,799,824]" pageId="18" pageNumber="119" value="1961" valueMax="1963">1961–1963</date>
|
||
), and
|
||
<collectingCountry id="87D45693FF9762248249C82AF8D31C32" name="South Africa" pageId="18" pageNumber="119">South Africa</collectingCountry>
|
||
(1963)—unfolded as a means to an end, that end being to afford Heinrich yet another opportunity to pursue his zoological first love, the natural history and taxonomic diversity of parasitic wasps or ichneumon flies (Insecta:
|
||
<taxonomicName id="38C36D80FF9762248785C8B8F9381CCA" box="[151,308,909,933]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
|
||
:
|
||
<taxonomicName id="38C36D80FF976224862CC8B9F9FC1CCA" box="[318,496,908,933]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="family">Ichneumonidae</taxonomicName>
|
||
). While just a teenager, Heinrich devoted himself to understanding this exceedingly diverse, systematically intractable family, and it remained a lifelong passion, somehow sustained during, between, and after the World Wars that sundered normalcy in Europe. He opportunistically promoted his service as professional collector, obtaining vertebrate specimens under the auspices of large, well funded museums in order to visit exotic faraway geographies and collect never-before-seen ichneumons. His body of publications and prolific taxonomic descriptions were foundational for understanding major groups of
|
||
<taxonomicName id="38C36D80FF9762248305CF0AFCC81B37" box="[1047,1220,1087,1112]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="family">Ichneumonidae</taxonomicName>
|
||
over vast areas of their distribution (e.g.,
|
||
<bibRefCitation id="9B526BF2FF97622486B5CF51FA771B13" author="Heinrich" box="[423,635,1124,1149]" pageId="18" pageNumber="119" refString="Heinrich, G. (1934) Die Ichneumoninae von Celebes bearbeitet auf Grund der Ausbeute der Celebes Expedition G. Heinrich 1930 - 1932. Mitteilungen aus dem Zoologischen Museum in Berlin, 20, 1 - 263." type="journal article" year="1934">G. Heinrich, 1934</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9B526BF2FF9762248599CF51FAC91B13" author="Heinrich" box="[651,709,1124,1148]" pageId="18" pageNumber="119" refString="Heinrich, G. (1938) Les Ichneumonides de Madagascar. III. Ichneumonidae, Ichneumoninae. Memoires de l'Academie Malgache, 25, 1 - 139." type="journal article" year="1938">1938</bibRefCitation>
|
||
,
|
||
<date id="8B7D30C3FF97622485C4CF51FB5A1B12" box="[726,854,1124,1149]" pageId="18" pageNumber="119" value="1961" valueMax="1962">
|
||
<bibRefCitation id="9B526BF2FF97622485C4CF51FB011B12" author="Heinrich" box="[726,781,1124,1149]" pageId="18" pageNumber="119" refString="Heinrich, G. (1961 - 1962) Synopsis of Nearctic Ichneumoninae Stenopneusticae with particular reference to the Northeastern Region (Hymenoptera). Canadian Entomologist. Supplement, 886 pp. [in 8 parts]" type="book" year="1961">1961</bibRefCitation>
|
||
–1962
|
||
</date>
|
||
,
|
||
<date id="8B7D30C3FF9762248475CF51FBE61B12" box="[871,1002,1124,1149]" pageId="18" pageNumber="119" value="1967" valueMax="1968">1967–1968</date>
|
||
). For more about Gerd Heinrich, his scientific contributions and improbable life, readers should consult
|
||
<bibRefCitation id="9B526BF2FF976224849DCFBCFC391BCF" author="Weems" box="[911,1077,1159,1184]" pageId="18" pageNumber="119" refString="Weems, H. V. Jr. (1986) In memoriam Gerd H. Heinrich. Florida Entomologist, 69 (1), 281 - 282." type="journal article" year="1986">Weems (1986)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9B526BF2FF9762248350CFBCFD161BCF" author="Heinrich" box="[1090,1306,1159,1184]" pageId="18" pageNumber="119" refString="Heinrich, B. (2007) The snoring bird: my family's journey through a century of biology. Harper Collins Publishers, New York, xvii + 461 pp." type="book" year="2007">B. Heinrich (2007)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation id="9B526BF2FF9762248244CFB2F8E91BAB" author="Collar" pageId="18" pageNumber="119" refString="Collar, N. J. (2009) Pioneer of Asian ornithology: Gerd Heinrich. Birding Asia, 11, 33 - 40." type="journal article" year="2009">Collar (2009)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="FF7C1603FF97622487D5CFFAF8EF19F7" blockId="18.[151,1437,151,1688]" pageId="18" pageNumber="119">
|
||
Although Gerd Heinrich’s personal energy in the field surpassed indefatigable, the successes of his many expeditions owed as much to the abilities and equally unflagging dedication of his female companions. Prior to World War II, his wife Anneliese Machatchek and her younger sister Liselotte accompanied him to Sulawesi (
|
||
<date id="8B7D30C3FF976224878DCE09F92C1A3B" box="[159,288,1340,1364]" pageId="18" pageNumber="119" value="1930" valueMax="1932">1930–1932</date>
|
||
) and
|
||
<collectingCountry id="87D45693FF9762248670CE08F9C31A3A" box="[354,463,1341,1365]" name="Myanmar" pageId="18" pageNumber="119">Myanmar</collectingCountry>
|
||
(
|
||
