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172 lines
22 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan & Machard, Patrice" docDate="2022" docId="0E9C3806E0B75CE39EE76CD094A8C9DF" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) insignitus Bedel 1902" docType="treatment" docVersion="2" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1655206106596" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gueorguiev, Borislav</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
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<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
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<mods:nameIdentifier type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wrase, David W.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
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<mods:affiliation>Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
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<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
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<mods:affiliation>Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Machard, Patrice</mods:namePart>
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<mods:affiliation>29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-06-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>69</mods:number>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:start>71</mods:start>
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<mods:end>122</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/dez.69.83840</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.69.83840</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-1324-1-71</mods:identifier>
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<mods:identifier type="ZooBank">7DEB0587A3734B06BAADCA6102EC6581</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:0E9C3806E0B75CE39EE76CD094A8C9DF" httpUri="http://treatment.plazi.org/id/0E9C3806E0B75CE39EE76CD094A8C9DF" lastPageNumber="71" pageId="0" pageNumber="71">
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<subSubSection pageId="0" pageNumber="71" type="nomenclature">
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<paragraph pageId="0" pageNumber="71">
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16.
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<taxonomicName LSID="0E9C3806-E0B7-5CE3-9EE7-6CD094A8C9DF" authority="Bedel, 1902" authorityName="Bedel" authorityYear="1902" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="insignitus" subGenus="Eremoderus">Platyderus (Eremoderus) insignitus Bedel, 1902</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="description">
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<paragraph pageId="0" pageNumber="71">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">Figs 4A</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">, 9E</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">, 11D</figureCitation>
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">, 15C</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="diagnosis">
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<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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This species differs from
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<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
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<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
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</taxonomicName>
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in pronotum less constricted to base (PW/PB: 1.12-1.17, vs. PW/PB: 1.21-1.28) and with apex appreciably narrower than base (PA/PB: 0.80-0.85, vs. PA/PB: 0.88-0.92). Median lobe of aedeagus somewhat longer than that in
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
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</taxonomicName>
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(s.l.), with ventral sclerite longer and not forming a distal kink (Figs
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9E</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="71" type="redescription">
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<paragraph pageId="0" pageNumber="71">Redescription.</paragraph>
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<paragraph pageId="0" pageNumber="71">
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
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.
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</emphasis>
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Moderately large species (BL: 6.60-8.70 mm; BW: 2.20-3.00 mm), with elongate, narrow and moderately convex body (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dorsal habitus. A. Platyderus (Eremoderus) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; B. P. (E.) irakensis, sp. nov., holotype; C. P. (E.) jordanensis, sp. nov., holotype; D. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype and its labels; E. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, " Jerusalem Syria ", Jerusalem District, Israel. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure4" httpUri="https://binary.pensoft.net/fig/695641" pageId="0" pageNumber="71">4A</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
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.
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</emphasis>
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See Table
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<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
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.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
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.
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</emphasis>
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Chestnut-colored as palpi, with antennae and legs usually lighter than rest of body, head somewhat darker than pronotum and elytra. Integument shiny.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
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.
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</emphasis>
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Pronotum and elytra with isodiametric sculpticells. Ventral surface with impressed slight stretched isodiametric sculpticells or with microsculpture less apparent or scarcely-visible. Head impunctate, usually smooth, but lateral areas adjacent to frontal furrows sometimes with fine wrinkles. Surface of pronotum impunctate, largely smooth, basal area medially behind posterior transverse impression longitudinally wrinkled, rarely apical part medially in front of anterior transverse impression also wrinkled; basal foveae and adjacent lateral areas smooth. Elytral intervals without apparent punctation. Abdominal ventrite 1 finely wrinkled in whole, 2-5 finely wrinkled at sides.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
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.
