233 lines
32 KiB
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233 lines
32 KiB
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<document id="ADC9EC0653B593DD560B7BCEA0F0FF72" ID-DOI="10.37828/em.2020.30.8" ID-ISSN="2336-9744" ID-Zenodo-Dep="13231368" ID-ZooBank="urn:lsid:zoobank.org:pub:789931C7-5667-4258-9986-121A74B53B9B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722912696433" checkinUser="felipe" docAuthor="László, Gyula M." docDate="2020" docId="183FDF25FFD2A56AFF6DF91CFB2FFECD" docLanguage="en" docName="EcolMontenegrina.30.82-92.pdf" docOrigin="Ecologica Montenegrina 30" docSource="http://dx.doi.org/10.37828/em.2020.30.8" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Ezishnola carcassoni László, 2020, sp. nov." docType="treatment" docUuid="E242BC96-AB35-4468-A9D9-66D533D459AE" docUuidSource="ZooBank" docVersion="2" lastPageNumber="92" masterDocId="E406A75DFFDAA560FFFAFFEBFFDAFFA8" masterDocTitle="On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae)" masterLastPageNumber="92" masterPageNumber="82" pageNumber="90" updateTime="1722913214493" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="8907848DBE62FF52550EB27CAE2103A4">
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<mods:title id="017FFAD46514019F4D3874CDE6396FED">On the taxonomy of Meganola cretacea (Hampson, 1914) with description of two new species (Lepidoptera, Nolidae, Nolinae)</mods:title>
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<mods:name id="09842D1702B61AB71AAC1DA53ABB5F38" type="personal">
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<mods:namePart id="267D5CDD85CF820D1B2C22E9397DF22C">László, Gyula M.</mods:namePart>
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<mods:typeOfResource id="250AC5D86E5DE038FFAF1E9715B4B150">text</mods:typeOfResource>
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<mods:title id="477B0F6DDFC5B7A8EE20BFDB930D7CE5">Ecologica Montenegrina</mods:title>
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<mods:part id="900C8A06F9F34D95A13DED261EE689D1">
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<mods:date id="20148D175A626E548F7F28B2A87D9315">2020</mods:date>
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<mods:number id="806F62C644D709D8CB21B850E47F1192">2020-04-18</mods:number>
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<mods:detail id="804F3C4669EC50DC90C9917C2B7782A0" type="volume">
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<mods:number id="722F6522AD100B3D154FA436242A3C6D">30</mods:number>
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<mods:start id="38DC9A1D5C4156F843D31F971D480749">82</mods:start>
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<mods:url id="96D1CB922028FE71687D4B157A79CF31">http://dx.doi.org/10.37828/em.2020.30.8</mods:url>
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</mods:location>
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<mods:classification id="509DACFC38BB6CCA5E5FDDB3C0FF50CF">journal article</mods:classification>
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<mods:identifier id="0F772B0803EAC3141E32BB21C9C76CDE" type="DOI">10.37828/em.2020.30.8</mods:identifier>
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<mods:identifier id="C83781C7837A8AC0DCE7DE7C82F8E55F" type="ISSN">2336-9744</mods:identifier>
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<mods:identifier id="1DBA05B3A79D45F45E68AA5EC6A3D7BB" type="ZooBank">urn:lsid:zoobank.org:pub:789931C7-5667-4258-9986-121A74B53B9B</mods:identifier>
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<treatment id="183FDF25FFD2A56AFF6DF91CFB2FFECD" ID-DOI="http://doi.org/10.5281/zenodo.13231384" ID-Zenodo-Dep="13231384" LSID="urn:lsid:zoobank.org:act:E242BC96-AB35-4468-A9D9-66D533D459AE" httpUri="http://treatment.plazi.org/id/183FDF25FFD2A56AFF6DF91CFB2FFECD" lastPageId="10" lastPageNumber="92" pageId="8" pageNumber="90">
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<subSubSection id="D88C3DB8FFD2A568FF6DF91CFD2CF8B9" box="[151,758,1783,1809]" pageId="8" pageNumber="90" type="nomenclature">
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<paragraph id="90296E33FFD2A568FF6DF91CFD2CF8B9" blockId="8.[151,758,1783,1809]" box="[151,758,1783,1809]" pageId="8" pageNumber="90">
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<heading id="CB61D95FFFD2A568FF6DF91CFD2CF8B9" box="[151,758,1783,1809]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="90" reason="2">
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<emphasis id="A2E2B221FFD2A568FF6DF91CFD2CF8B9" bold="true" box="[151,758,1783,1809]" pageId="8" pageNumber="90">
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<taxonomicName id="579615B0FFD2A568FF6DF91CFE42F8B9" ID-CoL="B9MD4" authorityName="László" authorityYear="2020" box="[151,408,1783,1809]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="90" phylum="Arthropoda" rank="species" species="carcassoni" status="sp. nov.">
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<emphasis id="A2E2B221FFD2A568FF6DF91CFE42F8B9" bold="true" box="[151,408,1783,1809]" italics="true" pageId="8" pageNumber="90">Ezishnola carcassoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B9D10F5AFFD2A568FE5AF91CFE25F8B9" box="[416,511,1783,1809]" pageId="8" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="08AD72B6FFD2A568FDF4F91CFD4CF8B9" box="[526,662,1783,1809]" captionStart="Figures 9-14" captionStartId="5.[151,236,1461,1485]" captionTargetBox="[158,1420,202,1404]" captionTargetId="figure-16@5.[151,1432,189,1442]" captionTargetPageId="5" captionText="Figures 9-14: adults. 9, Ezishnola inopinata sp. n., holotype, female, Zambia (ANHRT); 10, E. inopinata sp. n., paratype, female, Zambia (ANHRT); 11, E. inopinata sp. n., paratype, male, Zambia (ANHRT); 12, E. inopinata sp. n., paratype, male, Zambia (ANHRT); 13, E. carcassoni sp. n., holotype, female, Uganda (NHMUK); 14, E. carcassoni sp. n., paratype, male, Uganda (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13231372" httpUri="https://zenodo.org/record/13231372/files/figure.png" pageId="8" pageNumber="90">Figs 13, 14</figureCitation>
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,
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<figureCitation id="08AD72B6FFD2A568FD59F91CFD18F8B9" box="[675,706,1783,1809]" captionStart="Figures 15-18" captionStartId="6.[151,236,1903,1927]" captionTargetBox="[293,1306,199,1864]" captionTargetId="figure-11@6.[198,1388,189,1884]" captionTargetPageId="6" captionText="Figures 15-18: female genitalia. 15, Ezishnola cretacea (Hampson, 1914), LGNA 990, Ghana, Bunso Arboretum (HNHM); 16, E. inopinata sp. n., holotype, LGNA 692, Zambia, Lukwakwa (ANHRT); 17, E. carcassoni sp. n. holotype, NHMUK 010316559 (Gy. M. László), Uganda, Kalinzu Forest (NHMUK); 18, E. fuscographa László, Ronkay & Witt, 2010, paratype, LGN 971, Thailand, Bo Luang (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.13231376" httpUri="https://zenodo.org/record/13231376/files/figure.png" pageId="8" pageNumber="90">17</figureCitation>
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,
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<figureCitation id="08AD72B6FFD2A568FD35F91CFD34F8B9" box="[719,750,1783,1809]" captionStart="Figures 19-24" captionStartId="7.[151,236,1880,1904]" captionTargetBox="[225,1347,214,1844]" captionTargetId="figure-16@7.[170,1416,189,1862]" captionTargetPageId="7" captionText="Figures 19-24: male genitalia. 19, Ezishnola cretacea (Hampson, 1914), LGNA 835, Guinea, Nimba Mts (ANHRT); 20, E. cretacea (Hampson, 1914), LGNA 953, Liberia, Nimba Mts (ANHRT); 21, E. inopinata sp. n., paratype, LGNA 60, Zambia, Lukwakwa (ANHRT); 22, E. inopinata sp. n., paratype, LGNA 292, Zambia, Ikelenge (ANHRT); 23, E. carcassoni sp. n. paratype, NHMUK 010316558 (Gy. M. László), Uganda, Masaka (NHMUK); 24, E. fuscographa László, Ronkay & Witt, 2010, holotype, LGN 953, Thailand, Pua (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.13231378" httpUri="https://zenodo.org/record/13231378/files/figure.png" pageId="8" pageNumber="90">23</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D88C3DB8FFD2A568FF6DF8D1FBBEF856" pageId="8" pageNumber="90" type="materials_examined">
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<paragraph id="90296E33FFD2A568FF6DF8D1FEFBF830" blockId="8.[151,1435,1850,1944]" pageId="8" pageNumber="90">
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<materialsCitation id="20FE646EFFD2A568FF6DF8D1FEFBF830" collectingDate="1961-11" collectionCode="NHMUK" collectorName="R. H. Carcasson" country="Uganda" location="Kalinzu Forest Ankole" municipality="Kalinzu Forest" pageId="8" pageNumber="90" specimenCode="NHMUK 014046240, NHMUK 010316559" specimenCount="1" typeStatus="holotype">
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<emphasis id="A2E2B221FFD2A568FF6DF8D1FECAF8FC" bold="true" box="[151,272,1850,1876]" pageId="8" pageNumber="90">
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<typeStatus id="4F2DD091FFD2A568FF6DF8D1FED1F8FC" box="[151,267,1850,1876]" pageId="8" pageNumber="90" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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female, “
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<location id="954938E8FFD2A568FE79F8D1FD49F8FD" LSID="urn:lsid:plazi:treatment:183FDF25FFD2A56AFF6DF91CFB2FFECD:954938E8FFD2A568FE79F8D1FD49F8FD" box="[387,659,1850,1877]" country="Uganda" municipality="Kalinzu Forest" name="Kalinzu Forest Ankole" pageId="8" pageNumber="90">
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<collectingMunicipality id="704DF449FFD2A568FE79F8D1FDEFF8FD" box="[387,565,1850,1877]" pageId="8" pageNumber="90">Kalinzu Forest</collectingMunicipality>
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Ankole
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</location>
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<collectingCountry id="E8812EA3FFD2A568FD66F8D1FD22F8FD" box="[668,760,1850,1877]" name="Uganda" pageId="8" pageNumber="90">Uganda</collectingCountry>
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<date id="E42848F3FFD2A568FCFAF8D0FCA5F8FD" box="[768,895,1850,1877]" pageId="8" pageNumber="90" value="1961-11">
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<collectingDate id="F46CB11BFFD2A568FCFAF8D0FCA5F8FD" box="[768,895,1850,1877]" pageId="8" pageNumber="90" value="1961-11">Nov. 1961</collectingDate>
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</date>
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<collectorName id="3D630BE5FFD2A568FC7DF8D0FB96F8FD" box="[903,1100,1850,1877]" pageId="8" pageNumber="90">R. H. Carcasson</collectorName>
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”, “Coryndon Museum B.M. 1963-322”, QR code label with unique number:
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<specimenCode id="C030C648FFD2A568FD07F8B7FBD8F8DF" box="[765,1026,1884,1911]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="90" type="Museum">NHMUK 014046240</specimenCode>
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, slide No.:
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<specimenCode id="C030C648FFD2A568FB61F8B7FA41F8DF" box="[1179,1435,1884,1911]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="90" type="Museum">NHMUK 010316559</specimenCode>
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(
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<collectionCode id="F687F6F6FFD2A568FF65F896FECFF830" box="[159,277,1917,1944]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="90" type="Museum">NHMUK</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="90296E33FFD2A568FF6DF82AFBBEF856" blockId="8.[151,1436,1985,2046]" pageId="8" pageNumber="90">
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<materialsCitation id="20FE646EFFD2A568FF6DF82AFD8FF856" collectingDate="1960-10" collectionCode="NHMUK" collectorName="R. H. Carcasson" country="Uganda" location="Katera Sango Bay" pageId="8" pageNumber="90" specimenCode="NHMUK 014046239" specimenCount="1" specimenCount-male="1" stateProvince="Masaka" typeStatus="paratype">
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<emphasis id="A2E2B221FFD2A568FF6DF82AFE5FF873" bold="true" box="[151,389,1985,2011]" pageId="8" pageNumber="90">
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<typeStatus id="4F2DD091FFD2A568FF6DF82AFED1F873" box="[151,267,1985,2011]" pageId="8" pageNumber="90" type="paratype">Paratype</typeStatus>
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.
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<collectingCountry id="E8812EA3FFD2A568FEE6F82AFE5AF873" box="[284,384,1985,2011]" name="Uganda" pageId="8" pageNumber="90">Uganda</collectingCountry>
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.
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</emphasis>
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<specimenCount id="8690A5BAFFD2A568FE6BF82AFE3DF874" box="[401,487,1985,2012]" count="1" pageId="8" pageNumber="90" type="male">1 male</specimenCount>
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,
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<location id="954938E8FFD2A568FE0CF829FD0CF874" LSID="urn:lsid:plazi:treatment:183FDF25FFD2A56AFF6DF91CFB2FFECD:954938E8FFD2A568FE0CF829FD0CF874" box="[502,726,1985,2012]" country="Uganda" name="Katera Sango Bay" pageId="8" pageNumber="90" stateProvince="Masaka">Katera Sango Bay</location>
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,
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<collectingRegion id="5252A0D1FFD2A568FD1CF82AFC9EF874" box="[742,836,1985,2012]" country="Uganda" name="Masaka" pageId="8" pageNumber="90">Masaka</collectingRegion>
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,
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<date id="E42848F3FFD2A568FCA9F82AFC15F874" box="[851,975,1985,2012]" pageId="8" pageNumber="90" value="1960-10">
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<collectingDate id="F46CB11BFFD2A568FCA9F82AFC15F874" box="[851,975,1985,2012]" pageId="8" pageNumber="90" value="1960-10">Oct. 1960</collectingDate>
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</date>
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,
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<collectorName id="3D630BE5FFD2A568FC24F829FB46F874" box="[990,1180,1985,2012]" pageId="8" pageNumber="90">R.H. Carcasson</collectorName>
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, QR code label with unique number:
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<specimenCode id="C030C648FFD2A568FEADF808FD8FF856" box="[343,597,2019,2046]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="90" type="Museum">NHMUK 014046239</specimenCode>
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</materialsCitation>
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,
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<materialsCitation id="20FE646EFFD2A568FD9AF808FBBAF856" box="[608,1120,2019,2046]" collectionCode="NHMUK" country="Uganda" pageId="8" pageNumber="90" specimenCode="NHMUK 010316558" specimenCount="1" typeStatus="paratype">
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slide No.:
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<specimenCode id="C030C648FFD2A568FD23F808FC09F856" box="[729,979,2019,2046]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="90" type="Museum">NHMUK 010316558</specimenCode>
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(
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<collectionCode id="F687F6F6FFD2A568FC18F808FB82F856" box="[994,1112,2019,2046]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="90" type="Museum">NHMUK</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="D88C3DB8FFD3A569FF6DFF2FFD5EFD1F" pageId="9" pageNumber="91" type="diagnosis">
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<paragraph id="90296E33FFD3A569FF6DFF2FFD5EFD1F" blockId="9.[151,1437,195,695]" pageId="9" pageNumber="91">
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<emphasis id="A2E2B221FFD3A569FF6DFF2FFECDFF76" bold="true" box="[151,279,196,222]" pageId="9" pageNumber="91">Diagnosis.</emphasis>
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<taxonomicName id="579615B0FFD3A569FEDAFF2FFE18FF75" authorityName="László" authorityYear="2020" box="[288,450,196,221]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="carcassoni" status="sp. n.">
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<emphasis id="A2E2B221FFD3A569FEDAFF2FFE18FF75" box="[288,450,196,221]" italics="true" pageId="9" pageNumber="91">E. carcassoni</emphasis>
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</taxonomicName>
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<emphasis id="A2E2B221FFD3A569FE31FF2FFDD5FF76" bold="true" box="[459,527,196,222]" pageId="9" pageNumber="91">
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<taxonomicNameLabel id="B9D10F5AFFD3A569FE31FF2FFDD5FF76" box="[459,527,196,222]" pageId="9" pageNumber="91" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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is an East African allopatric sister species of
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<taxonomicName id="579615B0FFD3A569FBCDFF2FFB16FF75" baseAuthorityName="Hampson" baseAuthorityYear="1914" box="[1079,1228,196,221]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="creatacea">
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<emphasis id="A2E2B221FFD3A569FBCDFF2FFB16FF75" box="[1079,1228,196,221]" italics="true" pageId="9" pageNumber="91">E. creatacea</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="579615B0FFD3A569FAF3FF2FFA46FF75" authorityName="László" authorityYear="2020" box="[1289,1436,196,221]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="inopinata">
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<emphasis id="A2E2B221FFD3A569FAF3FF2FFA46FF75" box="[1289,1436,196,221]" italics="true" pageId="9" pageNumber="91">E. inopinata</emphasis>
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</taxonomicName>
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known so far only from
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<collectingCountry id="E8812EA3FFD3A569FE1BFF0EFD9AFEA8" box="[481,576,229,256]" name="Uganda" pageId="9" pageNumber="91">Uganda</collectingCountry>
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. The new species differ externally from both sister species by its conspicuously interrupted transverse lines consisting of a dense row of sharply defined blackish dots, whereas the crosslines are more continuous in the other species, although similar wing pattern is not unknown in
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<taxonomicName id="579615B0FFD3A569FED0FEA0FE6BFECC" authorityName="László" authorityYear="2020" baseAuthorityName="Hampson" baseAuthorityYear="1914" box="[298,433,331,356]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="cretacea">
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<emphasis id="A2E2B221FFD3A569FED0FEA0FE6BFECC" box="[298,433,331,356]" italics="true" pageId="9" pageNumber="91">E. cretacea</emphasis>
|
|
</taxonomicName>
|
|
. In the male genitalia,
|
|
<taxonomicName id="579615B0FFD3A569FD3BFEA0FCBBFECC" authorityName="László" authorityYear="2020" box="[705,865,331,356]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="carcassoni">
|
|
<emphasis id="A2E2B221FFD3A569FD3BFEA0FCBBFECC" box="[705,865,331,356]" italics="true" pageId="9" pageNumber="91">E. carcassoni</emphasis>
|
|
</taxonomicName>
|
|
differs from both sister species by its somewhat shorter uncus and valva, in addition more robust, almost straight harpe (those of the other two species are thinner and slightly arcuate) and considerably narrower, more elongate transtillae. In the female genitalia, the new species has somewhat shorter apophyses than those of
|
|
<taxonomicName id="579615B0FFD3A569FCA0FE5AFC3AFE62" authorityName="László" authorityYear="2020" baseAuthorityName="Hampson" baseAuthorityYear="1914" box="[858,992,433,458]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="cretacea">
|
|
<emphasis id="A2E2B221FFD3A569FCA0FE5AFC3AFE62" box="[858,992,433,458]" italics="true" pageId="9" pageNumber="91">E. cretacea</emphasis>
|
|
</taxonomicName>
|
|
, ovoid corpus bursae without tubular section and without sack-like cervix bursae (corpus bursae with conspicuously long tubular section and cervix bursae is sack-like in
|
|
<taxonomicName id="579615B0FFD3A569FE02FE1FFD5BFDA5" authorityName="László" authorityYear="2020" baseAuthorityName="Hampson" baseAuthorityYear="1914" box="[504,641,500,525]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="cretacea">
|
|
<emphasis id="A2E2B221FFD3A569FE02FE1FFD5BFDA5" box="[504,641,500,525]" italics="true" pageId="9" pageNumber="91">E. cretacea</emphasis>
|
|
</taxonomicName>
|
|
), in addition
|
|
<taxonomicName id="579615B0FFD3A569FCD1FE1FFC15FDA5" authorityName="László" authorityYear="2020" box="[811,975,500,525]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="carcassoni">
|
|
<emphasis id="A2E2B221FFD3A569FCD1FE1FFC15FDA5" box="[811,975,500,525]" italics="true" pageId="9" pageNumber="91">E. carcassoni</emphasis>
|
|
</taxonomicName>
|
|
lacks signum bursae. The differences between the female genitalia of the new species and
|
|
<taxonomicName id="579615B0FFD3A569FCF8FDFDFC4EFD87" authorityName="László" authorityYear="2020" box="[770,916,534,559]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="inopinata">
|
|
<emphasis id="A2E2B221FFD3A569FCF8FDFDFC4EFD87" box="[770,916,534,559]" italics="true" pageId="9" pageNumber="91">E. inopinata</emphasis>
|
|
</taxonomicName>
|
|
are as follows:
|
|
<taxonomicName id="579615B0FFD3A569FBB4FDFDFB35FD87" authorityName="László" authorityYear="2020" box="[1102,1263,534,559]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="carcassoni">
|
|
<emphasis id="A2E2B221FFD3A569FBB4FDFDFB35FD87" box="[1102,1263,534,559]" italics="true" pageId="9" pageNumber="91">E. carcassoni</emphasis>
|
|
</taxonomicName>
|
|
has somewhat longer apophyses, narrower ductus bursae with longer sclerotized medial section compared to those of
|
|
<taxonomicName id="579615B0FFD3A569FA7EFDD3FED2FDDA" authorityName="László" authorityYear="2020" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="inopinata">
|
|
<emphasis id="A2E2B221FFD3A569FA7EFDD3FED2FDDA" italics="true" pageId="9" pageNumber="91">E. inopinata</emphasis>
|
|
</taxonomicName>
|
|
, in addition less elongate, more rounded corpus bursae with simple cervix bursae, while the corpus bursae of
|
|
<taxonomicName id="579615B0FFD3A569FEEAFD90FE7EFD3C" authorityName="László" authorityYear="2020" box="[272,420,635,660]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="91" phylum="Arthropoda" rank="species" species="inopinata">
|
|
<emphasis id="A2E2B221FFD3A569FEEAFD90FE7EFD3C" box="[272,420,635,660]" italics="true" pageId="9" pageNumber="91">E. inopinata</emphasis>
|
|
</taxonomicName>
|
|
is more elongated with anteriorly dilated distal tubular section and its cervix bursae with a well-developed, membranous sack.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D88C3DB8FFD3A569FF6DFD0BFD00F854" pageId="9" pageNumber="91" type="description">
|
|
<paragraph id="90296E33FFD3A569FF6DFD0BFB93FCB4" blockId="9.[151,1436,735,1471]" pageId="9" pageNumber="91">
|
|
<emphasis id="A2E2B221FFD3A569FF6DFD0BFEF2FD52" bold="true" box="[151,296,736,762]" pageId="9" pageNumber="91">Description</emphasis>
|
|
(
|
|
<figureCitation id="08AD72B6FFD3A569FEC2FD34FE1AFD52" box="[312,448,735,762]" captionStart="Figures 9-14" captionStartId="5.[151,236,1461,1485]" captionTargetBox="[158,1420,202,1404]" captionTargetId="figure-16@5.[151,1432,189,1442]" captionTargetPageId="5" captionText="Figures 9-14: adults. 9, Ezishnola inopinata sp. n., holotype, female, Zambia (ANHRT); 10, E. inopinata sp. n., paratype, female, Zambia (ANHRT); 11, E. inopinata sp. n., paratype, male, Zambia (ANHRT); 12, E. inopinata sp. n., paratype, male, Zambia (ANHRT); 13, E. carcassoni sp. n., holotype, female, Uganda (NHMUK); 14, E. carcassoni sp. n., paratype, male, Uganda (NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13231372" httpUri="https://zenodo.org/record/13231372/files/figure.png" pageId="9" pageNumber="91">Figs 13, 14</figureCitation>
|
|
). Forewing length
|
|
<quantity id="576EC3D6FFD3A569FD67FD34FD2BFD52" box="[669,753,735,762]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="91" unit="mm" value="12.0">12 mm</quantity>
|
|
in male,
|
|
<quantity id="576EC3D6FFD3A569FCA7FD34FC6BFD52" box="[861,945,735,762]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="91" unit="mm" value="13.0">13 mm</quantity>
|
|
in female. Antenna filiform in both sexes with somewhat longer and denser ciliation in males compared to that of females.
|
|
</paragraph>
|
|