<date id="8B7D30C3FF97622486F2CE09FA681A3B" box="[480,612,1340,1364]" pageId="18" pageNumber="119" value="1937" valueMax="1938">1937–1938</date>
|
||
), laboring as indispensable field assistants. After emigrating to the
|
||
<collectingCountry id="87D45693FF9762248278CE09FD901A3B" box="[1386,1436,1340,1364]" name="United States of America" pageId="18" pageNumber="119">U.S.</collectingCountry>
|
||
following World War II, Heinrich resumed fieldwork in
|
||
<collectingCountry id="87D45693FF9762248407CE6AFB661A17" box="[789,874,1375,1400]" name="Mexico" pageId="18" pageNumber="119">Mexico</collectingCountry>
|
||
(
|
||
<date id="8B7D30C3FF9762248468CE6AFBF71A17" box="[890,1019,1375,1400]" pageId="18" pageNumber="119" value="1952" valueMax="1953">1952–1953</date>
|
||
) and those various countries of Sub- Saharan Africa (
|
||
<date id="8B7D30C3FF9762248642CEB1F9D81AF2" box="[336,468,1412,1437]" pageId="18" pageNumber="119" value="1953" valueMax="1963">1953–1963</date>
|
||
), where he was joined by his second spouse Hildegarde (Hilde) Buruvna, who shared in all aspects of camp maintenance, data collection, and specimen preparation. Gerd Heinrich’s efforts as mammal collector have been individually recognized for his earlier fieldwork in Sulawesi (
|
||
<taxonomicName id="38C36D80FF976224832ECEF8F9321967" authority="Archbold and Tate, 1935" authorityName="Archbold and Tate" authorityYear="1935" class="Mammalia" family="Sciuridae" genus="Hyosciurus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="heinrichi">
|
||
<emphasis id="CDB7CA11FF976224832ECEF8FD261A8A" box="[1084,1322,1484,1509]" italics="true" pageId="18" pageNumber="119">Hyosciurus heinrichi</emphasis>
|
||
Archbold and Tate, 1935
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="38C36D80FF976224865BCEC4FB631967" authority="Tate and Archbold, 1935" authorityName="Tate and Archbold" authorityYear="1935" box="[329,879,1519,1544]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="subSpecies" species="penitus" subSpecies="heinrichi">
|
||
<emphasis id="CDB7CA11FF976224865BCEC4FA581967" box="[329,596,1519,1544]" italics="true" pageId="18" pageNumber="119">Rattus penitus heinrichi</emphasis>
|
||
Tate and Archbold, 1935
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="38C36D80FF9762248487CEC4FCE71967" authority="J. A. Allen" authorityName="J. A. Allen" box="[917,1259,1519,1544]" class="Mammalia" family="Muridae" genus="Bunomys" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="andrewsi">
|
||
<emphasis id="CDB7CA11FF9762248487CEC4FC641967" box="[917,1128,1519,1544]" italics="true" pageId="18" pageNumber="119">Bunomys andrewsi</emphasis>
|
||
J. A. Allen
|
||
</taxonomicName>
|
||
]) and
|
||
<collectingCountry id="87D45693FF976224823DCEC4FD901966" box="[1327,1436,1521,1545]" name="Myanmar" pageId="18" pageNumber="119">Myanmar</collectingCountry>
|
||
(
|
||
<taxonomicName id="38C36D80FF97622487B2CD21FAA31942" authority="Tate, 1954" authorityName="Tate" authorityYear="1954" box="[160,687,1556,1581]" class="Mammalia" family="Sciuridae" genus="Callosciurus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="subSpecies" species="griseimanus" subSpecies="heinrichi">
|
||
<emphasis id="CDB7CA11FF97622487B2CD21FA231942" box="[160,559,1556,1581]" italics="true" pageId="18" pageNumber="119">Callosciurus griseimanus heinrichi</emphasis>
|
||
<bibRefCitation id="9B526BF2FF976224852ACD20FAA31942" author="Tate" box="[568,687,1556,1581]" pageId="18" pageNumber="119" refString="Tate, G. H. H. (1954) A new squirrel from Burma. American Museum Novitates, 1676, 1 - 2." type="journal article" year="1954">Tate, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="38C36D80FF97622485C8CD20FB931942" authority="Blyth" authorityName="Blyth" box="[730,927,1556,1581]" class="Mammalia" family="Sciuridae" genus="Callosciurus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="phayrei">
|
||
<emphasis id="CDB7CA11FF97622485C8CD20FAE11943" box="[730,749,1557,1580]" italics="true" pageId="18" pageNumber="119">C</emphasis>
|
||
.
|
||
<emphasis id="CDB7CA11FF97622485EECD21FB5C1942" box="[764,848,1556,1581]" italics="true" pageId="18" pageNumber="119">phayrei</emphasis>
|
||
Blyth
|
||
</taxonomicName>
|
||
]). Our specific patronym acknowledges both Gerd and Hilde and their complementary roles in generating these significant African collections. The species name
|
||
<taxonomicName id="38C36D80FF97622487C8CD69F97E191A" box="[218,370,1628,1653]" class="Mammalia" family="Muridae" genus="Hylomyscus" kingdom="Animalia" order="Rodentia" pageId="18" pageNumber="119" phylum="Chordata" rank="species" species="heinrichorum">
|
||
<emphasis id="CDB7CA11FF97622487C8CD69F97E191A" box="[218,370,1628,1653]" italics="true" pageId="18" pageNumber="119">heinrichorum</emphasis>
|
||
</taxonomicName>
|
||
thus combines the family surname Heinrich, a proper noun, and the Latin genitive plural suffix -orum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |