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<mods:title id="30476B12026D3A83C233FD27EF6682FE">Pagastia (P.) donoliveri sp. nov. - a new Nearctic alpine stream chironomid species (Diptera: Chironomidae: Diamesinae) from the Beartooth Mountains, Wyoming U. S. A.</mods:title>
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<subSubSection id="D8CD657DFFC2FF9DFF50A87DFF6E7AB6" pageId="4" pageNumber="175" type="nomenclature">
<paragraph id="906836F6FFC2FF9DFF50A87DFC1B7AFE" blockId="4.[151,938,621,719]" box="[151,938,621,647]" pageId="4" pageNumber="175">
<heading id="CB20819AFFC2FF9DFF50A87DFC1B7AFE" bold="true" box="[151,938,621,647]" fontSize="11" level="1" pageId="4" pageNumber="175" reason="1">
<emphasis id="A2A3EAE4FFC2FF9DFF50A87DFC1B7AFE" bold="true" box="[151,938,621,647]" pageId="4" pageNumber="175">
Key to the known species of
<taxonomicName id="57D74D75FFC2FF9DFE3EA87DFD067AFE" ID-CoL="6C9D" ID-ENA="994279" authority="Oliver" authorityName="Oliver" authorityYear="1959" box="[505,695,621,647]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="A2A3EAE4FFC2FF9DFE3EA87DFDD07AFE" bold="true" box="[505,609,621,647]" italics="true" pageId="4" pageNumber="175">Pagastia</emphasis>
Oliver
</taxonomicName>
of Holarctic region
</emphasis>
</heading>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50A8A5FF6E7AB6" blockId="4.[151,938,621,719]" box="[151,223,693,719]" pageId="4" pageNumber="175">
<heading id="CB20819AFFC2FF9DFF50A8A5FF6E7AB6" bold="true" box="[151,223,693,719]" fontSize="11" level="1" pageId="4" pageNumber="175" reason="1">
<emphasis id="A2A3EAE4FFC2FF9DFF50A8A5FF6E7AB6" bold="true" box="[151,223,693,719]" pageId="4" pageNumber="175">Males</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D8CD657DFFC2FF9DFF50A8EAFA2D7E51" pageId="4" pageNumber="175" type="key">
<keyStep id="2B232E53FFC2FF9DFF50A8EAFA2D7B30" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50A8EAFA2C7B54" blockId="4.[151,1437,761,1576]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50A8EAFA2C7B54" pageId="4" pageNumber="175">
1. Eyes hairy. Aedeagal lobes reduced (subgenus
<taxonomicName id="57D74D75FFC2FF9DFD64A8EAFC2C7B76" authority="Sublette" authorityName="Sublette" authorityYear="1967" box="[675,925,761,783]" class="Insecta" genus="Hesperodiamesa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="A2A3EAE4FFC2FF9DFD64A8EAFCF27B76" box="[675,835,762,783]" italics="true" pageId="4" pageNumber="175">Hesperodiamesa</emphasis>
Sublette
</taxonomicName>
). Basal lobe of gonocoxite small....................................................................
<taxonomicName id="57D74D75FFC2FF9DFCA2A908FBCD7B54" ID-CoL="6TKP5" authority="(Garrett, 1925)" baseAuthorityName="Garrett" baseAuthorityYear="1925" box="[869,1148,791,813]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="sequax" subGenus="H.">
<emphasis id="A2A3EAE4FFC2FF9DFCA2A908FCC97B54" box="[869,888,792,813]" italics="true" pageId="4" pageNumber="175">P.</emphasis>
(
<emphasis id="A2A3EAE4FFC2FF9DFCB9A908FC267B54" box="[894,919,792,813]" italics="true" pageId="4" pageNumber="175">H.</emphasis>
)
<emphasis id="A2A3EAE4FFC2FF9DFC63A908FC567B54" box="[932,999,792,813]" italics="true" pageId="4" pageNumber="175">sequax</emphasis>
(Garrett, 1925)
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFB4EA907FAF77B55" author="Makarchenko, E. A." box="[1161,1350,791,813]" pageId="4" pageNumber="175" pagination="115 - 128" refId="ref3335" refString="Makarchenko, E. A. (2019) Review of the genus Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from North America, with description of P. (P.) subletteorum sp. nov. Zootaxa, 4664 (1), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4664.1.5" type="journal article" year="2019">Makarchenko 2019</bibRefCitation>
, Fig. 18)
</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50A925FA2D7B30" blockId="4.[151,1437,761,1576]" box="[151,1436,820,842]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50A925FA2D7B30" box="[151,1436,820,842]" pageId="4" pageNumber="175">
Eyes bare or pubescent. One or two aedeagal lobes present (subgenus
<taxonomicName id="57D74D75FFC2FF9DFC44A925FB917B33" ID-CoL="6C9D" authority="Oliver" authorityName="Oliver" authorityYear="1959" box="[899,1056,820,842]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="genus">
<emphasis id="A2A3EAE4FFC2FF9DFC44A925FC697B33" box="[899,984,821,842]" italics="true" pageId="4" pageNumber="175">Pagastia</emphasis>
Oliver
</taxonomicName>
). Basal lobe of gonocoxite large.... 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B232E53FFC2FF9DFF50A942FA2D7BFC" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50A942FA2D7B1E" blockId="4.[151,1437,761,1576]" box="[151,1436,849,871]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50A942FA2D7B1E" box="[151,1436,849,871]" pageId="4" pageNumber="175">2. Both median and lateral aedeagal lobes present. AR 2.5-4.2................................................... 3</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50A97FFA2D7BFC" blockId="4.[151,1437,761,1576]" box="[151,1436,879,901]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50A97FFA2D7BFC" box="[151,1436,879,901]" pageId="4" pageNumber="175"> Only lateral aedeagal lobe present. AR 1.02.42............................................................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B232E53FFC2FF9DFF50A99CFA2D7BA4" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50A99CFA2D7BD8" blockId="4.[151,1437,761,1576]" box="[151,1436,908,930]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50A99CFA2D7BD8" box="[151,1436,908,930]" pageId="4" pageNumber="175">3. Antepronotum with dorsal and lateral setae in widely separated groups. Median aedeagal lobe digitated and widest distally. 4</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50A9BAFA2D7BA4" blockId="4.[151,1437,761,1576]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50A9BAFA2D7BA4" pageId="4" pageNumber="175">
Antepronotum completely covered with setae. Median aedeagal lobe widest medially, then tapering abruptly and with the apex forming a sharp hook..........................
<taxonomicName id="57D74D75FFC2FF9DFD29A9D8FBE47BA4" ID-CoL="75KP4" authority="(Roback, 1957)" baseAuthorityName="Roback" baseAuthorityYear="1957" box="[750,1109,967,989]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="partica" subGenus="Pagastia">
<emphasis id="A2A3EAE4FFC2FF9DFD29A9D8FD4C7BA4" box="[750,765,968,989]" italics="true" pageId="4" pageNumber="175">P</emphasis>
. (
<emphasis id="A2A3EAE4FFC2FF9DFCD6A9D8FCD77BA4" box="[785,870,968,989]" italics="true" pageId="4" pageNumber="175">Pagastia</emphasis>
)
<emphasis id="A2A3EAE4FFC2FF9DFCB3A9D8FC087BA4" box="[884,953,968,989]" italics="true" pageId="4" pageNumber="175">partica</emphasis>
(Roback, 1957)
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFBA6A9D7FAAE7BA5" author="Makarchenko, E. A." box="[1121,1311,967,989]" pageId="4" pageNumber="175" pagination="115 - 128" refId="ref3335" refString="Makarchenko, E. A. (2019) Review of the genus Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from North America, with description of P. (P.) subletteorum sp. nov. Zootaxa, 4664 (1), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4664.1.5" type="journal article" year="2019">Makarchenko 2019</bibRefCitation>
, Figs.1617)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B232E53FFC2FF9DFF50A9F4FA2D7C6E" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50A9F4FA2D7B83" blockId="4.[151,1437,761,1576]" box="[151,1436,996,1018]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50A9F4FA2D7B83" box="[151,1436,996,1018]" pageId="4" pageNumber="175">
4. Gonostylus subapically with “heel”................
<emphasis id="A2A3EAE4FFC2FF9DFCD6A9F5FC917B83" box="[785,800,997,1018]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.