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</emphasis>
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About two thirds as wide as pronotum (PW/HW= 1.35-1.45). Eye long, convex. Labrum slightly shorter than clypeus, subrectangular, with anterior margin concave medially. Frontoclypeal suture more or less distinct in middle, indistinct or disappeared at sides. Frontal furrows very small, shallow, punctiform. Paraorbital sulci fine, backward not reaching level of posterior supraorbital pore.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
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. Pronotum about a tenth to a fifth wider than long (PW/PL= 1.06-1.17), widest point at second quarter. Anterior transverse impression absent, posterior transverse impression very shallow, barely distinct. Sides convex medially and anteriorly, slightly concave to base (somewhat more concave than in
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<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
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</taxonomicName>
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); anterior bead present laterally, reduced or lack in medial eight to tenth; basal bead present laterally, reduced to absent in medial fifth to half. Metepisternum slightly longer than wide or nearly as long as wide, MA/MI= 0.79-0.86.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
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. Oblong, about one-and-a-half times, or a little more, as long as elytra (EL/EW= 1.57-1.65), two-and-a-half times as long as pronotum (EL/PL= 2.41-2.61), and a half as wide as pronotum (EW/PW= 1.40-1.51), with widest point at medial third. Parascutellar striole and striae 1-7 rather shallow, impunctate or slightly punctate, stria 8 deeper than other striae; parascutellar striole not joining stria 1; bases of striae 1-7 mostly reduced, not reaching basal bead, rarely reaching it. Interval 3 with three discal setiferous punctures. Umbilicate setiferous series with 16 or 17 punctures on each side.
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
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.
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</emphasis>
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Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur mostly with 5-6 setiferous punctures on anterior side ventrally, rarely with 4 punctures. Anterior side of metafemur ventrally mostly with one seta in basal third and one in medial third or apical half (single specimen with two setae in basal half and two in apical half on one femur).
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<emphasis bold="true" pageId="0" pageNumber="71">
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<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
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.
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</emphasis>
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Median lobe of aedeagus in lateral view arcuate, with narrow basal bulb, long shaft and short, straight apex (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9E</figureCitation>
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); same in ventral view straight, about 3.6-3.7 times longer than wide; apical lamella (dorsal view) short, almost symmetrical, rounded at tip and slightly concave at each side. Internal sac in lateral view (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Median lobe of aedeagus, left lateral view (black arrows on D, F-H indicate convexity of ventral margin of apex of median lobe). A. Platyderus (Platyderus) sp., male specimen, Boz Daglar Mtn., Turkey; B. P. (Eremoderus) afghanistanicus, sp. nov., holotype; C. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; D. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; E. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; F. P. (E.) jordanensis, sp. nov., holotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; H. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure9" httpUri="https://binary.pensoft.net/fig/695646" pageId="0" pageNumber="71">9E</figureCitation>
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) with ventral sclerite long, slightly broadened distally; same in ventral view (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Median lobe of aedeagus, ventral view. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; C. P. (E.) brunneus ferrantei Reitter, 1909, male specimen, Holot Haluza, Southern District, Israel; D. P. (E.) insignitus insignitus Bedel, 1902, male specimen, SW Tiznit, Sous-Massa Region, Morocco; E. P. (E.) jordanensis, sp. nov., holotype; F. P. (E.) languidus (Reiche & Saulcy, 1855), lectotype; G. P. (E.) languidus (Reiche & Saulcy, 1855), male specimen, Na ẖal Prat, Judea and Samaria Area, Israel. Scale bars: 0.2 mm." figureDoi="10.3897/dez.69.83840.figure11" httpUri="https://binary.pensoft.net/fig/695648" pageId="0" pageNumber="71">11D</figureCitation>
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), with dorsal sclerite large, forming two small left-sided protuberances and a broad right-sided protuberance, and with ventral sclerite narrow, straight and slightly broadened distally.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) brunneus brunneus Karsch, 1881, female specimen, Bukamash, Nuqat al Khams District, Libya; B. P. (E.) brunneus ferrantei Reitter, 1909, female specimen, Holot Haluza, Southern District, Israel; C. P. (E.) insignitus insignitus Bedel, 1902, female specimen, Plage Aglou, Sous-Massa Region, Morocco; D. P. (E.) irakensis, sp. nov., holotype; E. P. (E.) jordanensis, sp. nov., female paratype, N Wadi Musa, Ma'an Governorate, Jordan; F. P. (E.) languidus (Reiche & Saulcy, 1855), female specimens, Israel, Netanya 22. xii. 1996, Central District, Israel. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure15" httpUri="https://binary.pensoft.net/fig/695652" pageId="0" pageNumber="71">15C</figureCitation>
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). Apical gonocoxite with rounded apex and one dorsolateral ensiform seta. Spermathecal canal connected in medial third of receptaculum.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |