194 lines
20 KiB
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194 lines
20 KiB
XML
<document id="AB85903F6070317BDF9863DDD3090A01" ID-CLB-Dataset="20341" ID-DOI="10.11646/zootaxa.5105.3.1" ID-GBIF-Dataset="797fca93-60bf-40ce-bb55-e5cb695f6356" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332761" ID-ZooBank="187B04D7-4C35-4E27-9B2D-A616BF59F380" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646597857429" checkinUser="plazi" docAuthor="Watson, Gillian W." docDate="2022" docId="03D687D0FF82987E19E8CB82FA2FE5B2" docLanguage="en" docName="zootaxa.5105.3.1.pdf" docOrigin="Zootaxa 5105 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudaspidoproctus Morrison" docType="treatment" docVersion="9" lastPageNumber="337" masterDocId="FFEFFFA8FFA6985A197FCA59FFB2E100" masterDocTitle="Towards identification of the scale insects (Hemiptera: Coccomorpha) of continental Africa: 2. Checklists and keys to six archaeococcoid families" masterLastPageNumber="356" masterPageNumber="301" pageNumber="337" updateTime="1699317191505" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="05E4D893F556BB8151F4CF8306ECFFCB">
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<mods:title id="CDA45E87B13E0619130C94AA7A4321B1">Towards identification of the scale insects (Hemiptera: Coccomorpha) of continental Africa: 2. Checklists and keys to six archaeococcoid families</mods:title>
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<mods:name id="107A7149F2E7188EB7A38FC20030C589" type="personal">
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<mods:namePart id="C02323DAD0E989A6C45AF3DAA4C19043">Watson, Gillian W.</mods:namePart>
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<mods:typeOfResource id="58F2EF22A1BD2167C2FEFE2C53636B44">text</mods:typeOfResource>
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<mods:title id="D2A35727B0AE4BFEF5E5E8F74BAE33F3">Zootaxa</mods:title>
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<mods:part id="6FCE6AFF0A2776BD810B9ED8CAB90FAB">
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<mods:date id="FED16038C17E8D1481B8B53D56BC89D0">2022</mods:date>
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<mods:number id="343FEF781E14F1AC5E29AA40D18B0F60">2022-03-04</mods:number>
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<mods:number id="038B4EF6324790300A2F3EB7954F7515">5105</mods:number>
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<mods:number id="AB8B88C84FBCD5545DE295194A1D579D">3</mods:number>
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<mods:identifier id="99808E80E24C965F595D448DB101CFB0" type="DOI">10.11646/zootaxa.5105.3.1</mods:identifier>
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<mods:identifier id="8E7EBA3C3A351A0F697450F617ECE5DF" type="GBIF-Dataset">797fca93-60bf-40ce-bb55-e5cb695f6356</mods:identifier>
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<treatment id="03D687D0FF82987E19E8CB82FA2FE5B2" ID-DOI="http://doi.org/10.5281/zenodo.6343214" ID-GBIF-Taxon="193890941" ID-Zenodo-Dep="6343214" LSID="urn:lsid:plazi:treatment:03D687D0FF82987E19E8CB82FA2FE5B2" httpUri="http://treatment.plazi.org/id/03D687D0FF82987E19E8CB82FA2FE5B2" lastPageNumber="337" pageId="36" pageNumber="337">
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<subSubSection id="C365654DFF82987E19E8CB82FDE5E0F5" box="[151,599,475,502]" pageId="36" pageNumber="337" type="nomenclature">
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<paragraph id="8BC036C6FF82987E19E8CB82FDE5E0F5" blockId="36.[151,599,475,502]" box="[151,599,475,502]" pageId="36" pageNumber="337">
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<heading id="D08881AAFF82987E19E8CB82FDE5E0F5" bold="true" box="[151,599,475,502]" fontSize="11" level="1" pageId="36" pageNumber="337" reason="1">
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<emphasis id="B90BEAD4FF82987E19E8CB82FDE5E0F5" bold="true" box="[151,599,475,502]" pageId="36" pageNumber="337">
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Genus
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<taxonomicName id="4C7F4D45FF82987E1991CB82FDE5E0F5" ID-CoL="6438Z" authority="Morrison" authorityName="Morrison" box="[238,599,475,502]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B90BEAD4FF82987E1991CB82FE69E0F6" bold="true" box="[238,475,475,502]" italics="true" pageId="36" pageNumber="337">Pseudaspidoproctus</emphasis>
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Morrison
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C365654DFF82987E19E8C87AFA3FE239" pageId="36" pageNumber="337" type="reference_group">
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<paragraph id="8BC036C6FF82987E19E8C87AFD19E3F1" blockId="36.[151,1437,547,826]" pageId="36" pageNumber="337">
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<taxonomicName id="4C7F4D45FF82987E19E8C87AFEC7E33D" ID-CoL="6438Z" authorityName="Morrison" box="[151,373,547,573]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B90BEAD4FF82987E19E8C87AFEC7E33D" box="[151,373,547,573]" italics="true" pageId="36" pageNumber="337">Pseudaspidoproctus</emphasis>
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</taxonomicName>
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is an African genus of nine described species that have not been thoroughly studied, two of which have not been recorded from continental Africa (
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<bibRefCitation id="EFEE4B37FF82987E1BD2C81EFC6DE362" author="Garcia Morales, M. & Denno, B. D. & Miller, D. R. & Miller, G. L. & Ben-Dov, Y. & Hardy, N. B." box="[685,991,583,610]" pageId="36" pageNumber="337" refId="ref29245" refString="Garcia Morales, M., Denno, B. D., Miller, D. R., Miller, G. L., Ben-Dov, Y. & Hardy, N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 12 January 2022) https: // doi. org / 10.1093 / database / bav 118." type="url" year="2016">
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García Morales
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<emphasis id="B90BEAD4FF82987E1A1CC811FC27E361" box="[867,917,583,609]" italics="true" pageId="36" pageNumber="337">et al</emphasis>
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. 2016
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</bibRefCitation>
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). Now that
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<taxonomicName id="4C7F4D45FF82987E1D21C81EFF6EE385" ID-CoL="4NLGN" baseAuthorityName="Newstead" baseAuthorityYear="1911" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="zimmermanni">
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<emphasis id="B90BEAD4FF82987E1D21C81EFF6EE385" italics="true" pageId="36" pageNumber="337">Pseudaspidoproctus zimmermanni</emphasis>
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</taxonomicName>
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has been transferred to
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<taxonomicName id="4C7F4D45FF82987E18A2C832FDC9E385" ID-CoL="344Y" authorityName="Newstead" box="[477,635,619,645]" class="Insecta" family="Margarodidae" genus="Aspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B90BEAD4FF82987E18A2C832FDC9E385" box="[477,635,619,645]" italics="true" pageId="36" pageNumber="337">Aspidoproctus</emphasis>
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</taxonomicName>
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(see the new combination made above), there are six species found on the continent. They are quite large for scale insects; the anterior margin of the marsupium lacks a dense band of pores, so in life the marsupial opening is not covered by a flap of felted wax. The insects prefer woody hosts (
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<bibRefCitation id="EFEE4B37FF82987E1D92C8EAFEA2E3F2" author="Garcia Morales, M. & Denno, B. D. & Miller, D. R. & Miller, G. L. & Ben-Dov, Y. & Hardy, N. B." pageId="36" pageNumber="337" refId="ref29245" refString="Garcia Morales, M., Denno, B. D., Miller, D. R., Miller, G. L., Ben-Dov, Y. & Hardy, N. B. (2016) ScaleNet: a literature-based model of scale insect biology and systematics. Database. Available from: http: // scalenet. info (accessed 12 January 2022) https: // doi. org / 10.1093 / database / bav 118." type="url" year="2016">
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García Morales
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<emphasis id="B90BEAD4FF82987E19E8C881FF7AE3F1" box="[151,200,727,753]" italics="true" pageId="36" pageNumber="337">et al</emphasis>
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. 2016
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</bibRefCitation>
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) and are of no economic importance.
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</paragraph>
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<paragraph id="8BC036C6FF82987E19B8C8A2FA3FE239" blockId="36.[151,1437,547,826]" pageId="36" pageNumber="337">
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The material studied included
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<typeStatus id="54C48864FF82987E1B6FC8A5FDC2E216" box="[528,624,764,790]" pageId="36" pageNumber="337" type="syntype">syntypes</typeStatus>
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of
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<taxonomicName id="4C7F4D45FF82987E1BEDC8A5FCA1E215" ID-CoL="448L3" baseAuthorityName="Newstead" baseAuthorityYear="1912" box="[658,787,763,789]" class="Insecta" family="Margarodidae" genus="Monophleboides" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="africanus">
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<emphasis id="B90BEAD4FF82987E1BEDC8A5FCA1E215" box="[658,787,763,789]" italics="true" pageId="36" pageNumber="337">P. africanus</emphasis>
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</taxonomicName>
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, the
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<typeStatus id="54C48864FF82987E1A39C8A2FC19E215" box="[838,939,763,789]" pageId="36" pageNumber="337" type="lectotype">lectotype</typeStatus>
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of
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<emphasis id="B90BEAD4FF82987E1AB3C8A5FB36E215" box="[972,1156,763,789]" italics="true" pageId="36" pageNumber="337">
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<taxonomicName id="4C7F4D45FF82987E1AB3C8A5FB32E215" ID-CoL="4NLGH" baseAuthorityName="Hall" baseAuthorityYear="1925" box="[972,1152,763,789]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="hyphaeniacus">P. hyphaeniacus</taxonomicName>
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,
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</emphasis>
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and a
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<typeStatus id="54C48864FF82987E1DB7C8A5FA95E216" box="[1224,1319,764,790]" pageId="36" pageNumber="337" type="paratype">paratype</typeStatus>
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of
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<taxonomicName id="4C7F4D45FF82987E1C36C8A5FF4EE239" ID-CoL="4NLGL" authorityName="Ghesquiere" authorityYear="1943" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="vayssieriellus">
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<emphasis id="B90BEAD4FF82987E1C36C8A5FF4EE239" italics="true" pageId="36" pageNumber="337">P. vayssieriellus</emphasis>
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</taxonomicName>
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but no type material of
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<taxonomicName id="4C7F4D45FF82987E1B7CC979FDC1E239" ID-CoL="4NLG8" baseAuthorityName="Joubert" baseAuthorityYear="1925" box="[515,627,800,825]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="acaciae">
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<emphasis id="B90BEAD4FF82987E1B7CC979FDC1E239" box="[515,627,800,825]" italics="true" pageId="36" pageNumber="337">P. acaciae</emphasis>
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</taxonomicName>
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. No material of
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<taxonomicName id="4C7F4D45FF82987E1A57C979FCCFE239" ID-CoL="4NLGD" baseAuthorityName="Cockerell" baseAuthorityYear="1901" box="[808,893,799,825]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="fortis">
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<emphasis id="B90BEAD4FF82987E1A57C979FCCFE239" box="[808,893,799,825]" italics="true" pageId="36" pageNumber="337">P. fortis</emphasis>
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</taxonomicName>
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was seen, so it is not included in the key below.
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</paragraph>
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</subSubSection>
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<subSubSection id="C365654DFF82987E19E8C9D2FA2FE5B2" pageId="36" pageNumber="337" type="key">
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<paragraph id="8BC036C6FF82987E19E8C9D2FBFAE2A5" blockId="36.[151,1096,907,934]" box="[151,1096,907,934]" pageId="36" pageNumber="337">
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<heading id="D08881AAFF82987E19E8C9D2FBFAE2A5" box="[151,1096,907,934]" pageId="36" pageNumber="337">
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<emphasis id="B90BEAD4FF82987E19E8C9D2FCF5E2A5" bold="true" box="[151,839,907,934]" pageId="36" pageNumber="337">
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Key to species of
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<taxonomicName id="4C7F4D45FF82987E181BC9D2FDF8E2A6" ID-CoL="6438Z" box="[356,586,907,934]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="genus">
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<emphasis id="B90BEAD4FF82987E181BC9D2FDF8E2A6" bold="true" box="[356,586,907,934]" italics="true" pageId="36" pageNumber="337">Pseudaspidoproctus</emphasis>
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</taxonomicName>
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in continental Africa
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</emphasis>
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, not including
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<taxonomicName id="4C7F4D45FF82987E1A8EC9D5FBFAE2A5" ID-CoL="4NLGD" baseAuthorityName="Cockerell" baseAuthorityYear="1901" box="[1009,1096,907,933]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="fortis">
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<emphasis id="B90BEAD4FF82987E1A8EC9D5FBFAE2A5" box="[1009,1096,907,933]" italics="true" pageId="36" pageNumber="337">P. fortis</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<key id="FFEE61F7FF82987E19E8C989FA2FE5B2" pageId="36" pageNumber="337">
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<keyStep id="308B2E63FF82987E19E8C989FA2EE503" pageId="36" pageNumber="337">
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<paragraph id="8BC036C6FF82987E19E8C989FA2EE2E5" blockId="36.[151,1437,976,1203]" box="[151,1436,976,998]" pageId="36" pageNumber="337">
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<keyLead id="308E95F3FF82987E19E8C989FA2EE2E5" box="[151,1436,976,998]" pageId="36" pageNumber="337">1(0) Dermal spines present. With 3 oblong cicatrices. Internal ring on anal tube with pores.............................. 2</keyLead>
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</paragraph>
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<paragraph id="8BC036C6FF82987E19E8C9B7FA2EE503" blockId="36.[151,1437,976,1203]" box="[151,1436,1005,1027]" pageId="36" pageNumber="337">
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<keyLead id="308E95F3FF82987E19E8C9B7FA2EE503" box="[151,1436,1005,1027]" pageId="36" pageNumber="337">- Dermal spines absent. With 1 or 3 subcircular cicatrices. Internal ring on anal tube without pores...................... 3</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308B2E63FF82987E19E8CE52FA2EE53E" pageId="36" pageNumber="337">
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<paragraph id="8BC036C6FF82987E19E8CE52FA2EE520" blockId="36.[151,1437,976,1203]" box="[151,1436,1034,1057]" pageId="36" pageNumber="337">
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<keyLead id="308E95F3FF82987E19E8CE52FA2EE520" box="[151,1436,1034,1057]" pageId="36" pageNumber="337">
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2(1) Marsupial opening U-shaped, anterior marsupial band not developed........................................
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<taxonomicName id="4C7F4D45FF82987E1C20CE53FA2EE520" ID-CoL="4NLGF" baseAuthorityName="Cockerell" baseAuthorityYear="1901" box="[1375,1436,1034,1056]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="fulleri">
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<emphasis id="B90BEAD4FF82987E1C20CE53FA2EE520" bold="true" box="[1375,1436,1034,1056]" italics="true" pageId="36" pageNumber="337">fulleri</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BC036C6FF82987E19E8CE71FA2EE53E" blockId="36.[151,1437,976,1203]" box="[151,1436,1064,1086]" pageId="36" pageNumber="337">
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<keyLead id="308E95F3FF82987E19E8CE71FA2EE53E" box="[151,1436,1064,1086]" pageId="36" pageNumber="337">
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- Marsupial opening V-shaped, anterior marsupial band sclerotized.............................
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<taxonomicName id="4C7F4D45FF82987E1DC6CE71FAF0E53E" ID-CoL="4NLGH" baseAuthorityName="Hall" baseAuthorityYear="1925" box="[1209,1346,1064,1086]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="hyphaeniacus">
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<emphasis id="B90BEAD4FF82987E1DC6CE71FAF0E53E" bold="true" box="[1209,1346,1064,1086]" italics="true" pageId="36" pageNumber="337">hyphaeniacus</emphasis>
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</taxonomicName>
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(
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<figureCitation id="13442A43FF82987E1C30CE71FA27E53E" box="[1359,1429,1064,1086]" captionStart="FIGURE 19" captionStartId="29.[151,250,1791,1815]" captionTargetBox="[160,1427,195,1757]" captionTargetId="figure-15@29.[151,1436,181,1768]" captionTargetPageId="29" captionText="FIGURE 19. Pseudaspidoproctus hyphaeniacus (Hall), adult female, developed from Morrison (1928: 139, Fig. 66). C: cicatrices; M: marsupium opening." figureDoi="http://doi.org/10.5281/zenodo.6332799" httpUri="https://zenodo.org/record/6332799/files/figure.png" pageId="36" pageNumber="337">Fig. 19</figureCitation>
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)
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308B2E63FF82987E19E8CE1CFA2EE578" pageId="36" pageNumber="337">
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<paragraph id="8BC036C6FF82987E19E8CE1CFA2EE55B" blockId="36.[151,1437,976,1203]" box="[151,1436,1093,1115]" pageId="36" pageNumber="337">
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<keyLead id="308E95F3FF82987E19E8CE1CFA2EE55B" box="[151,1436,1093,1115]" pageId="36" pageNumber="337">
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3(1) With 1 subcircular cicatrice. Marsupial opening V-shaped but distal acute angle rounded..................
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<taxonomicName id="4C7F4D45FF82987E1C67CE1CFA2EE55B" ID-CoL="4NLGL" authorityName="Ghesquiere" authorityYear="1943" box="[1304,1436,1093,1115]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="vayssieriellus">
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<emphasis id="B90BEAD4FF82987E1C67CE1CFA2EE55B" bold="true" box="[1304,1436,1093,1115]" italics="true" pageId="36" pageNumber="337">vayssieriellus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BC036C6FF82987E19E8CE3AFA2EE578" blockId="36.[151,1437,976,1203]" box="[151,1436,1123,1145]" pageId="36" pageNumber="337">
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<keyLead id="308E95F3FF82987E19E8CE3AFA2EE578" box="[151,1436,1123,1145]" pageId="36" pageNumber="337">- With 3 subcircular cicatrices. Marsupial opening sharply V-shaped or absent...................................... 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308B2E63FF82987E19E8CED9FA2FE5B2" pageId="36" pageNumber="337">
|
|
<paragraph id="8BC036C6FF82987E19E8CED9FA2FE596" blockId="36.[151,1437,976,1203]" box="[151,1437,1152,1174]" pageId="36" pageNumber="337">
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<keyLead id="308E95F3FF82987E19E8CED9FA2FE596" box="[151,1437,1152,1174]" pageId="36" pageNumber="337">
|
|
4(3) Marsupial opening sharply V-shaped. Mature female sclerotized, cuticle brown.............................
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<taxonomicName id="4C7F4D45FF82987E1C41CED9FA2FE596" ID-CoL="448L3" baseAuthorityName="Newstead" baseAuthorityYear="1912" box="[1342,1437,1152,1174]" class="Insecta" family="Margarodidae" genus="Monophleboides" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="africanus">
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<emphasis id="B90BEAD4FF82987E1C41CED9FA2FE596" bold="true" box="[1342,1437,1152,1174]" italics="true" pageId="36" pageNumber="337">africanus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="8BC036C6FF82987E19E8CEC7FA2FE5B2" blockId="36.[151,1437,976,1203]" box="[151,1437,1181,1203]" pageId="36" pageNumber="337">
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|
<keyLead id="308E95F3FF82987E19E8CEC7FA2FE5B2" box="[151,1437,1181,1203]" pageId="36" pageNumber="337">
|
|
- Marsupial opening absent. Mature female cuticle membranous............................................
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<taxonomicName id="4C7F4D45FF82987E1C2DCEC4FA2FE5B2" ID-CoL="4NLG8" baseAuthorityName="Joubert" baseAuthorityYear="1925" box="[1362,1437,1181,1202]" class="Insecta" family="Margarodidae" genus="Pseudaspidoproctus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="337" phylum="Arthropoda" rank="species" species="acaciae">
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<emphasis id="B90BEAD4FF82987E1C2DCEC4FA2FE5B2" bold="true" box="[1362,1437,1181,1202]" italics="true" pageId="36" pageNumber="337">acaciae</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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</key>
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</subSubSection>
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</treatment>
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</document> |