<taxonomicName id="57D74D75FFC2FF9DFCEBA9F4FB847B83" ID-CoL="6TKP3" authority="(Tokunaga, 1936)" baseAuthorityName="Tokunaga" baseAuthorityYear="1936" box="[812,1077,996,1018]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="nivis" subGenus="Pagastia">
(
<emphasis id="A2A3EAE4FFC2FF9DFCF3A9F5FCF27B83" box="[820,835,997,1018]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFC90A9F5FC357B83" box="[855,900,997,1018]" italics="true" pageId="4" pageNumber="175">nivis</emphasis>
(Tokunaga, 1936)
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFB86A9F4FB4F7B83" author="Makarchenko, E. A." box="[1089,1278,996,1018]" pageId="4" pageNumber="175" pagination="607 - 621" refId="ref3268" refString="Makarchenko, E. A. (2006) 3. Subfamily Diamesinae. In: Lelej, A. (Ed.), Key to the insects of Russian Far East. Vol. 6. Diptera and Siphonaptera. Pt 4. Dal'nauka, Vladivostok, pp. 253 - 276 + 468 - 480 + 607 - 621. [in Russian]" type="book chapter" year="2006">Makarchenko 2006</bibRefCitation>
, Fig. 180, 510)
</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50AE12FA2D7C6E" blockId="4.[151,1437,761,1576]" box="[151,1436,1025,1047]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AE12FA2D7C6E" box="[151,1436,1025,1047]" pageId="4" pageNumber="175"> Gonostylus subapically without “heel”.................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B232E53FFC2FF9DFF50AE0FFA2D7C09" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50AE0FFA2D7C4C" blockId="4.[151,1437,761,1576]" box="[151,1436,1055,1077]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AE0FFA2D7C4C" box="[151,1436,1055,1077]" pageId="4" pageNumber="175">
5. Apex of the gonostylus angled.................
<emphasis id="A2A3EAE4FFC2FF9DFD13AE30FD567C4C" box="[724,743,1056,1077]" italics="true" pageId="4" pageNumber="175">P.</emphasis>
<taxonomicName id="57D74D75FFC2FF9DFD2BAE0FFBF17C4C" ID-CoL="75KP3" authority="(Tshernovskij, 1949)" baseAuthorityName="Tshernovskij" baseAuthorityYear="1949" box="[748,1088,1055,1077]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="orientalis" subGenus="Pagastia">
(
<emphasis id="A2A3EAE4FFC2FF9DFD33AE30FCB27C4C" box="[756,771,1056,1077]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFCD0AE0FFCC27C4D" box="[791,883,1055,1076]" italics="true" pageId="4" pageNumber="175">orientalis</emphasis>
(Tshernovskij, 1949)
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFB8AAE0FFABB7C4C" author="Makarchenko, E. A." box="[1101,1290,1055,1077]" pageId="4" pageNumber="175" pagination="607 - 621" refId="ref3268" refString="Makarchenko, E. A. (2006) 3. Subfamily Diamesinae. In: Lelej, A. (Ed.), Key to the insects of Russian Far East. Vol. 6. Diptera and Siphonaptera. Pt 4. Dal'nauka, Vladivostok, pp. 253 - 276 + 468 - 480 + 607 - 621. [in Russian]" type="book chapter" year="2006">Makarchenko 2006</bibRefCitation>
, Fig. 181, 15)
</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50AE2DFA2D7C09" blockId="4.[151,1437,761,1576]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AE2DFA2D7C09" pageId="4" pageNumber="175">
Apex of the gonostylus broadly rounded................................................................................................
<emphasis id="A2A3EAE4FFC2FF9DFDECAE4AFD8B7C16" box="[555,570,1114,1135]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.
<taxonomicName id="57D74D75FFC2FF9DFD81AE4AFBE37C16" ID-CoL="75L2C" authority="Makarchenko, Kerkis et Ivanchenko, 1997" authorityName="Makarchenko, Kerkis et Ivanchenko" authorityYear="1997" box="[582,1106,1113,1135]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="altaica" subGenus="Pagastia">
(
<emphasis id="A2A3EAE4FFC2FF9DFD89AE4AFDEC7C16" box="[590,605,1114,1135]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFDB7AE4AFD027C16" box="[624,691,1114,1135]" italics="true" pageId="4" pageNumber="175">altaica</emphasis>
<bibRefCitation id="F4464B07FFC2FF9DFD7EAE49FBE37C16" author="Makarchenko, E. A. &amp; Kerkis, I. E. &amp; Ivanchenko, O. V." box="[697,1106,1113,1135]" pageId="4" pageNumber="175" pagination="1 - 8" refId="ref3398" refString="Makarchenko, E. A., Kerkis, I. E. &amp; Ivanchenko, O. V. (1997) Morphokaryological description of Pagastia altaica sp. n. (Diptera, Chironomidae) from Altai Mountains, with the key to Holarctic species of Pagastia Oliver. Far Eastern Entomologist, 43, 1 - 8." type="journal article" year="1997">Makarchenko, Kerkis et Ivanchenko, 1997</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFB98AE49FAE37C16" author="Makarchenko, E. A. &amp; Kerkis, I. E. &amp; Ivanchenko, O. V." box="[1119,1362,1113,1135]" pageId="4" pageNumber="175" pagination="1 - 8" refId="ref3398" refString="Makarchenko, E. A., Kerkis, I. E. &amp; Ivanchenko, O. V. (1997) Morphokaryological description of Pagastia altaica sp. n. (Diptera, Chironomidae) from Altai Mountains, with the key to Holarctic species of Pagastia Oliver. Far Eastern Entomologist, 43, 1 - 8." type="journal article" year="1997">
Makarchenko
<emphasis id="A2A3EAE4FFC2FF9DFB2DAE4AFAAA7C16" box="[1258,1307,1114,1135]" italics="true" pageId="4" pageNumber="175">et al.</emphasis>
1997
</bibRefCitation>
,
<figureCitation id="08EC2A73FFC2FF9DFA9BAE49FA227C09" box="[1372,1427,1113,1136]" captionStart="FIGURES 13" captionStartId="1.[151,264,1325,1350]" captionTargetBox="[178,1421,352,1271]" captionTargetId="figure@1.[151,1435,342,1297]" captionTargetPageId="1" captionText="FIGURES 13. Pagastia (P.) donoliveri sp. nov., male. 1, head in frontal view; 2, pronotum and mesonotum in dorsal view; 3, part of thorax in lateral view." figureDoi="http://doi.org/10.5281/zenodo.3724321" httpUri="https://zenodo.org/record/3724321/files/figure.png" pageId="4" pageNumber="175">Fig. 1</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B232E53FFC2FF9DFF50AE67FA2D7CBE" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50AE67FA2D7CD3" blockId="4.[151,1437,761,1576]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AE67FA2D7CD3" pageId="4" pageNumber="175">
6. Anal point absent. Alula as well as M
<subScript id="0C5334B3FFC2FF9DFD8DAE95FDD17CE8" attach="right" box="[586,608,1157,1169]" fontSize="5" pageId="4" pageNumber="175">1+2</subScript>
, M
<subScript id="0C5334B3FFC2FF9DFD46AE95FD277CE8" attach="left" box="[641,662,1157,1169]" fontSize="5" pageId="4" pageNumber="175">3+4</subScript>
, and Cu
<subScript id="0C5334B3FFC2FF9DFD2FAE95FD5E7CE8" attach="left" box="[744,751,1157,1169]" fontSize="5" pageId="4" pageNumber="175">1</subScript>
of wing with setae................................................................................ ..
<emphasis id="A2A3EAE4FFC2FF9DFD15AE85FD507CD3" box="[722,737,1173,1194]" italics="true" pageId="4" pageNumber="175">P</emphasis>
. (
<emphasis id="A2A3EAE4FFC2FF9DFD32AE85FCB57CD3" box="[757,772,1173,1194]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFCDFAE84FC247CD0" box="[792,917,1172,1193]" italics="true" pageId="4" pageNumber="175">subletteorum</emphasis>
Makarchenko (
<bibRefCitation id="F4464B07FFC2FF9DFBECAE84FB587CD0" author="Makarchenko, E. A." box="[1067,1257,1172,1194]" pageId="4" pageNumber="175" pagination="115 - 128" refId="ref3335" refString="Makarchenko, E. A. (2019) Review of the genus Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from North America, with description of P. (P.) subletteorum sp. nov. Zootaxa, 4664 (1), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4664.1.5" type="journal article" year="2019">Makarchenko 2019</bibRefCitation>
,
<figureCitation id="08EC2A73FFC2FF9DFB34AE84FAFC7CD3" box="[1267,1357,1172,1194]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 45" captionStart-2="FIGURES 69" captionStartId-0="1.[151,264,1325,1350]" captionStartId-1="2.[151,264,1863,1888]" captionStartId-2="3.[151,264,1476,1501]" captionTargetBox-0="[178,1421,352,1271]" captionTargetBox-1="[164,1421,193,1825]" captionTargetBox-2="[154,1435,183,1445]" captionTargetId-0="figure@1.[151,1435,342,1297]" captionTargetId-1="figure@2.[151,1436,179,1836]" captionTargetId-2="figure@3.[151,1435,179,1448]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 13. Pagastia (P.) donoliveri sp. nov., male. 1, head in frontal view; 2, pronotum and mesonotum in dorsal view; 3, part of thorax in lateral view." captionText-1="FIGURES 45. Pagastia (P.) donoliveri sp. nov., male. 4, hypopygium in dorsal view; 5, lateral aedeagal lobes. Scale bars: 50 µm." captionText-2="FIGURES 69. Pagastia (P.) donoliveri sp. nov., male. 67, parts of hypopygium in dorsal view; 8, apex of anal point with peg; 9, gonostylus." figureDoi-0="http://doi.org/10.5281/zenodo.3724321" figureDoi-1="http://doi.org/10.5281/zenodo.3724323" figureDoi-2="http://doi.org/10.5281/zenodo.3724325" httpUri-0="https://zenodo.org/record/3724321/files/figure.png" httpUri-1="https://zenodo.org/record/3724323/files/figure.png" httpUri-2="https://zenodo.org/record/3724325/files/figure.png" pageId="4" pageNumber="175">Figs. 16</figureCitation>
, 1920)
</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50AEA2FA2D7CBE" blockId="4.[151,1437,761,1576]" box="[151,1436,1201,1228]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AEA2FA2D7CBE" box="[151,1436,1201,1228]" pageId="4" pageNumber="175">
Anal point present. Alula and M
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, M
<subScript id="0C5334B3FFC2FF9DFD89AED0FDD57CB5" attach="left" box="[590,612,1216,1228]" fontSize="5" pageId="4" pageNumber="175">3+4</subScript>
, Cu
<subScript id="0C5334B3FFC2FF9DFD4BAED0FD227CB5" attach="left" box="[652,659,1216,1228]" fontSize="5" pageId="4" pageNumber="175">1</subScript>
without setae.................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B232E53FFC2FF9DFF50AEDFFA2D7D78" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50AEDFFA2D7C9D" blockId="4.[151,1437,761,1576]" box="[151,1436,1231,1253]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AEDFFA2D7C9D" box="[151,1436,1231,1253]" pageId="4" pageNumber="175">7. Anal point almost parallel-sided in dorsal view; lateral aedeagal lobe tapering to thin apex........................... 8</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50AEFDFA2D7D78" blockId="4.[151,1437,761,1576]" box="[151,1436,1260,1282]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AEFDFA2D7D78" box="[151,1436,1260,1282]" pageId="4" pageNumber="175"> Anal point tapering to apex; lateral aedeagal lobe broad in distal part........................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B232E53FFC2FF9DFF50AF1AFA2D7D45" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50AF1AFA2D7D59" blockId="4.[151,1437,761,1576]" box="[151,1436,1289,1312]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AF1AFA2D7D59" box="[151,1436,1289,1312]" pageId="4" pageNumber="175">
8. Anal point with rounded apex and without apical peg..
<emphasis id="A2A3EAE4FFC2FF9DFCCEAF1AFCA97D66" box="[777,792,1290,1311]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.
<taxonomicName id="57D74D75FFC2FF9DFCE4AF1AFB867D66" ID-CoL="6TKP4" authority="Oliver, 1959" authorityName="Oliver" authorityYear="1959" box="[803,1079,1289,1311]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="orthogonia" subGenus="Pagastia">
(
<emphasis id="A2A3EAE4FFC2FF9DFCECAF1AFC8B7D66" box="[811,826,1290,1311]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFC89AF1AFC087D66" box="[846,953,1290,1311]" italics="true" pageId="4" pageNumber="175">orthogonia</emphasis>
<bibRefCitation id="F4464B07FFC2FF9DFC78AF19FB867D66" author="Oliver, D. R." box="[959,1079,1289,1311]" pageId="4" pageNumber="175" pagination="48 - 64" refId="ref3459" refString="Oliver, D. R. (1959) Some Diamesini (Chironomidae) from the Nearctic and Palaearctic. Entomologisk Tidskrift, 80, 48 - 64." type="journal article" year="1959">Oliver, 1959</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFB84AF19FAB07D66" author="Makarchenko, E. A." box="[1091,1281,1289,1311]" pageId="4" pageNumber="175" pagination="115 - 128" refId="ref3335" refString="Makarchenko, E. A. (2019) Review of the genus Pagastia Oliver (Diptera: Chironomidae: Diamesinae) from North America, with description of P. (P.) subletteorum sp. nov. Zootaxa, 4664 (1), 115 - 128. https: // doi. org / 10.11646 / zootaxa. 4664.1.5" type="journal article" year="2019">Makarchenko 2019</bibRefCitation>
,
<figureCitation id="08EC2A73FFC2FF9DFACCAF19FAFD7D66" box="[1291,1356,1289,1311]" captionStart="FIGURES 69" captionStartId="3.[151,264,1476,1501]" captionTargetBox="[154,1435,183,1445]" captionTargetId="figure@3.[151,1435,179,1448]" captionTargetPageId="3" captionText="FIGURES 69. Pagastia (P.) donoliveri sp. nov., male. 67, parts of hypopygium in dorsal view; 8, apex of anal point with peg; 9, gonostylus." figureDoi="http://doi.org/10.5281/zenodo.3724325" httpUri="https://zenodo.org/record/3724325/files/figure.png" pageId="4" pageNumber="175">Figs. 7</figureCitation>
13, 21)
</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50AF37FA2D7D45" blockId="4.[151,1437,761,1576]" box="[151,1436,1319,1341]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AF37FA2D7D45" box="[151,1436,1319,1341]" pageId="4" pageNumber="175"> Anal point with pointed apical peg........................................................................ 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B232E53FFC2FF9DFF50AF54FA2D7DCB" pageId="4" pageNumber="175">
<paragraph id="906836F6FFC2FF9DFF50AF54FA2C7D0E" blockId="4.[151,1437,761,1576]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AF54FA2C7D0E" pageId="4" pageNumber="175">
9. Gonostylus with small basal lobe in outer side; apically rounded, without “heel”. AR 1.561.75.............................................................................
<emphasis id="A2A3EAE4FFC2FF9DFCABAF72FCCE7D0E" box="[876,895,1378,1399]" italics="true" pageId="4" pageNumber="175">P.</emphasis>
<taxonomicName id="57D74D75FFC2FF9DFC43AF72FB097D0E" ID-CoL="6TKP2" authority="Endo, 2004" authorityName="Endo" authorityYear="2004" box="[900,1208,1377,1399]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="hidakamontana" subGenus="Pagastia">
(
<emphasis id="A2A3EAE4FFC2FF9DFC4BAF72FC2A7D0E" box="[908,923,1378,1399]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFC69AF72FBF27D0E" box="[942,1091,1378,1399]" italics="true" pageId="4" pageNumber="175">hidakamontana</emphasis>
<bibRefCitation id="F4464B07FFC2FF9DFB8DAF71FB097D0E" author="Endo, K." box="[1098,1208,1377,1399]" pageId="4" pageNumber="175" pagination="277 - 289" refId="ref3123" refString="Endo, K. (2004) Genus Pagastia Oliver (Diptera: Chironomidae) from Japan, with description of a new species. Entomological Science, 7, 277 - 289. https: // doi. org / 10.1111 / j. 1479 - 8298.2004.00074. x" type="journal article" year="2004">Endo, 2004</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFB02AF71FA807D0E" author="Endo, K." box="[1221,1329,1377,1399]" pageId="4" pageNumber="175" pagination="277 - 289" refId="ref3123" refString="Endo, K. (2004) Genus Pagastia Oliver (Diptera: Chironomidae) from Japan, with description of a new species. Entomological Science, 7, 277 - 289. https: // doi. org / 10.1111 / j. 1479 - 8298.2004.00074. x" type="journal article" year="2004">Endo 2004</bibRefCitation>
,
<figureCitation id="08EC2A73FFC2FF9DFAFCAF71FA247D0E" box="[1339,1429,1377,1399]" captionStart-0="FIGURES 13" captionStart-1="FIGURES 45" captionStart-2="FIGURES 69" captionStartId-0="1.[151,264,1325,1350]" captionStartId-1="2.[151,264,1863,1888]" captionStartId-2="3.[151,264,1476,1501]" captionTargetBox-0="[178,1421,352,1271]" captionTargetBox-1="[164,1421,193,1825]" captionTargetBox-2="[154,1435,183,1445]" captionTargetId-0="figure@1.[151,1435,342,1297]" captionTargetId-1="figure@2.[151,1436,179,1836]" captionTargetId-2="figure@3.[151,1435,179,1448]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURES 13. Pagastia (P.) donoliveri sp. nov., male. 1, head in frontal view; 2, pronotum and mesonotum in dorsal view; 3, part of thorax in lateral view." captionText-1="FIGURES 45. Pagastia (P.) donoliveri sp. nov., male. 4, hypopygium in dorsal view; 5, lateral aedeagal lobes. Scale bars: 50 µm." captionText-2="FIGURES 69. Pagastia (P.) donoliveri sp. nov., male. 67, parts of hypopygium in dorsal view; 8, apex of anal point with peg; 9, gonostylus." figureDoi-0="http://doi.org/10.5281/zenodo.3724321" figureDoi-1="http://doi.org/10.5281/zenodo.3724323" figureDoi-2="http://doi.org/10.5281/zenodo.3724325" httpUri-0="https://zenodo.org/record/3724321/files/figure.png" httpUri-1="https://zenodo.org/record/3724323/files/figure.png" httpUri-2="https://zenodo.org/record/3724325/files/figure.png" pageId="4" pageNumber="175">Figs. 16</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50AF6FFA2D7DCB" blockId="4.[151,1437,761,1576]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AF6FFA2D7DCB" pageId="4" pageNumber="175">
Gonostylus with rounded basal lobe in outer side and with lobe like “heel” apically. AR 1.221.26........................................................................................
<emphasis id="A2A3EAE4FFC2FF9DFBC2AF8DFBA97DCB" box="[1029,1048,1437,1458]" italics="true" pageId="4" pageNumber="175">P.</emphasis>
<taxonomicName id="57D74D75FFC2FF9DFBDAAF8CFB1A7DC8" ID-CoL="9J242" authority="Makarchenko &amp; Hansen, 2020" authorityName="Makarchenko &amp; Hansen" authorityYear="2020" box="[1053,1195,1436,1458]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="donoliveri" status="sp. nov." subGenus="Pagastia">
(
<emphasis id="A2A3EAE4FFC2FF9DFBE2AF8DFB857DCB" box="[1061,1076,1437,1458]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFB8FAF8CFB1A7DC8" box="[1096,1195,1436,1457]" italics="true" pageId="4" pageNumber="175">donoliveri</emphasis>
</taxonomicName>
<emphasis id="A2A3EAE4FFC2FF9DFB77AF8CFB4C7DC8" bold="true" box="[1200,1277,1436,1457]" pageId="4" pageNumber="175">
<taxonomicNameLabel id="B990579FFFC2FF9DFB77AF8CFB4C7DC8" box="[1200,1277,1436,1457]" pageId="4" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="08EC2A73FFC2FF9DFACDAF8CFAFF7DC8" box="[1290,1358,1436,1458]" captionStart="FIGURES 45" captionStartId="2.[151,264,1863,1888]" captionTargetBox="[164,1421,193,1825]" captionTargetId="figure@2.[151,1436,179,1836]" captionTargetPageId="2" captionText="FIGURES 45. Pagastia (P.) donoliveri sp. nov., male. 4, hypopygium in dorsal view; 5, lateral aedeagal lobes. Scale bars: 50 µm." figureDoi="http://doi.org/10.5281/zenodo.3724323" httpUri="https://zenodo.org/record/3724323/files/figure.png" pageId="4" pageNumber="175">Figs. 4</figureCitation>
,
<figureCitation id="08EC2A73FFC2FF9DFA9FAF8CFA247DCB" box="[1368,1429,1436,1458]" captionStart="FIGURES 69" captionStartId="3.[151,264,1476,1501]" captionTargetBox="[154,1435,183,1445]" captionTargetId="figure@3.[151,1435,179,1448]" captionTargetPageId="3" captionText="FIGURES 69. Pagastia (P.) donoliveri sp. nov., male. 67, parts of hypopygium in dorsal view; 8, apex of anal point with peg; 9, gonostylus." figureDoi="http://doi.org/10.5281/zenodo.3724325" httpUri="https://zenodo.org/record/3724325/files/figure.png" pageId="4" pageNumber="175">6, 7, 9</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
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<keyLead id="2B2695C3FFC2FF9DFF50AFAAFA2D7D94" pageId="4" pageNumber="175">
10. Anal point narrow, tapering to pointed apex, without apical peg; gonostylus in basal part with outer angle-shaped projection. AR 2.182.42.................
<emphasis id="A2A3EAE4FFC2FF9DFDFEAFC8FDFD7D94" box="[569,588,1496,1517]" italics="true" pageId="4" pageNumber="175">P.</emphasis>
<taxonomicName id="57D74D75FFC2FF9DFD96AFC7FB9D7D95" ID-CoL="9J2T8" authority="Makarchenko et Wang, 2017" authorityName="Makarchenko et Wang" authorityYear="2017" box="[593,1068,1495,1517]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="tianmumontana" subGenus="Pagastia">
(
<emphasis id="A2A3EAE4FFC2FF9DFD9EAFC8FDD97D94" box="[601,616,1496,1517]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFDBCAFC8FCA37D94" box="[635,786,1496,1517]" italics="true" pageId="4" pageNumber="175">tianmumontana</emphasis>
<bibRefCitation id="F4464B07FFC2FF9DFCDFAFC7FB9D7D95" author="Makarchenko, E. A. &amp; Wang, X." box="[792,1068,1495,1517]" pageId="4" pageNumber="175" pagination="13 - 15" refId="ref3221" refString="Makarchenko, E. A. &amp; Wang, X. (2017) Pagastia tianmumontana sp. n. - a new species of chironomids (Diptera: Chironomidae: Diamesinae) from South China. Far Eastern Entomologist, 336, 13 - 15." type="journal article" year="2017">
Makarchenko
<emphasis id="A2A3EAE4FFC2FF9DFC65AFC8FC027D94" box="[930,947,1496,1517]" italics="true" pageId="4" pageNumber="175">et</emphasis>
Wang, 2017
</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFBFEAFC7FAF77D95" author="Makarchenko, E. A. &amp; Wang, X." box="[1081,1350,1495,1517]" pageId="4" pageNumber="175" pagination="13 - 15" refId="ref3221" refString="Makarchenko, E. A. &amp; Wang, X. (2017) Pagastia tianmumontana sp. n. - a new species of chironomids (Diptera: Chironomidae: Diamesinae) from South China. Far Eastern Entomologist, 336, 13 - 15." type="journal article" year="2017">Makarchenko &amp;Wang 2017</bibRefCitation>
, Fig. 17)
</keyLead>
</paragraph>
<paragraph id="906836F6FFC2FF9DFF50AFE5FA2D7E51" blockId="4.[151,1437,761,1576]" pageId="4" pageNumber="175">
<keyLead id="2B2695C3FFC2FF9DFF50AFE5FA2D7E51" pageId="4" pageNumber="175">
Anal point widest in basal part and thin apically, pointed and sometimes with apical peg; gonostylus without outer basal projection and with rounded apex. AR 1.812.10........
<emphasis id="A2A3EAE4FFC2FF9DFD22AC02FD457E5E" box="[741,756,1554,1575]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.
<taxonomicName id="57D74D75FFC2FF9DFCC7AC02FBF07E5E" ID-CoL="75KP2" authority="(Tokunaga, 1936)" baseAuthorityName="Tokunaga" baseAuthorityYear="1936" box="[768,1089,1553,1575]" class="Insecta" family="Chironomidae" genus="Pagastia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="lanceolata" subGenus="Pagastia">
(
<emphasis id="A2A3EAE4FFC2FF9DFCCFAC02FCA67E5E" box="[776,791,1554,1575]" italics="true" pageId="4" pageNumber="175">P</emphasis>
.)
<emphasis id="A2A3EAE4FFC2FF9DFCECAC02FC217E5E" box="[811,912,1554,1575]" italics="true" pageId="4" pageNumber="175">lanceolata</emphasis>
(Tokunaga, 1936)
</taxonomicName>
(
<bibRefCitation id="F4464B07FFC2FF9DFB8AAC01FABB7E5E" author="Makarchenko, E. A." box="[1101,1290,1553,1575]" pageId="4" pageNumber="175" pagination="607 - 621" refId="ref3268" refString="Makarchenko, E. A. (2006) 3. Subfamily Diamesinae. In: Lelej, A. (Ed.), Key to the insects of Russian Far East. Vol. 6. Diptera and Siphonaptera. Pt 4. Dal'nauka, Vladivostok, pp. 253 - 276 + 468 - 480 + 607 - 621. [in Russian]" type="book chapter" year="2006">Makarchenko 2006</bibRefCitation>
, Fig. 180, 14)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>