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<document id="E36A4ECD7C42A0D3A616D99A90529DCC" ID-DOI="10.11646/zootaxa.3861.1.3" ID-GBIF-Dataset="28f4fb2e-c0ed-4d2c-a842-47c03fc6fb3d" ID-ISSN="1175-5326" ID-Zenodo-Dep="287094" ID-ZooBank="92A437D0-A49A-48E6-A02F-C88F8599665C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460908450667" checkinUser="plazi" docAuthor="Stroiński, Adam &amp; Świerczewski, Dariusz" docDate="2014" docId="6849879D4B14FFCCFF69FC30188D442C" docLanguage="en" docName="zt03861p075.pdf" docOrigin="Zootaxa 3861 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Griveaudus Stroiński &amp; Świerczewski, 2014, gen. nov." docType="treatment" docVersion="5" lastPageNumber="68" masterDocId="9470FFE54B15FFCBFFFEFF9419784531" masterDocTitle="Griveaudus gen. nov. (Hemiptera: Fulgoromorpha: Flatidae) from Tsaratanana Massif supports the biodiversity of montane flatids in Madagascar" masterLastPageNumber="75" masterPageNumber="61" pageNumber="62" updateTime="1698361019604" updateUser="plazi">
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<mods:title id="0B04AA09593D98EC63542B306C697BC7">Griveaudus gen. nov. (Hemiptera: Fulgoromorpha: Flatidae) from Tsaratanana Massif supports the biodiversity of montane flatids in Madagascar</mods:title>
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<mods:namePart id="E1E50B37265AA87D294B9A2C5A4B103F">Stroiński, Adam</mods:namePart>
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<mods:namePart id="B7437EB7E046A206E97F73F8FBDACDBB">Świerczewski, Dariusz</mods:namePart>
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<mods:date id="2D445318C76D812E7DF525C7D14C817E">2014</mods:date>
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<mods:number id="DF341FF15B023B7D5DB00728B11DBC63">3861</mods:number>
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<mods:identifier id="59C1F7E50679B02AD7D8D104338AC471" type="DOI">10.11646/zootaxa.3861.1.3</mods:identifier>
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<treatment id="6849879D4B14FFCCFF69FC30188D442C" ID-DOI="http://doi.org/10.5281/zenodo.6144848" ID-GBIF-Taxon="127669852" ID-Zenodo-Dep="6144848" LSID="urn:lsid:plazi:treatment:6849879D4B14FFCCFF69FC30188D442C" httpUri="http://treatment.plazi.org/id/6849879D4B14FFCCFF69FC30188D442C" lastPageId="7" lastPageNumber="68" pageId="1" pageNumber="62">
<subSubSection id="A8FA65004B14FFCAFF69FC30186046D3" pageId="1" pageNumber="62" type="nomenclature">
<paragraph id="E05F368B4B14FFCAFF69FC3018EC468F" blockId="1.[151,404,932,994]" box="[151,404,932,958]" pageId="1" pageNumber="62">
<heading id="BB1781E74B14FFCAFF69FC3018EC468F" bold="true" box="[151,404,932,958]" fontSize="10" level="2" pageId="1" pageNumber="62" reason="2">
<emphasis id="D294EA994B14FFCAFF69FC3018EC468F" bold="true" box="[151,404,932,958]" pageId="1" pageNumber="62">
<taxonomicName id="27E04D084B14FFCAFF69FC301865468C" ID-CoL="4R6B" box="[151,285,932,957]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D294EA994B14FFCAFF69FC301865468C" bold="true" box="[151,285,932,957]" italics="true" pageId="1" pageNumber="62">Griveaudus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9A757E24B14FFCAFED5FC3118EC468F" box="[299,404,933,958]" pageId="1" pageNumber="62" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E05F368B4B14FFCAFF69FC5D186046D3" blockId="1.[151,404,932,994]" box="[151,280,969,994]" pageId="1" pageNumber="62">
(
<figureCitation id="78DB2A0E4B14FFCAFF61FC5D186846D3" box="[159,272,969,994]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 10" captionStart-10="FIGURES 65 – 70" captionStart-2="FIGURES 11 – 16" captionStart-3="FIGURES 17 – 22" captionStart-4="FIGURES 23 – 28" captionStart-5="FIGURES 29 – 35" captionStart-6="FIGURES 36 – 43" captionStart-7="FIGURES 44 – 49" captionStart-8="FIGURES 50 – 55" captionStart-9="FIGURES 56 – 64" captionStartId-0="2.[151,264,1972,1995]" captionStartId-1="3.[151,264,1684,1707]" captionStartId-10="13.[151,264,1713,1736]" captionStartId-2="4.[151,264,1663,1686]" captionStartId-3="5.[151,264,1684,1707]" captionStartId-4="6.[151,264,1684,1707]" captionStartId-5="7.[151,264,1903,1926]" captionStartId-6="8.[151,264,1868,1891]" captionStartId-7="9.[151,264,1674,1697]" captionStartId-8="10.[151,264,1895,1918]" captionStartId-9="11.[151,264,1884,1907]" captionTargetBox-0="[155,1429,579,1939]" captionTargetBox-1="[151,1436,193,1653]" captionTargetBox-10="[161,1413,209,1680]" captionTargetBox-2="[151,1436,183,1643]" captionTargetBox-3="[151,1436,193,1653]" captionTargetBox-4="[151,1436,193,1654]" captionTargetBox-5="[151,1436,339,1873]" captionTargetBox-6="[176,1415,201,1832]" captionTargetBox-7="[151,1436,183,1644]" captionTargetBox-8="[151,1436,409,1870]" captionTargetBox-9="[166,1419,203,1844]" captionTargetId-0="figure@2.[151,1436,567,1951]" captionTargetId-1="figure@3.[151,1436,193,1664]" captionTargetId-10="figure@13.[151,1436,195,1692]" captionTargetId-2="figure@4.[151,1436,183,1649]" captionTargetId-3="figure@5.[151,1436,193,1664]" captionTargetId-4="figure@6.[151,1436,193,1664]" captionTargetId-5="figure@7.[151,1436,339,1882]" captionTargetId-6="figure@8.[151,1436,193,1847]" captionTargetId-7="figure@9.[151,1436,183,1654]" captionTargetId-8="figure@10.[151,1436,409,1875]" captionTargetId-9="figure@11.[151,1436,193,1863]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="13" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="FIGURES 1 – 4. Griveaudus issidiformis gen. et sp. nov., habitus, female. (1) lateral view; (2) dorsal view; (3) anterior part, dorsal view; (4) anterior part, lateral view." captionText-1="FIGURES 5 – 10. Griveaudus issidiformis gen. et sp. nov. (5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9 – 10) anterior part, dorsal view. (8 – 9) male, (5 – 7, 10) female." captionText-10="FIGURES 65 – 70. Griveaudus tsaratananae gen. et sp. nov., male. (65) periandrium, dorsal view; (66) periandrium, ventral view; (67) periandrium, dorso-lateral view; (68) periandrium, frontal view; (69) aedeagus, dorsal view; (70) aedeagus, lateral view." captionText-2="FIGURES 11 – 16. Griveaudus tsaratananae gen. et sp. nov., male. (11) anterior part, frontal view; (12 – 13) frons, general view; (14) frons, sensory and secretory structures; (15) hind tibia, apical part; (16) hind tarsomeres, ventral view." captionText-3="FIGURES 17 – 22. Griveaudus issidiformis gen. et sp. nov., female. (17) vertex, sensory and secretory structures; (18 – 20) antenna, general view; (21 – 22) antennal plate organs." captionText-4="FIGURES 23 – 28. Griveaudus issidiformis gen. et sp. nov., tegmen, female. (23) upper posterior part; (24) anterior part; (25) lower posterior part; (26) basal part; (27 – 28) sensory and secretory structures: (27) tegmen; (28) pronotum." captionText-5="FIGURES 29 – 35. Griveaudus issidiformis gen. et sp. nov. (29) male, abdomen, lateral view; (30) male, genital capsule, lateral view; (31) male, genital capsule, dorsal view; (32) male, genital capsule, ventral view; (33) female, bursa copulatrix; (34) female, bursa copulatrix, cells; (35) female, spermatheca." captionText-6="FIGURES 36 – 43. Griveaudus issidiformis gen. et sp. nov., male. (36) pygofer, lateral view; (37) anal tube, lateral view; (38) genital style, lateral view; (39) anal tube, dorsal view; (40) periandrium, dorsal view; (41) periandrium, dorso-lateral view; (42) periandrium, lateral view, right side; (43) periandrium, lateral view, left side." captionText-7="FIGURES 44 – 49. Griveaudus issidiformis gen. et sp. nov., female. (44) abdomen, ventral view; (45 – 47) pregenital sternite: (45) ventral view, (46) lateral view, (47) posterior view; (46 – 47) pregenital sternite, ventral view, gonoplacs, frontal view; (48 – 49) sternites with sensory structures." captionText-8="FIGURES 50 – 55. Griveaudus issidiformis gen. et sp. nov., female. (50) abdomen, lateral view; (51) genital terminalia, lateral view; (52) genital terminalia, posterior view; (53) posterior margin of the gonoplac, posterior view; (54) posterior margin of pregenital sternite and gonapophysis VIII and endogonocoxal process, basal part, ventral view; (55) endogonocoxal process, microsculpture." captionText-9="FIGURES 56 – 64. Griveaudus issidiformis gen. et sp. nov., female. (56) pregenital sternite; (57) anal tube, dorsal view; (58) terminalia, lateral view; (59) terminalia, latero-ventral view; (60) gonoplac, ventral view; (61) gonapophysis VIII, lateral view; (62) gonapophysis VIII, latero-dorsal view; (63 – 64) gonapophyses IX and gonospiculum bridge: (63) lateral view, (64) dorsal view." httpUri-0="https://zenodo.org/record/287095/files/figure.png" httpUri-1="https://zenodo.org/record/287096/files/figure.png" httpUri-10="https://zenodo.org/record/287105/files/figure.png" httpUri-2="https://zenodo.org/record/287097/files/figure.png" httpUri-3="https://zenodo.org/record/287098/files/figure.png" httpUri-4="https://zenodo.org/record/287099/files/figure.png" httpUri-5="https://zenodo.org/record/287100/files/figure.png" httpUri-6="https://zenodo.org/record/287101/files/figure.png" httpUri-7="https://zenodo.org/record/287102/files/figure.png" httpUri-8="https://zenodo.org/record/287103/files/figure.png" httpUri-9="https://zenodo.org/record/287104/files/figure.png" pageId="1" pageNumber="62">Figs 1–70</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A8FA65004B14FFCAFF69FB861A674116" box="[151,799,1041,1063]" pageId="1" pageNumber="62" type="materials_examined">
<paragraph id="E05F368B4B14FFCAFF69FB861A674116" blockId="1.[151,799,1041,1063]" box="[151,799,1041,1063]" pageId="1" pageNumber="62">
<typeStatus id="3F5B88294B14FFCAFF69FB8619B14116" box="[151,201,1042,1063]" pageId="1" pageNumber="62">Type</typeStatus>
species:
<taxonomicName id="27E04D084B14FFCAFED8FB851B604116" box="[294,536,1041,1063]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="issidiformis" status="sp. nov.">
<emphasis id="D294EA994B14FFCAFED8FB851B604116" box="[294,536,1041,1063]" italics="true" pageId="1" pageNumber="62">Griveaudus issidiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C9A757E24B14FFCAFDE1FB861B084116" box="[543,624,1042,1063]" pageId="1" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="A8FA65004B14FFCAFF69FBC01D7B41A2" pageId="1" pageNumber="62" type="etymology">
<paragraph id="E05F368B4B14FFCAFF69FBC01D7B41A2" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="62">
<emphasis id="D294EA994B14FFCAFF69FBC01866415C" bold="true" box="[151,286,1108,1133]" pageId="1" pageNumber="62">Etymology.</emphasis>
The generic name is after Paul Griveaud (
<date id="945E104B4B14FFCAFD05FBC01A06415D" box="[763,894,1108,1132]" pageId="1" pageNumber="62" value="1907" valueMax="1980">1907–1980</date>
), a well-known French collector of Madagascan insects. Gender: masculine (Genetivus singularis:
<emphasis id="D294EA994B14FFCAFD36FBEF1A2941A2" box="[712,849,1147,1171]" italics="true" pageId="1" pageNumber="62">
<taxonomicName id="27E04D084B14FFCAFD36FBEF1A3541A2" box="[712,845,1147,1171]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">Griveaudus</taxonomicName>
,
</emphasis>
4th declension).
</paragraph>
</subSubSection>
<subSubSection id="A8FA65004B14FFCAFF39FB0A1AB44012" pageId="1" pageNumber="62" type="diagnosis">
<paragraph id="E05F368B4B14FFCAFF39FB0A1AB44012" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="62">
<emphasis id="D294EA994B14FFCAFF39FB0A18394186" bold="true" box="[199,321,1182,1207]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
The newly described genus
<taxonomicName id="27E04D084B14FFCAFD64FB0B1A654186" box="[666,797,1183,1207]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D294EA994B14FFCAFD64FB0B1A654186" box="[666,797,1183,1207]" italics="true" pageId="1" pageNumber="62">Griveaudus</emphasis>
</taxonomicName>
differs from other
<collectingCountry id="98F7761B4B14FFCAFBF8FB0A1DF54186" box="[1030,1165,1182,1207]" name="Madagascar" pageId="1" pageNumber="62">Madagascar</collectingCountry>
Selizini genera by the combination of the following characters: frons with short median, single carina, mesonotum smooth (frons with Yshaped carina, mesonotum with gibbosities—
<taxonomicName id="27E04D084B14FFCAFD5CFB7C1ADF41CE" authority="Melichar, 1902" authorityName="Melichar" authorityYear="1902" box="[674,935,1254,1279]" class="Insecta" family="Flatidae" genus="Urana" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="D294EA994B14FFCAFD5CFB7C1B9341CE" box="[674,747,1256,1279]" italics="true" pageId="1" pageNumber="62">Urana</emphasis>
<bibRefCitation id="84714B7A4B14FFCAFD0BFB721ADF41CE" author="Melichar" box="[757,935,1254,1279]" pageId="1" pageNumber="62" refString="Melichar, L. (1902) Monographie der Acanaloniiden und Flatiden (Homoptera) (Fortsetzung). Annalen des K. K. Naturhistorischen Hofmuseum in Wien, 17, 1 - 256." type="journal article" year="1902">Melichar, 1902</bibRefCitation>
</taxonomicName>
), vertex extremely incised medially (vertex slightly incised medially—
<taxonomicName id="27E04D084B14FFCAFE38FA9F1AB84012" authority="Stroinski et Swierczewski, 2013" authorityName="Stroinski et Swierczewski" authorityYear="2013" box="[454,960,1290,1315]" class="Insecta" family="Flatidae" genus="Peyrierasus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="D294EA994B14FFCAFE38FA9F1B344013" box="[454,588,1291,1314]" italics="true" pageId="1" pageNumber="62">Peyrierasus</emphasis>
Stroiński et Świerczewski, 2013
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="A8FA65004B14FFCCFF39FABA1A3A45C9" lastPageId="7" lastPageNumber="68" pageId="1" pageNumber="62" type="description">
<paragraph id="E05F368B4B14FFCAFF39FABA1995405A" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="62">
<emphasis id="D294EA994B14FFCAFF39FABA18214076" bold="true" box="[199,345,1326,1351]" pageId="1" pageNumber="62">Description.</emphasis>
Body robust. Head truncate, with compound eyes in dorsal view about as wide as thorax (
<figureCitation id="78DB2A0E4B14FFCAFA94FABA19BB405A" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[155,1429,579,1939]" captionTargetId="figure@2.[151,1436,567,1951]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Griveaudus issidiformis gen. et sp. nov., habitus, female. (1) lateral view; (2) dorsal view; (3) anterior part, dorsal view; (4) anterior part, lateral view." httpUri="https://zenodo.org/record/287095/files/figure.png" pageId="1" pageNumber="62">Figs 2–3</figureCitation>
,
<figureCitation id="78DB2A0E4B14FFCAFF31FAC619A7405A" box="[207,223,1362,1387]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@3.[151,1436,193,1664]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Griveaudus issidiformis gen. et sp. nov. (5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9 – 10) anterior part, dorsal view. (8 – 9) male, (5 – 7, 10) female." httpUri="https://zenodo.org/record/287096/files/figure.png" pageId="1" pageNumber="62">5</figureCitation>
).
</paragraph>
<paragraph id="E05F368B4B14FFCAFF39FAE31A8C439E" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="62">
HEAD. Vertex in form of hourglass, extremely narrow medially; posterior margin elevated, anterior margin obliterated laterally, lateral margins subparallel (
<figureCitation id="78DB2A0E4B14FFCAFD49FA0E1A654082" box="[695,797,1434,1459]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@3.[151,1436,193,1664]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Griveaudus issidiformis gen. et sp. nov. (5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9 – 10) anterior part, dorsal view. (8 – 9) male, (5 – 7, 10) female." httpUri="https://zenodo.org/record/287096/files/figure.png" pageId="1" pageNumber="62">Figs 7–8</figureCitation>
), surface with sensory and secretory structures (
<figureCitation id="78DB2A0E4B14FFCAFAC5FA0E1CF74082" box="[1339,1423,1434,1459]" captionStart="FIGURES 17 – 22" captionStartId="5.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@5.[151,1436,193,1664]" captionTargetPageId="5" captionText="FIGURES 17 – 22. Griveaudus issidiformis gen. et sp. nov., female. (17) vertex, sensory and secretory structures; (18 – 20) antenna, general view; (21 – 22) antennal plate organs." httpUri="https://zenodo.org/record/287098/files/figure.png" pageId="1" pageNumber="62">Fig. 17</figureCitation>
). Frons (
<figureCitation id="78DB2A0E4B14FFCAFF1BFA2A181C40E6" box="[229,356,1470,1495]" captionStart="FIGURES 11 – 16" captionStartId="4.[151,264,1663,1686]" captionTargetBox="[151,1436,183,1643]" captionTargetId="figure@4.[151,1436,183,1649]" captionTargetPageId="4" captionText="FIGURES 11 – 16. Griveaudus tsaratananae gen. et sp. nov., male. (11) anterior part, frontal view; (12 – 13) frons, general view; (14) frons, sensory and secretory structures; (15) hind tibia, apical part; (16) hind tarsomeres, ventral view." httpUri="https://zenodo.org/record/287097/files/figure.png" pageId="1" pageNumber="62">Figs 11–13</figureCitation>
) as long as wide, the widest at the level of compound eyes; frons with short and wide protrusion in the upper part of head; disc of frons with short median carina; lateral margins of frons carinate and elevated; disc of frons irregularly rugose, with sensory and secretory organs (
<figureCitation id="78DB2A0E4B14FFCAFC92F9921ABA432E" box="[876,962,1542,1567]" captionStart="FIGURES 11 – 16" captionStartId="4.[151,264,1663,1686]" captionTargetBox="[151,1436,183,1643]" captionTargetId="figure@4.[151,1436,183,1649]" captionTargetPageId="4" captionText="FIGURES 11 – 16. Griveaudus tsaratananae gen. et sp. nov., male. (11) anterior part, frontal view; (12 – 13) frons, general view; (14) frons, sensory and secretory structures; (15) hind tibia, apical part; (16) hind tarsomeres, ventral view." httpUri="https://zenodo.org/record/287097/files/figure.png" pageId="1" pageNumber="62">Fig. 14</figureCitation>
). Compound eyes oval, with small callus placed at lower-posterior margin. Ocelli absent (
<figureCitation id="78DB2A0E4B14FFCAFD4FF9BE1A6E4372" box="[689,790,1578,1603]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@3.[151,1436,193,1664]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Griveaudus issidiformis gen. et sp. nov. (5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9 – 10) anterior part, dorsal view. (8 – 9) male, (5 – 7, 10) female." httpUri="https://zenodo.org/record/287096/files/figure.png" pageId="1" pageNumber="62">Figs 6–7</figureCitation>
). Antennal pedicel short, widened apically, with setae and plate organs mainly restricted to a hollow area at the top and partly on upper surface (
<figureCitation id="78DB2A0E4B14FFCAFB5AF9DA1C544356" box="[1188,1324,1614,1639]" captionStart="FIGURES 17 – 22" captionStartId="5.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@5.[151,1436,193,1664]" captionTargetPageId="5" captionText="FIGURES 17 – 22. Griveaudus issidiformis gen. et sp. nov., female. (17) vertex, sensory and secretory structures; (18 – 20) antenna, general view; (21 – 22) antennal plate organs." httpUri="https://zenodo.org/record/287098/files/figure.png" pageId="1" pageNumber="62">Figs 18–20</figureCitation>
). Sensilla placodea of the clover-leaf like
<typeStatus id="3F5B88294B14FFCAFE06F9E71B5043BA" box="[504,552,1651,1675]" pageId="1" pageNumber="62">type</typeStatus>
(
<figureCitation id="78DB2A0E4B14FFCAFDC9F9E61BC243BA" box="[567,698,1650,1675]" captionStart="FIGURES 17 – 22" captionStartId="5.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@5.[151,1436,193,1664]" captionTargetPageId="5" captionText="FIGURES 17 – 22. Griveaudus issidiformis gen. et sp. nov., female. (17) vertex, sensory and secretory structures; (18 – 20) antenna, general view; (21 – 22) antennal plate organs." httpUri="https://zenodo.org/record/287098/files/figure.png" pageId="1" pageNumber="62">Figs 21–22</figureCitation>
). Clypeus narrower than frons, without carinae (
<figureCitation id="78DB2A0E4B14FFCAFB2BF9E61C5F43BA" box="[1237,1319,1650,1675]" captionStart="FIGURES 11 – 16" captionStartId="4.[151,264,1663,1686]" captionTargetBox="[151,1436,183,1643]" captionTargetId="figure@4.[151,1436,183,1649]" captionTargetPageId="4" captionText="FIGURES 11 – 16. Griveaudus tsaratananae gen. et sp. nov., male. (11) anterior part, frontal view; (12 – 13) frons, general view; (14) frons, sensory and secretory structures; (15) hind tibia, apical part; (16) hind tarsomeres, ventral view." httpUri="https://zenodo.org/record/287097/files/figure.png" pageId="1" pageNumber="62">Fig. 11</figureCitation>
). Rostrum with apical segment a little shorter than subapical, apex reaching hind coxae.
</paragraph>
<paragraph id="E05F368B4B14FFCAFF39F92F18604252" blockId="1.[151,1437,1108,2035]" pageId="1" pageNumber="62">
THORAX. Pronotum distinctly longer than vertex at midline, with short, median carina placed posteriorly (
<figureCitation id="78DB2A0E4B14FFCAFA94F92E19DF43C6" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[155,1429,579,1939]" captionTargetId="figure@2.[151,1436,567,1951]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Griveaudus issidiformis gen. et sp. nov., habitus, female. (1) lateral view; (2) dorsal view; (3) anterior part, dorsal view; (4) anterior part, lateral view." httpUri="https://zenodo.org/record/287095/files/figure.png" pageId="1" pageNumber="62">Figs 3</figureCitation>
,
<figureCitation id="78DB2A0E4B14FFCAFF48F974198943C6" box="[182,241,1759,1783]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@3.[151,1436,193,1664]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Griveaudus issidiformis gen. et sp. nov. (5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9 – 10) anterior part, dorsal view. (8 – 9) male, (5 – 7, 10) female." httpUri="https://zenodo.org/record/287096/files/figure.png" pageId="1" pageNumber="62">7, 10</figureCitation>
) or in some specimens median carina absent (
<figureCitation id="78DB2A0E4B14FFCAFCF3F94A1A2C43C6" box="[781,852,1758,1783]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@3.[151,1436,193,1664]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Griveaudus issidiformis gen. et sp. nov. (5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9 – 10) anterior part, dorsal view. (8 – 9) male, (5 – 7, 10) female." httpUri="https://zenodo.org/record/287096/files/figure.png" pageId="1" pageNumber="62">Fig. 9</figureCitation>
); disc of pronotum medially depressed, postocular eminences absent. Mesonotum deltoid (
<figureCitation id="78DB2A0E4B14FFCAFDAEF8961BE0422A" box="[592,664,1794,1819]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[155,1429,579,1939]" captionTargetId="figure@2.[151,1436,567,1951]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Griveaudus issidiformis gen. et sp. nov., habitus, female. (1) lateral view; (2) dorsal view; (3) anterior part, dorsal view; (4) anterior part, lateral view." httpUri="https://zenodo.org/record/287095/files/figure.png" pageId="1" pageNumber="62">Figs 3</figureCitation>
,
<figureCitation id="78DB2A0E4B14FFCAFD5AF8971B98422A" box="[676,736,1795,1819]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@3.[151,1436,193,1664]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Griveaudus issidiformis gen. et sp. nov. (5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9 – 10) anterior part, dorsal view. (8 – 9) male, (5 – 7, 10) female." httpUri="https://zenodo.org/record/287096/files/figure.png" pageId="1" pageNumber="62">9–10</figureCitation>
); disc anteriorly with median groove and short obsolete carina; lateral carinae internally arcuate and elevated, reaching posterior margin; lateral parts of mesonotum without gibbosities.
</paragraph>
<paragraph id="E05F368B4B14FFC9FF39F8E41CFB445C" blockId="1.[151,1437,1108,2035]" lastBlockId="2.[151,1436,151,515]" lastPageId="2" lastPageNumber="63" pageId="1" pageNumber="62">
Tegmen (
<figureCitation id="78DB2A0E4B14FFCAFECCF8FA18E542B6" box="[306,413,1902,1927]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1972,1995]" captionTargetBox="[155,1429,579,1939]" captionTargetId="figure@2.[151,1436,567,1951]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Griveaudus issidiformis gen. et sp. nov., habitus, female. (1) lateral view; (2) dorsal view; (3) anterior part, dorsal view; (4) anterior part, lateral view." httpUri="https://zenodo.org/record/287095/files/figure.png" pageId="1" pageNumber="62">Figs 1, 4</figureCitation>
,
<figureCitation id="78DB2A0E4B14FFCAFE52F8FA18C442B6" box="[428,444,1902,1927]" captionStart="FIGURES 5 – 10" captionStartId="3.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1653]" captionTargetId="figure@3.[151,1436,193,1664]" captionTargetPageId="3" captionText="FIGURES 5 – 10. Griveaudus issidiformis gen. et sp. nov. (5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9 – 10) anterior part, dorsal view. (8 – 9) male, (5 – 7, 10) female." httpUri="https://zenodo.org/record/287096/files/figure.png" pageId="1" pageNumber="62">6</figureCitation>
,
<figureCitation id="78DB2A0E4B14FFCAFE35F8FB1B6A42B6" box="[459,530,1903,1927]" captionStart="FIGURES 23 – 28" captionStartId="6.[151,264,1684,1707]" captionTargetBox="[151,1436,193,1654]" captionTargetId="figure@6.[151,1436,193,1664]" captionTargetPageId="6" captionText="FIGURES 23 – 28. Griveaudus issidiformis gen. et sp. nov., tegmen, female. (23) upper posterior part; (24) anterior part; (25) lower posterior part; (26) basal part; (27 – 28) sensory and secretory structures: (27) tegmen; (28) pronotum." httpUri="https://zenodo.org/record/287099/files/figure.png" pageId="1" pageNumber="62">23–28</figureCitation>
) coriaceous and convex, with well visible venation and bulla, with apical line only; transverse veinlets forming irregular net on whole tegmen. Costal and posterior margins arcuate, costal and sutural angles bluntly rounded; postclaval sutural margin absent. Costal area narrower than costal cell, with transverse veinlets, terminating at the level of end of clavus. Costal cell with several transverse veinlets. Basal cell long and narrow. Sc+R forked before bulla and distinctly basad of M forking; ScRA elevated,
<collectingCountry id="98F7761B4B17FFC9FB44FF0C1DA4459E" box="[1210,1244,152,175]" name="Poland" pageId="2" pageNumber="63">RP</collectingCountry>
obsolete basally, M fork before half of tegmen; M1+2 and M3+4 forks at the same level in apical part of tegmen; CuA bifurcated near the midlength of tegmen. Claval veins Pcu and A1 fused before end of clavus, vein A1 strongly elevated; transverse veinlets between Pcu and CuP. Terminal forks in subapical part of tegmen. Terminals: CuA—2, M1+2—2, M3+4—2, RP—2, ScRA—2. Sensory and secretory structures present on the whole surface of tegmen. Tubercles with concentration on costal area, alongside apical margin, between basal
<collectingCountry id="98F7761B4B17FFC9FC62FEC11AC6445D" box="[924,958,341,364]" name="Poland" pageId="2" pageNumber="63">RP</collectingCountry>
and M veins and in basal part of clavus.
</paragraph>
<paragraph id="E05F368B4B17FFC9FF39FEED18354732" blockId="2.[151,1436,151,515]" pageId="2" pageNumber="63">
Femora shorter than tibiae; hind tibia arcuate and partly flattened laterally with 2 lateral spines placed after midlength, apically with row of well-developed 7 teeth (
<figureCitation id="78DB2A0E4B17FFC9FCF1FE0B1A1A4489" box="[783,866,415,440]" captionStart="FIGURES 11 – 16" captionStartId="4.[151,264,1663,1686]" captionTargetBox="[151,1436,183,1643]" captionTargetId="figure@4.[151,1436,183,1649]" captionTargetPageId="4" captionText="FIGURES 11 – 16. Griveaudus tsaratananae gen. et sp. nov., male. (11) anterior part, frontal view; (12 – 13) frons, general view; (14) frons, sensory and secretory structures; (15) hind tibia, apical part; (16) hind tarsomeres, ventral view." httpUri="https://zenodo.org/record/287097/files/figure.png" pageId="2" pageNumber="63">Fig. 15</figureCitation>
); basitarsomere as long as cumulative length of 2nd and 3rd tarsomeres with row of apical spines in formula 2 (longer) + 6–7 (shorter), second tarsomere with 2 lateral spines (
<figureCitation id="78DB2A0E4B17FFC9FF12FE7E18474732" box="[236,319,490,515]" captionStart="FIGURES 11 – 16" captionStartId="4.[151,264,1663,1686]" captionTargetBox="[151,1436,183,1643]" captionTargetId="figure@4.[151,1436,183,1649]" captionTargetPageId="4" captionText="FIGURES 11 – 16. Griveaudus tsaratananae gen. et sp. nov., male. (11) anterior part, frontal view; (12 – 13) frons, general view; (14) frons, sensory and secretory structures; (15) hind tibia, apical part; (16) hind tarsomeres, ventral view." httpUri="https://zenodo.org/record/287097/files/figure.png" pageId="2" pageNumber="63">Fig. 16</figureCitation>
).
</paragraph>
<caption id="B49F66034B17FFC9FF69F8201B3B42D8" httpUri="https://zenodo.org/record/287095/files/figure.png" pageId="2" pageNumber="63" targetBox="[155,1429,579,1939]" targetPageId="2">
<paragraph id="E05F368B4B17FFC9FF69F8201B3B42D8" blockId="2.[151,1436,1972,2025]" pageId="2" pageNumber="63">
<emphasis id="D294EA994B17FFC9FF69F820184442F8" bold="true" box="[151,316,1972,1995]" pageId="2" pageNumber="63">FIGURES 1–4.</emphasis>
<taxonomicName id="27E04D084B17FFC9FEBDF8201B4F42FB" box="[323,567,1972,1994]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="issidiformis">
<emphasis id="D294EA994B17FFC9FEBDF8201B4F42FB" box="[323,567,1972,1994]" italics="true" pageId="2" pageNumber="63">Griveaudus issidiformis</emphasis>
</taxonomicName>
<emphasis id="D294EA994B17FFC9FDC1F8211B9942FB" bold="true" box="[575,737,1973,1994]" pageId="2" pageNumber="63">gen. et sp. nov.</emphasis>
, habitus, female. (1) lateral view; (2) dorsal view; (3) anterior part, dorsal view; (4) anterior part, lateral view.
</paragraph>
</caption>
<caption id="B49F66034B16FFC8FF69F9001DB443FB" httpUri="https://zenodo.org/record/287096/files/figure.png" pageId="3" pageNumber="64" targetBox="[151,1436,193,1653]" targetPageId="3">
<paragraph id="E05F368B4B16FFC8FF69F9001DB443FB" blockId="3.[151,1436,1684,1738]" pageId="3" pageNumber="64">
<emphasis id="D294EA994B16FFC8FF69F9001830439B" bold="true" box="[151,328,1684,1707]" pageId="3" pageNumber="64">FIGURES 5–10.</emphasis>
<taxonomicName id="27E04D084B16FFC8FEB1F9011B3A439A" box="[335,578,1685,1707]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="issidiformis">
<emphasis id="D294EA994B16FFC8FEB1F9011B3A439A" box="[335,578,1685,1707]" italics="true" pageId="3" pageNumber="64">Griveaudus issidiformis</emphasis>
</taxonomicName>
<emphasis id="D294EA994B16FFC8FDB4F9021B92439A" bold="true" box="[586,746,1686,1707]" pageId="3" pageNumber="64">gen. et sp. nov.</emphasis>
(5) habitus, dorsal view; (6) habitus, lateral view; (7) anterior part, dorso-lateral view; (8) vertex, dorsal view; (9–10) anterior part, dorsal view. (8–9) male, (5–7, 10) female.
</paragraph>
</caption>
<paragraph id="E05F368B4B16FFC8FF39F9661A4C4263" blockId="3.[151,1436,1777,2018]" pageId="3" pageNumber="64">
MALE. Anal tube (in lateral view,
<figureCitation id="78DB2A0E4B16FFC8FDA0F9651B9A423B" box="[606,738,1777,1802]" captionStart="FIGURES 29 – 35" captionStartId="7.[151,264,1903,1926]" captionTargetBox="[151,1436,339,1873]" captionTargetId="figure@7.[151,1436,339,1882]" captionTargetPageId="7" captionText="FIGURES 29 – 35. Griveaudus issidiformis gen. et sp. nov. (29) male, abdomen, lateral view; (30) male, genital capsule, lateral view; (31) male, genital capsule, dorsal view; (32) male, genital capsule, ventral view; (33) female, bursa copulatrix; (34) female, bursa copulatrix, cells; (35) female, spermatheca." httpUri="https://zenodo.org/record/287100/files/figure.png" pageId="3" pageNumber="64">Figs 29–30</figureCitation>
,
<figureCitation id="78DB2A0E4B16FFC8FD0CF9661A77423B" box="[754,783,1778,1802]" captionStart="FIGURES 36 – 43" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[176,1415,201,1832]" captionTargetId="figure@8.[151,1436,193,1847]" captionTargetPageId="8" captionText="FIGURES 36 – 43. Griveaudus issidiformis gen. et sp. nov., male. (36) pygofer, lateral view; (37) anal tube, lateral view; (38) genital style, lateral view; (39) anal tube, dorsal view; (40) periandrium, dorsal view; (41) periandrium, dorso-lateral view; (42) periandrium, lateral view, right side; (43) periandrium, lateral view, left side." httpUri="https://zenodo.org/record/287101/files/figure.png" pageId="3" pageNumber="64">37</figureCitation>
) elongate, distinctly tapering apicad; anus placed about midlength. Anal tube (in dorsal view,
<figureCitation id="78DB2A0E4B16FFC8FDBDF8811BED421F" box="[579,661,1813,1838]" captionStart="FIGURES 29 – 35" captionStartId="7.[151,264,1903,1926]" captionTargetBox="[151,1436,339,1873]" captionTargetId="figure@7.[151,1436,339,1882]" captionTargetPageId="7" captionText="FIGURES 29 – 35. Griveaudus issidiformis gen. et sp. nov. (29) male, abdomen, lateral view; (30) male, genital capsule, lateral view; (31) male, genital capsule, dorsal view; (32) male, genital capsule, ventral view; (33) female, bursa copulatrix; (34) female, bursa copulatrix, cells; (35) female, spermatheca." httpUri="https://zenodo.org/record/287100/files/figure.png" pageId="3" pageNumber="64">Fig. 31</figureCitation>
,
<figureCitation id="78DB2A0E4B16FFC8FD5FF8821BC6421F" box="[673,702,1814,1838]" captionStart="FIGURES 36 – 43" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[176,1415,201,1832]" captionTargetId="figure@8.[151,1436,193,1847]" captionTargetPageId="8" captionText="FIGURES 36 – 43. Griveaudus issidiformis gen. et sp. nov., male. (36) pygofer, lateral view; (37) anal tube, lateral view; (38) genital style, lateral view; (39) anal tube, dorsal view; (40) periandrium, dorsal view; (41) periandrium, dorso-lateral view; (42) periandrium, lateral view, right side; (43) periandrium, lateral view, left side." httpUri="https://zenodo.org/record/287101/files/figure.png" pageId="3" pageNumber="64">39</figureCitation>
) elongate, basal part distinctly narrower than median part; apical margin with shallow incision; anus placed about midlength.
</paragraph>
<paragraph id="E05F368B4B16FFC8FF39F8C91A8A42A8" blockId="3.[151,1436,1777,2018]" pageId="3" pageNumber="64">
Pygofer (in lateral view) distinctly higher than wide, narrowing dorsally (
<figureCitation id="78DB2A0E4B16FFC8FBDAF8C91DD24247" box="[1060,1194,1885,1910]" captionStart="FIGURES 29 – 35" captionStartId="7.[151,264,1903,1926]" captionTargetBox="[151,1436,339,1873]" captionTargetId="figure@7.[151,1436,339,1882]" captionTargetPageId="7" captionText="FIGURES 29 – 35. Griveaudus issidiformis gen. et sp. nov. (29) male, abdomen, lateral view; (30) male, genital capsule, lateral view; (31) male, genital capsule, dorsal view; (32) male, genital capsule, ventral view; (33) female, bursa copulatrix; (34) female, bursa copulatrix, cells; (35) female, spermatheca." httpUri="https://zenodo.org/record/287100/files/figure.png" pageId="3" pageNumber="64">Figs 29–30</figureCitation>
,
<figureCitation id="78DB2A0E4B16FFC8FB47F8C91DA14247" box="[1209,1241,1885,1910]" captionStart="FIGURES 36 – 43" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[176,1415,201,1832]" captionTargetId="figure@8.[151,1436,193,1847]" captionTargetPageId="8" captionText="FIGURES 36 – 43. Griveaudus issidiformis gen. et sp. nov., male. (36) pygofer, lateral view; (37) anal tube, lateral view; (38) genital style, lateral view; (39) anal tube, dorsal view; (40) periandrium, dorsal view; (41) periandrium, dorso-lateral view; (42) periandrium, lateral view, right side; (43) periandrium, lateral view, left side." httpUri="https://zenodo.org/record/287101/files/figure.png" pageId="3" pageNumber="64">36</figureCitation>
); dorso-posterior angle almost right; anterior margin sinuate, posterior margin weakly arcuate.
</paragraph>
<paragraph id="E05F368B4B16FFC8FF39F8311C0B42D3" blockId="3.[151,1436,1777,2018]" pageId="3" pageNumber="64">
Genital styles (in ventral view,
<figureCitation id="78DB2A0E4B16FFC8FDDAF8311B0E428F" box="[548,630,1957,1982]" captionStart="FIGURES 29 – 35" captionStartId="7.[151,264,1903,1926]" captionTargetBox="[151,1436,339,1873]" captionTargetId="figure@7.[151,1436,339,1882]" captionTargetPageId="7" captionText="FIGURES 29 – 35. Griveaudus issidiformis gen. et sp. nov. (29) male, abdomen, lateral view; (30) male, genital capsule, lateral view; (31) male, genital capsule, dorsal view; (32) male, genital capsule, ventral view; (33) female, bursa copulatrix; (34) female, bursa copulatrix, cells; (35) female, spermatheca." httpUri="https://zenodo.org/record/287100/files/figure.png" pageId="3" pageNumber="64">Fig. 32</figureCitation>
; in lateral view,
<figureCitation id="78DB2A0E4B16FFC8FCD0F8311A06428F" box="[814,894,1957,1982]" captionStart="FIGURES 36 – 43" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[176,1415,201,1832]" captionTargetId="figure@8.[151,1436,193,1847]" captionTargetPageId="8" captionText="FIGURES 36 – 43. Griveaudus issidiformis gen. et sp. nov., male. (36) pygofer, lateral view; (37) anal tube, lateral view; (38) genital style, lateral view; (39) anal tube, dorsal view; (40) periandrium, dorsal view; (41) periandrium, dorso-lateral view; (42) periandrium, lateral view, right side; (43) periandrium, lateral view, left side." httpUri="https://zenodo.org/record/287101/files/figure.png" pageId="3" pageNumber="64">Fig. 38</figureCitation>
) longer than wide and bearing distinct, long and sharp capitulum; upper margin near base of capitulum strongly convex; ventro-posterior angle widely rounded.
</paragraph>
<caption id="B49F66034B11FFCFFF69F9EB1C764384" httpUri="https://zenodo.org/record/287097/files/figure.png" pageId="4" pageNumber="65" targetBox="[151,1436,183,1643]" targetPageId="4">
<paragraph id="E05F368B4B11FFCFFF69F9EB1C764384" blockId="4.[151,1436,1663,1717]" pageId="4" pageNumber="65">
<emphasis id="D294EA994B11FFCFFF69F9EB182D43A4" bold="true" box="[151,341,1663,1686]" pageId="4" pageNumber="65">FIGURES 11–16.</emphasis>
<taxonomicName id="27E04D084B11FFCFFEA0F9141B1A43A7" box="[350,610,1664,1686]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="tsaratananae">
<emphasis id="D294EA994B11FFCFFEA0F9141B1A43A7" box="[350,610,1664,1686]" italics="true" pageId="4" pageNumber="65">Griveaudus tsaratananae</emphasis>
</taxonomicName>
<emphasis id="D294EA994B11FFCFFD92F9141A6B43A4" bold="true" box="[620,787,1664,1685]" pageId="4" pageNumber="65">gen. et sp. nov.</emphasis>
, male. (11) anterior part, frontal view; (12–13) frons, general view; (14) frons, sensory and secretory structures; (15) hind tibia, apical part; (16) hind tarsomeres, ventral view.
</paragraph>
</caption>
<paragraph id="E05F368B4B11FFCFFF39F9481A5642C0" blockId="4.[151,1436,1756,2033]" pageId="4" pageNumber="65">
Phallic complex. Periandrium closed basally, open dorsally with long lateral split (
<figureCitation id="78DB2A0E4B11FFCFFB9FF9481D9B43C4" box="[1121,1251,1756,1781]" captionStart="FIGURES 36 – 43" captionStartId="8.[151,264,1868,1891]" captionTargetBox="[176,1415,201,1832]" captionTargetId="figure@8.[151,1436,193,1847]" captionTargetPageId="8" captionText="FIGURES 36 – 43. Griveaudus issidiformis gen. et sp. nov., male. (36) pygofer, lateral view; (37) anal tube, lateral view; (38) genital style, lateral view; (39) anal tube, dorsal view; (40) periandrium, dorsal view; (41) periandrium, dorso-lateral view; (42) periandrium, lateral view, right side; (43) periandrium, lateral view, left side." httpUri="https://zenodo.org/record/287101/files/figure.png" pageId="4" pageNumber="65">Figs 40–43</figureCitation>
,
<figureCitation id="78DB2A0E4B11FFCFFB10F9481C4043C4" box="[1262,1336,1756,1781]" captionStart="FIGURES 65 – 70" captionStartId="13.[151,264,1713,1736]" captionTargetBox="[161,1413,209,1680]" captionTargetId="figure@13.[151,1436,195,1692]" captionTargetPageId="13" captionText="FIGURES 65 – 70. Griveaudus tsaratananae gen. et sp. nov., male. (65) periandrium, dorsal view; (66) periandrium, ventral view; (67) periandrium, dorso-lateral view; (68) periandrium, frontal view; (69) aedeagus, dorsal view; (70) aedeagus, lateral view." httpUri="https://zenodo.org/record/287105/files/figure.png" pageId="4" pageNumber="65">65–68</figureCitation>
). Ventral part apically tripartite with median sharp process and lateral folds; vertical keel and ventrally pointed tooth present; apical lateral folds with several teeth. Dorsal part trilobate; median part very short with vertical, bifurcate process oriented dorso-basad; lateral lobes as long as ventral part, with two groups of lateral teeth, median and apical processes; median process huge and multiforked apically, oriented basad; apical process long and narrow, well sclerotised basally with short tooth and bifurcate apically, with or without median short process. Aedeagus divided into dorsal and ventral parts (
<figureCitation id="78DB2A0E4B11FFCFFE16F8201B1542FC" box="[488,621,1972,1997]" captionStart="FIGURES 65 – 70" captionStartId="13.[151,264,1713,1736]" captionTargetBox="[161,1413,209,1680]" captionTargetId="figure@13.[151,1436,195,1692]" captionTargetPageId="13" captionText="FIGURES 65 – 70. Griveaudus tsaratananae gen. et sp. nov., male. (65) periandrium, dorsal view; (66) periandrium, ventral view; (67) periandrium, dorso-lateral view; (68) periandrium, frontal view; (69) aedeagus, dorsal view; (70) aedeagus, lateral view." httpUri="https://zenodo.org/record/287105/files/figure.png" pageId="4" pageNumber="65">Figs 69–70</figureCitation>
), lateral split extending midlenght. Dorsal part about as long as ventral part, medially with lateral lobe, apically distinctly bilobate.
</paragraph>
<caption id="B49F66034B10FFCEFF69F9001BCE43FB" httpUri="https://zenodo.org/record/287098/files/figure.png" pageId="5" pageNumber="66" targetBox="[151,1436,193,1653]" targetPageId="5">
<paragraph id="E05F368B4B10FFCEFF69F9001BCE43FB" blockId="5.[151,1435,1684,1738]" pageId="5" pageNumber="66">
<emphasis id="D294EA994B10FFCEFF69F900182E439B" bold="true" box="[151,342,1684,1707]" pageId="5" pageNumber="66">FIGURES 17–22.</emphasis>
<taxonomicName id="27E04D084B10FFCEFE9EF9011B2E439A" box="[352,598,1685,1707]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="issidiformis">
<emphasis id="D294EA994B10FFCEFE9EF9011B2E439A" box="[352,598,1685,1707]" italics="true" pageId="5" pageNumber="66">Griveaudus issidiformis</emphasis>
</taxonomicName>
<emphasis id="D294EA994B10FFCEFDA1F9021A7E439A" bold="true" box="[607,774,1686,1707]" pageId="5" pageNumber="66">gen. et sp. nov.</emphasis>
, female. (17) vertex, sensory and secretory structures; (18–20) antenna, general view; (21–22) antennal plate organs.
</paragraph>
</caption>
<paragraph id="E05F368B4B10FFCEFF39F9661D8F4263" blockId="5.[151,1436,1777,1982]" pageId="5" pageNumber="66">
FEMALE. Pregenital sternite massive, lateral lobes weakly separated (
<figureCitation id="78DB2A0E4B10FFCEFC2BF9651D2E423B" box="[981,1110,1777,1802]" captionStart="FIGURES 44 – 49" captionStartId="9.[151,264,1674,1697]" captionTargetBox="[151,1436,183,1644]" captionTargetId="figure@9.[151,1436,183,1654]" captionTargetPageId="9" captionText="FIGURES 44 – 49. Griveaudus issidiformis gen. et sp. nov., female. (44) abdomen, ventral view; (45 – 47) pregenital sternite: (45) ventral view, (46) lateral view, (47) posterior view; (46 – 47) pregenital sternite, ventral view, gonoplacs, frontal view; (48 – 49) sternites with sensory structures." httpUri="https://zenodo.org/record/287102/files/figure.png" pageId="5" pageNumber="66">Figs 44–47</figureCitation>
,
<figureCitation id="78DB2A0E4B10FFCEFB9CF9651DFA423B" box="[1122,1154,1777,1802]" captionStart="FIGURES 56 – 64" captionStartId="11.[151,264,1884,1907]" captionTargetBox="[166,1419,203,1844]" captionTargetId="figure@11.[151,1436,193,1863]" captionTargetPageId="11" captionText="FIGURES 56 – 64. Griveaudus issidiformis gen. et sp. nov., female. (56) pregenital sternite; (57) anal tube, dorsal view; (58) terminalia, lateral view; (59) terminalia, latero-ventral view; (60) gonoplac, ventral view; (61) gonapophysis VIII, lateral view; (62) gonapophysis VIII, latero-dorsal view; (63 – 64) gonapophyses IX and gonospiculum bridge: (63) lateral view, (64) dorsal view." httpUri="https://zenodo.org/record/287104/files/figure.png" pageId="5" pageNumber="66">56</figureCitation>
); anterior margin weakly concave, medially with sclerotized lobe; posterior margin medially with two bluntly triangular lobes separated by shallow concavity, sometimes shifted. Abdominal sternites with scattered sensory pits (
<figureCitation id="78DB2A0E4B10FFCEFB99F8AD1D934263" box="[1127,1259,1849,1874]" captionStart="FIGURES 44 – 49" captionStartId="9.[151,264,1674,1697]" captionTargetBox="[151,1436,183,1644]" captionTargetId="figure@9.[151,1436,183,1654]" captionTargetPageId="9" captionText="FIGURES 44 – 49. Griveaudus issidiformis gen. et sp. nov., female. (44) abdomen, ventral view; (45 – 47) pregenital sternite: (45) ventral view, (46) lateral view, (47) posterior view; (46 – 47) pregenital sternite, ventral view, gonoplacs, frontal view; (48 – 49) sternites with sensory structures." httpUri="https://zenodo.org/record/287102/files/figure.png" pageId="5" pageNumber="66">Figs 48–49</figureCitation>
).
</paragraph>
<paragraph id="E05F368B4B10FFCEFF39F8C9182D428F" blockId="5.[151,1436,1777,1982]" pageId="5" pageNumber="66">
Anal tube (in lateral view,
<figureCitation id="78DB2A0E4B10FFCEFE08F8C91B0E4247" box="[502,630,1885,1910]" captionStart="FIGURES 50 – 55" captionStartId="10.[151,264,1895,1918]" captionTargetBox="[151,1436,409,1870]" captionTargetId="figure@10.[151,1436,409,1875]" captionTargetPageId="10" captionText="FIGURES 50 – 55. Griveaudus issidiformis gen. et sp. nov., female. (50) abdomen, lateral view; (51) genital terminalia, lateral view; (52) genital terminalia, posterior view; (53) posterior margin of the gonoplac, posterior view; (54) posterior margin of pregenital sternite and gonapophysis VIII and endogonocoxal process, basal part, ventral view; (55) endogonocoxal process, microsculpture." httpUri="https://zenodo.org/record/287103/files/figure.png" pageId="5" pageNumber="66">Figs 50–51</figureCitation>
) flattened, elongate and narrow, reaching end of gonoplac; anus placed about midlength; ventral surface with long setae. Anal tube (in dorsal view,
<figureCitation id="78DB2A0E4B10FFCEFBF8F8151D2142AB" box="[1030,1113,1921,1946]" captionStart="FIGURES 56 – 64" captionStartId="11.[151,264,1884,1907]" captionTargetBox="[166,1419,203,1844]" captionTargetId="figure@11.[151,1436,193,1863]" captionTargetPageId="11" captionText="FIGURES 56 – 64. Griveaudus issidiformis gen. et sp. nov., female. (56) pregenital sternite; (57) anal tube, dorsal view; (58) terminalia, lateral view; (59) terminalia, latero-ventral view; (60) gonoplac, ventral view; (61) gonapophysis VIII, lateral view; (62) gonapophysis VIII, latero-dorsal view; (63 – 64) gonapophyses IX and gonospiculum bridge: (63) lateral view, (64) dorsal view." httpUri="https://zenodo.org/record/287104/files/figure.png" pageId="5" pageNumber="66">Fig. 57</figureCitation>
) oval or ovoid; anus placed about midlength.
</paragraph>
<caption id="B49F66034B13FFCDFF69F9001DAE43FB" httpUri="https://zenodo.org/record/287099/files/figure.png" pageId="6" pageNumber="67" targetBox="[151,1436,193,1654]" targetPageId="6">
<paragraph id="E05F368B4B13FFCDFF69F9001DAE43FB" blockId="6.[151,1436,1684,1738]" pageId="6" pageNumber="67">
<emphasis id="D294EA994B13FFCDFF69F900182D439B" bold="true" box="[151,341,1684,1707]" pageId="6" pageNumber="67">FIGURES 23–28.</emphasis>
<taxonomicName id="27E04D084B13FFCDFEA5F9011B36439A" box="[347,590,1685,1707]" class="Insecta" family="Flatidae" genus="Griveaudus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="issidiformis">
<emphasis id="D294EA994B13FFCDFEA5F9011B36439A" box="[347,590,1685,1707]" italics="true" pageId="6" pageNumber="67">Griveaudus issidiformis</emphasis>
</taxonomicName>
<emphasis id="D294EA994B13FFCDFDABF9021B8D439A" bold="true" box="[597,757,1686,1707]" pageId="6" pageNumber="67">gen. et sp. nov.</emphasis>
, tegmen, female. (23) upper posterior part; (24) anterior part; (25) lower posterior part; (26) basal part; (27–28) sensory and secretory structures: (27) tegmen; (28) pronotum.
</paragraph>
</caption>
<paragraph id="E05F368B4B13FFCDFF39F962185C4266" blockId="6.[151,1437,1782,2023]" pageId="6" pageNumber="67">
Gonoplac unilobate, laterally flattened, elongate (
<figureCitation id="78DB2A0E4B13FFCDFD0AF9621A00423E" box="[756,888,1782,1807]" captionStart="FIGURES 56 – 64" captionStartId="11.[151,264,1884,1907]" captionTargetBox="[166,1419,203,1844]" captionTargetId="figure@11.[151,1436,193,1863]" captionTargetPageId="11" captionText="FIGURES 56 – 64. Griveaudus issidiformis gen. et sp. nov., female. (56) pregenital sternite; (57) anal tube, dorsal view; (58) terminalia, lateral view; (59) terminalia, latero-ventral view; (60) gonoplac, ventral view; (61) gonapophysis VIII, lateral view; (62) gonapophysis VIII, latero-dorsal view; (63 – 64) gonapophyses IX and gonospiculum bridge: (63) lateral view, (64) dorsal view." httpUri="https://zenodo.org/record/287104/files/figure.png" pageId="6" pageNumber="67">Figs 58–59</figureCitation>
); posterior margin rounded with single row of 5- 6 well-developed teeth (
<figureCitation id="78DB2A0E4B13FFCDFE5AF88E1B5D4202" box="[420,549,1818,1843]" captionStart="FIGURES 50 – 55" captionStartId="10.[151,264,1895,1918]" captionTargetBox="[151,1436,409,1870]" captionTargetId="figure@10.[151,1436,409,1875]" captionTargetPageId="10" captionText="FIGURES 50 – 55. Griveaudus issidiformis gen. et sp. nov., female. (50) abdomen, lateral view; (51) genital terminalia, lateral view; (52) genital terminalia, posterior view; (53) posterior margin of the gonoplac, posterior view; (54) posterior margin of pregenital sternite and gonapophysis VIII and endogonocoxal process, basal part, ventral view; (55) endogonocoxal process, microsculpture." httpUri="https://zenodo.org/record/287103/files/figure.png" pageId="6" pageNumber="67">Figs 52–53</figureCitation>
,
<figureCitation id="78DB2A0E4B13FFCDFDCEF88E1B284202" box="[560,592,1818,1843]" captionStart="FIGURES 56 – 64" captionStartId="11.[151,264,1884,1907]" captionTargetBox="[166,1419,203,1844]" captionTargetId="figure@11.[151,1436,193,1863]" captionTargetPageId="11" captionText="FIGURES 56 – 64. Griveaudus issidiformis gen. et sp. nov., female. (56) pregenital sternite; (57) anal tube, dorsal view; (58) terminalia, lateral view; (59) terminalia, latero-ventral view; (60) gonoplac, ventral view; (61) gonapophysis VIII, lateral view; (62) gonapophysis VIII, latero-dorsal view; (63 – 64) gonapophyses IX and gonospiculum bridge: (63) lateral view, (64) dorsal view." httpUri="https://zenodo.org/record/287104/files/figure.png" pageId="6" pageNumber="67">60</figureCitation>
); narrow, membranous part placed alongside ventral margin, extending half of gonoplac.
</paragraph>
<paragraph id="E05F368B4B13FFCDFF39F8F61D6A42F2" blockId="6.[151,1437,1782,2023]" pageId="6" pageNumber="67">
Gonapophysis VIII sabre-shaped and laterally flattened, tapering apicad (
<figureCitation id="78DB2A0E4B13FFCDFBEEF8F61DED424A" box="[1040,1173,1890,1915]" captionStart="FIGURES 56 – 64" captionStartId="11.[151,264,1884,1907]" captionTargetBox="[166,1419,203,1844]" captionTargetId="figure@11.[151,1436,193,1863]" captionTargetPageId="11" captionText="FIGURES 56 – 64. Griveaudus issidiformis gen. et sp. nov., female. (56) pregenital sternite; (57) anal tube, dorsal view; (58) terminalia, lateral view; (59) terminalia, latero-ventral view; (60) gonoplac, ventral view; (61) gonapophysis VIII, lateral view; (62) gonapophysis VIII, latero-dorsal view; (63 – 64) gonapophyses IX and gonospiculum bridge: (63) lateral view, (64) dorsal view." httpUri="https://zenodo.org/record/287104/files/figure.png" pageId="6" pageNumber="67">Figs 61–62</figureCitation>
); apical part of ventral margin folded externally, apical part of dorsal margin with 4 teeth. Endogonocoxal process shorter than gonapophysis VIII, sabre-shaped with spiniferous microsculpture (
<figureCitation id="78DB2A0E4B13FFCDFC7CF83E1D7E42F2" box="[898,1030,1962,1987]" captionStart="FIGURES 50 – 55" captionStartId="10.[151,264,1895,1918]" captionTargetBox="[151,1436,409,1870]" captionTargetId="figure@10.[151,1436,409,1875]" captionTargetPageId="10" captionText="FIGURES 50 – 55. Griveaudus issidiformis gen. et sp. nov., female. (50) abdomen, lateral view; (51) genital terminalia, lateral view; (52) genital terminalia, posterior view; (53) posterior margin of the gonoplac, posterior view; (54) posterior margin of pregenital sternite and gonapophysis VIII and endogonocoxal process, basal part, ventral view; (55) endogonocoxal process, microsculpture." httpUri="https://zenodo.org/record/287103/files/figure.png" pageId="6" pageNumber="67">Figs 54–55</figureCitation>
).
</paragraph>
<paragraph id="E05F368B4B13FFCDFF39F85A1AF142D6" blockId="6.[151,1437,1782,2023]" box="[199,905,1998,2023]" pageId="6" pageNumber="67">
Gonaphophyses IX and gonospiculum bridge as in
<figureCitation id="78DB2A0E4B13FFCDFCFAF85A1AFC42D6" box="[772,900,1998,2023]" captionStart="FIGURES 56 – 64" captionStartId="11.[151,264,1884,1907]" captionTargetBox="[166,1419,203,1844]" captionTargetId="figure@11.[151,1436,193,1863]" captionTargetPageId="11" captionText="FIGURES 56 – 64. Griveaudus issidiformis gen. et sp. nov., female. (56) pregenital sternite; (57) anal tube, dorsal view; (58) terminalia, lateral view; (59) terminalia, latero-ventral view; (60) gonoplac, ventral view; (61) gonapophysis VIII, lateral view; (62) gonapophysis VIII, latero-dorsal view; (63 – 64) gonapophyses IX and gonospiculum bridge: (63) lateral view, (64) dorsal view." httpUri="https://zenodo.org/record/287104/files/figure.png" pageId="6" pageNumber="67">Figs 63–64</figureCitation>
.
</paragraph>
<paragraph id="E05F368B4B12FFCCFF39FF0C1A3A45C9" blockId="7.[151,1436,151,285]" pageId="7" pageNumber="68">
Bursa copulatrix of single, elongately oval, huge pouch and narrow basis (
<figureCitation id="78DB2A0E4B12FFCCFBEEFF031D1C4581" box="[1040,1124,151,176]" captionStart="FIGURES 29 – 35" captionStartId="7.[151,264,1903,1926]" captionTargetBox="[151,1436,339,1873]" captionTargetId="figure@7.[151,1436,339,1882]" captionTargetPageId="7" captionText="FIGURES 29 – 35. Griveaudus issidiformis gen. et sp. nov. (29) male, abdomen, lateral view; (30) male, genital capsule, lateral view; (31) male, genital capsule, dorsal view; (32) male, genital capsule, ventral view; (33) female, bursa copulatrix; (34) female, bursa copulatrix, cells; (35) female, spermatheca." httpUri="https://zenodo.org/record/287100/files/figure.png" pageId="7" pageNumber="68">Fig. 33</figureCitation>
); cells well visible, without ornamentation (
<figureCitation id="78DB2A0E4B12FFCCFEAAFF2818CD45E5" box="[340,437,188,213]" captionStart="FIGURES 29 – 35" captionStartId="7.[151,264,1903,1926]" captionTargetBox="[151,1436,339,1873]" captionTargetId="figure@7.[151,1436,339,1882]" captionTargetPageId="7" captionText="FIGURES 29 – 35. Griveaudus issidiformis gen. et sp. nov. (29) male, abdomen, lateral view; (30) male, genital capsule, lateral view; (31) male, genital capsule, dorsal view; (32) male, genital capsule, ventral view; (33) female, bursa copulatrix; (34) female, bursa copulatrix, cells; (35) female, spermatheca." httpUri="https://zenodo.org/record/287100/files/figure.png" pageId="7" pageNumber="68">Fig. 34</figureCitation>
). Spermatheca well developed;
<emphasis id="D294EA994B12FFCCFCB7FF281D5A45E5" box="[841,1058,188,212]" italics="true" pageId="7" pageNumber="68">ductus receptaculi</emphasis>
ribbed and widened apically,
<emphasis id="D294EA994B12FFCCFF69FF74180C45C9" box="[151,372,224,248]" italics="true" pageId="7" pageNumber="68">diverticulum ductus</emphasis>
smooth and narrow with bulba (
<figureCitation id="78DB2A0E4B12FFCCFD1FFF4B1A4D45C9" box="[737,821,223,248]" captionStart="FIGURES 29 – 35" captionStartId="7.[151,264,1903,1926]" captionTargetBox="[151,1436,339,1873]" captionTargetId="figure@7.[151,1436,339,1882]" captionTargetPageId="7" captionText="FIGURES 29 – 35. Griveaudus issidiformis gen. et sp. nov. (29) male, abdomen, lateral view; (30) male, genital capsule, lateral view; (31) male, genital capsule, dorsal view; (32) male, genital capsule, ventral view; (33) female, bursa copulatrix; (34) female, bursa copulatrix, cells; (35) female, spermatheca." httpUri="https://zenodo.org/record/287100/files/figure.png" pageId="7" pageNumber="68">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A8FA65004B12FFCCFF39FE90188D442C" box="[199,501,260,285]" pageId="7" pageNumber="68" type="distribution">
<paragraph id="E05F368B4B12FFCCFF39FE90188D442C" blockId="7.[151,1436,151,285]" box="[199,501,260,285]" pageId="7" pageNumber="68">
<emphasis id="D294EA994B12FFCCFF39FE901818442C" bold="true" box="[199,352,260,285]" pageId="7" pageNumber="68">Distribution.</emphasis>
<collectingCountry id="98F7761B4B12FFCCFE99FE901889442C" box="[359,497,260,285]" name="Madagascar" pageId="7" pageNumber="68">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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