<paragraph id="90296E33FFD3A569FF0FFCC8FEB4FC4E" blockId="9.[151,1436,735,1471]" pageId="9" pageNumber="91">Head relatively large, labial palps medium long, curved dorsad, inner surface brownish white, lateral and ventral surface of first and second segments brownish grey, third segment pale creamy white; frons and vertex pale brownish white; compound eyes moderately large, globular. Thorax brownish grey, basal twothird of collar darker greyish, abdomen brownish white. Intraspecific variability limited. Sexual dimorphism is moderate, expressed in size (female slightly larger) and in coloration, (female hindwing somewhat darker than that of male).</paragraph>
|
|
<paragraph id="90296E33FFD3A569FF0FFC06FCC3FA90" blockId="9.[151,1436,735,1471]" pageId="9" pageNumber="91">Forewing relatively long and broad, rather quadrangular, apically rounded. Forewing ground colour pale brownish grey, with some darker brown irregular patches in the postmedian area. Sub-basal and basal lines diffuse, shadow-like, represented by groups of brownish hair scales; antemedial line fine, brownish grey, interrupted, almost straight, running angularly towards costa; medial line rather thick, interrupted, diffuse, present only in the ventral third; Orbicular stigma rather large, rounded, consisting of raised pale whitish grey scales. Postmedial line relatively thick, interrupted, very sharply defined by dense row of dark brown scales, upper section broadly arched, lower section running parallel with medial line. Subterminal line rather undulating, interrupted, consisting of dark brown triangular patches of different size; terminal line thick, interrupted, consisting of a row of dark brown streaks; terminal area with some red-brown suffusion; cilia pale brownish grey chequered with darker brown.</paragraph>
|
|
<paragraph id="90296E33FFD3A569FF0FFAD4FCB5FA17" blockId="9.[151,1436,735,1471]" pageId="9" pageNumber="91">Hindwing pale greyish white, with some darker brownish suffusion terminally in males, darker grey in females, turning to brownish grey in the terminal area; cilia whitish grey in both sexes. Underside of forewing uniformly dark brownish grey, that of hindwing paler brownish grey, darkened along the margin in males, similar but darker in females, without traces of pattern.</paragraph>
|
|
<paragraph id="90296E33FFD3A569FF6DFA03FC2CF8B8" blockId="9.[151,1436,1511,1808]" pageId="9" pageNumber="91">
|
|
<emphasis id="A2E2B221FFD3A569FF6DFA03FE96F9AA" bold="true" box="[151,332,1512,1538]" pageId="9" pageNumber="91">Male genitalia</emphasis>
|
|
(
|
|
<figureCitation id="08AD72B6FFD3A569FE9AFA0CFE61F9AA" box="[352,443,1511,1538]" captionStart="Figures 19-24" captionStartId="7.[151,236,1880,1904]" captionTargetBox="[225,1347,214,1844]" captionTargetId="figure-16@7.[170,1416,189,1862]" captionTargetPageId="7" captionText="Figures 19-24: male genitalia. 19, Ezishnola cretacea (Hampson, 1914), LGNA 835, Guinea, Nimba Mts (ANHRT); 20, E. cretacea (Hampson, 1914), LGNA 953, Liberia, Nimba Mts (ANHRT); 21, E. inopinata sp. n., paratype, LGNA 60, Zambia, Lukwakwa (ANHRT); 22, E. inopinata sp. n., paratype, LGNA 292, Zambia, Ikelenge (ANHRT); 23, E. carcassoni sp. n. paratype, NHMUK 010316558 (Gy. M. László), Uganda, Masaka (NHMUK); 24, E. fuscographa László, Ronkay & Witt, 2010, holotype, LGN 953, Thailand, Pua (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.13231378" httpUri="https://zenodo.org/record/13231378/files/figure.png" pageId="9" pageNumber="91">Fig. 23</figureCitation>
|
|
). Uncus relatively short, rather narrow, gradually tapered distally, apically with a short, fine, pointed hook; subscaphium rather long and narrow poorly sclerotized; tegumen short and broad, rather triangular; valvae weakly sclerotized, relatively short, medially moderately dilated, apically broadly rounded, tongue-shaped; harpe thorn-like, rather long and robust, almost straight, apically curved and pointed, situated in the central plate of valva; transtillae well-developed, elongate-ovoid, medially divided; sacculus long and broad, basal third more strongly sclerotized than apical one; fultura inferior (juxta) very small, with a pair of ear-like rounded lobes; vinculum rather broad and long, broad V-shaped, saccus short, apically acutely pointed. Aedeagus short and narrow, caecum penis rather long, evenly rounded; carina without sclerotization; vesica membranous in full length, without cornuti.
|
|
</paragraph>
|
|
<paragraph id="90296E33FFD3A569FF6DF8D2FD00F854" blockId="9.[151,1437,1848,2044]" pageId="9" pageNumber="91">
|
|
<emphasis id="A2E2B221FFD3A569FF6DF8D2FEBCF8FB" bold="true" box="[151,358,1849,1875]" pageId="9" pageNumber="91">Female genitalia</emphasis>
|
|
(
|
|
<figureCitation id="08AD72B6FFD3A569FE82F8D3FE0AF8FB" box="[376,464,1848,1875]" captionStart="Figures 15-18" captionStartId="6.[151,236,1903,1927]" captionTargetBox="[293,1306,199,1864]" captionTargetId="figure-11@6.[198,1388,189,1884]" captionTargetPageId="6" captionText="Figures 15-18: female genitalia. 15, Ezishnola cretacea (Hampson, 1914), LGNA 990, Ghana, Bunso Arboretum (HNHM); 16, E. inopinata sp. n., holotype, LGNA 692, Zambia, Lukwakwa (ANHRT); 17, E. carcassoni sp. n. holotype, NHMUK 010316559 (Gy. M. László), Uganda, Kalinzu Forest (NHMUK); 18, E. fuscographa László, Ronkay & Witt, 2010, paratype, LGN 971, Thailand, Bo Luang (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.13231376" httpUri="https://zenodo.org/record/13231376/files/figure.png" pageId="9" pageNumber="91">Fig. 17</figureCitation>
|
|
). Ovipositor short, trapezoidal, papillae anales conical; apophyses posteriores and anteriores short, relatively thin, of same length; 8
|
|
<superScript id="67E3C37BFFD3A569FD11F8B3FD20F8C1" attach="left" box="[747,762,1880,1897]" fontSize="7" pageId="9" pageNumber="91">th</superScript>
|
|
tergite conspicuously short, weakly sclerotized, ribbon-like; ostium bursae moderately sclerotized, goblet-shaped, relatively short and narrow; ductus bursae very short, relatively narrow, medial half more strongly sclerotized than distal and proximal quarters, cervix bursae membranous, rather thick, without sack-like protrusion; corpus bursae membranous, elongate-ovoid without distal tubular part; signum bursae absent.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D88C3DB8FFD0A56AFF6DFF2FFEA7FEA8" pageId="10" pageNumber="92" type="etymology">
|
|
<paragraph id="90296E33FFD0A56AFF6DFF2FFEA7FEA8" blockId="10.[151,1434,195,256]" pageId="10" pageNumber="92">
|
|
<emphasis id="A2E2B221FFD0A56AFF6DFF2FFEFCFF76" bold="true" box="[151,294,196,222]" pageId="10" pageNumber="92">Etymology.</emphasis>
|
|
The new species is named after the British entomologist Robert Herbert Carcasson, collector of the
|
|
<typeStatus id="4F2DD091FFD0A56AFF38FF0DFF2FFEA8" box="[194,245,230,256]" pageId="10" pageNumber="92">type</typeStatus>
|
|
specimens.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="D88C3DB8FFD0A56AFF6DFEC2FB2FFECD" pageId="10" pageNumber="92" type="discussion">
|
|
<paragraph id="90296E33FFD0A56AFF6DFEC2FB2FFECD" blockId="10.[151,1435,297,357]" pageId="10" pageNumber="92">
|
|
<emphasis id="A2E2B221FFD0A56AFF6DFEC2FEDEFEEB" bold="true" box="[151,260,297,323]" pageId="10" pageNumber="92">Remark.</emphasis>
|
|
An unidentified specimen is illustrated in
|
|
<bibRefCitation id="F40713C2FFD0A56AFCEAFEC2FBDDFEEC" author="Hacker, H. H. & Schreier, H. P. & Goater, B." box="[784,1031,297,324]" pageId="10" pageNumber="92" pagination="7 - 612" refId="ref6010" refString="Hacker, H. H., Schreier, H. P. & Goater, B. (2012) Revision of the tribe Nolini of Africa and the Western Palaearctic Region (Lepidoptera, Noctuoidea, Noctuidae, Nolinae). Esperiana, 17, 7 - 612." type="journal article" year="2012">
|
|
Hacker
|
|
<emphasis id="A2E2B221FFD0A56AFC8AFEC1FC7DFEEC" box="[880,935,297,324]" italics="true" pageId="10" pageNumber="92">et al</emphasis>
|
|
. (2012)
|
|
</bibRefCitation>
|
|
on p. 529 from Budongo Forest,
|
|
<collectingCountry id="E8812EA3FFD0A56AFF6DFEA1FF2CFECD" box="[151,246,330,357]" name="Uganda" pageId="10" pageNumber="92">Uganda</collectingCountry>
|
|
. Based on its external habitus, this specimen almost certainly belongs to
|
|
<taxonomicName id="579615B0FFD0A56AFBB7FEA0FB37FECC" authorityName="László" authorityYear="2020" box="[1101,1261,331,356]" class="Insecta" family="Nolidae" genus="Ezishnola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="92" phylum="Arthropoda" rank="species" species="carcassoni">
|
|
<emphasis id="A2E2B221FFD0A56AFBB7FEA0FB37FECC" box="[1101,1261,331,356]" italics="true" pageId="10" pageNumber="92">E. carcassoni</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |