549 lines
89 KiB
XML
549 lines
89 KiB
XML
<document id="08542695F8A25C10FB7E3A746256997F" ID-CLB-Dataset="30402" ID-DOI="10.11646/zootaxa.4402.2.3" ID-GBIF-Dataset="7139708e-922d-41d4-ad42-a5d5f7076f1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="10070885" ID-ZooBank="CEA5700E-9F94-42A8-9067-BD8F4D301B9D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1522215469332" checkinUser="plazi" docAuthor="Huynh, Cuong" docDate="2018" docId="03AB2952FFE45E5D0FBDFB8AFD8F0B51" docLanguage="en" docName="zootaxa.4402.2.3.pdf.imf" docOrigin="Zootaxa 4402 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Monographis phuquocensis Huynh 2018, sp. n." docType="treatment" docVersion="8" lastPageNumber="298" masterDocId="FF92512AFFEE5E520F2AFFE1FFB20949" masterDocTitle="Two new species of penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam" masterLastPageNumber="302" masterPageNumber="283" pageNumber="293" updateTime="1699558794197" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="684038812930192D3CF49E85A7B4B7E4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="977B744A8DFE2FB64558B70DBD081319">
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<mods:title id="34394262A8E5AED6FACB81EAFC92293B">Two new species of penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam</mods:title>
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</mods:titleInfo>
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<mods:name id="B37EB1F34F182D454F7B95A918A6E92E" type="personal">
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<mods:roleTerm id="1CAA6340836D6BBA692EA9CDA5495DFA">Author</mods:roleTerm>
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<mods:namePart id="9EC1DC993E235BB05ED1E86F24ABE126">Huynh, Cuong</mods:namePart>
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<mods:title id="AC8E58CD61F3C8EC17AC2017393BBF79">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B0D4F398D13DC389877CC913B9C01991">
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<mods:date id="B36C4C702696624BF1DCFCBDC5F6E0B1">2018</mods:date>
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<mods:number id="2824978BFFB2D970C190AFC620109F9F">2018-03-28</mods:number>
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<mods:number id="DA812AF7C1CB15FC997B812D4C973F61">4402</mods:number>
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<mods:detail id="73CA92AEDDA75C5182783C77621E97F1" type="issue">
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<mods:number id="DD5F8C4A8E50BCE16BB9BFABE32305CE">2</mods:number>
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<mods:start id="F5FCE98C042E7F509ABB6525ACDF49EA">283</mods:start>
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<mods:classification id="D5FACCC2F5A13F97DB13043D5912438F">journal article</mods:classification>
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<mods:identifier id="11020C577C5270FB672B6BEBB0365144" type="DOI">10.11646/zootaxa.4402.2.3</mods:identifier>
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<mods:identifier id="EE94A2F26689075D074D82752AAFD173" type="GBIF-Dataset">7139708e-922d-41d4-ad42-a5d5f7076f1a</mods:identifier>
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</mods:mods>
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<treatment id="03AB2952FFE45E5D0FBDFB8AFD8F0B51" ID-GBIF-Taxon="163748925" LSID="urn:lsid:plazi:treatment:03AB2952FFE45E5D0FBDFB8AFD8F0B51" httpUri="http://treatment.plazi.org/id/03AB2952FFE45E5D0FBDFB8AFD8F0B51" lastPageId="15" lastPageNumber="298" pageId="10" pageNumber="293">
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<subSubSection id="C318CBCFFFE45E580FBDFB8AFD830DCF" box="[151,561,1131,1158]" pageId="10" pageNumber="293" type="nomenclature">
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<paragraph id="8BBD9844FFE45E580FBDFB8AFD830DCF" blockId="10.[151,561,1131,1191]" box="[151,561,1131,1158]" pageId="10" pageNumber="293">
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<heading id="D0F52F28FFE45E580FBDFB8AFD830DCF" bold="true" box="[151,561,1131,1158]" fontSize="11" level="1" pageId="10" pageNumber="293" reason="1">
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<emphasis id="B9764456FFE45E580FBDFB8AFD830DCF" bold="true" box="[151,561,1131,1158]" pageId="10" pageNumber="293">
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<taxonomicName id="4C02E3C7FFE45E580FBDFB8AFE550DCC" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[151,487,1131,1157]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="phuquocensis" status="sp. nov.">
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<emphasis id="B9764456FFE45E580FBDFB8AFE550DCC" bold="true" box="[151,487,1131,1157]" italics="true" pageId="10" pageNumber="293">Monographis phuquocensis</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A245F92DFFE45E580EC4FB8DFD830DCF" box="[494,561,1132,1158]" pageId="10" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C318CBCFFFE45E580FBDFB6FFF590DEE" box="[151,235,1166,1191]" pageId="10" pageNumber="293" type="description">
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<paragraph id="8BBD9844FFE45E580FBDFB6FFF590DEE" blockId="10.[151,561,1131,1191]" box="[151,235,1166,1191]" pageId="10" pageNumber="293">
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<emphasis id="B9764456FFE45E580FBDFB6FFF120DEE" bold="true" box="[151,160,1166,1191]" pageId="10" pageNumber="293">(</emphasis>
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<figureCitation id="133984C1FFE45E580F8AFB6FFF500DEE" box="[160,226,1166,1191]" captionStart="FIGURE 8" captionStartId="11.[151,250,1118,1140]" captionTargetBox="[202,1384,301,1097]" captionTargetId="figure@11.[202,1384,301,1097]" captionTargetPageId="11" captionText="FIGURE 8. A living specimen of Monographis phuquocensis sp. n. from Da Ban stream, Phu Quoc Island, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.10092680" httpUri="https://zenodo.org/record/10092680/files/figure.png" pageId="10" pageNumber="293">Fig. 8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C318CBCFFFE45E580FBDFB37FC630F7A" pageId="10" pageNumber="293" type="materials_examined">
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<paragraph id="8BBD9844FFE45E580FBDFB37FCA60C12" blockId="10.[151,1437,1238,2019]" pageId="10" pageNumber="293">
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<materialsCitation id="3B6A9219FFE45E580FBDFB37FCA60C12" ID-GBIF-Occurrence="2608820667" collectedFrom="leaf litter" collectingDate="2015-07-02" collectionCode="QMS" country="Australia" county="Brisbane" elevation="86" latitude="10.183044" location="Tranh Stream (Suoi Tranh)" longLatPrecision="1" longitude="104.0152" municipality="Museum" pageId="10" pageNumber="293" specimenCode="QMS 108531" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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||
<emphasis id="B9764456FFE45E580FBDFB37FEB90DA6" bold="true" box="[151,267,1238,1263]" pageId="10" pageNumber="293">
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||
<typeStatus id="54B926E6FFE45E580FBDFB37FEB40DA6" box="[151,262,1238,1263]" pageId="10" pageNumber="293" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
<specimenCount id="9D0453CDFFE45E580E3EFB37FEE20DA6" box="[276,336,1238,1263]" pageId="10" pageNumber="293" type="male">Male</specimenCount>
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,
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<taxonomicName id="4C02E3C7FFE45E580E75FB37FD2B0DA6" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[351,665,1238,1263]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="phuquocensis" status="sp. nov.">
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||
<emphasis id="B9764456FFE45E580E75FB37FD2B0DA6" box="[351,665,1238,1263]" italics="true" pageId="10" pageNumber="293">Monographis phuquocensis</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B9764456FFE45E580D88FB37FD560DA6" bold="true" box="[674,740,1238,1263]" pageId="10" pageNumber="293">
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||
<taxonomicNameLabel id="A245F92DFFE45E580D88FB37FD560DA6" box="[674,740,1238,1263]" pageId="10" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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was collected by the first author from
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<collectedFrom id="381EDBB9FFE45E580B87FB37FAA40DA6" box="[1197,1302,1238,1263]" pageId="10" pageNumber="293">leaf litter</collectedFrom>
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near
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||
<location id="8EDDCE9FFFE45E580A70FB37FE300C5A" LSID="urn:lsid:plazi:treatment:03AB2952FFE45E5D0FBDFB8AFD8F0B51:8EDDCE9FFFE45E580A70FB37FE300C5A" country="Australia" county="Brisbane" latitude="10.183044" longLatPrecision="1" longitude="104.0152" municipality="Museum" name="Tranh Stream (Suoi Tranh)" pageId="10" pageNumber="293" stateProvince="Queensland">Tranh Stream (Suoi Tranh)</location>
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,
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||
<geoCoordinate id="EE36FE83FFE45E580EA5FB1BFD8A0C5A" box="[399,568,1274,1299]" degrees="10" direction="north" minutes="10" orientation="latitude" pageId="10" pageNumber="293" precision="1" seconds="58.96" value="10.183044">10°10'58.96"N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE36FE83FFE45E580D6FFB1AFD470C5A" box="[581,757,1274,1299]" degrees="104" direction="east" minutes="0" orientation="longitude" pageId="10" pageNumber="293" precision="1" seconds="54.72" value="104.0152">104° 0'54.72"E</geoCoordinate>
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||
;
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||
<elevation id="002F7F77FFE45E580C28FB1BFC1D0C5B" box="[770,943,1274,1299]" metricMagnitude="1" metricUnit="m" metricValue="8.6" pageId="10" pageNumber="293" unit="m" value="86.0">
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||
elevation
|
||
<quantity id="4CFA35A1FFE45E580C58FB1BFC1D0C5B" box="[882,943,1274,1299]" metricMagnitude="1" metricUnit="m" metricValue="8.6" pageId="10" pageNumber="293" unit="m" value="86.0">86 m</quantity>
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||
</elevation>
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||
;
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||
<location id="8EDDCE9FFFE45E580C97FB1AFDAE0C7E" LSID="urn:lsid:plazi:treatment:03AB2952FFE45E5D0FBDFB8AFD8F0B51:8EDDCE9FFFE45E580C97FB1AFDAE0C7E" country="Australia" county="Brisbane" latitude="10.183044" longLatPrecision="1" longitude="104.0152" municipality="Museum" name="7 km southeast of the main town - Duong Dong town on Phu Quoc Island" pageId="10" pageNumber="293" stateProvince="Queensland">
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||
<quantity id="4CFA35A1FFE45E580C97FB1AFC4A0C5A" box="[957,1016,1274,1299]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="293" unit="km" value="7.0">7 km</quantity>
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||
southeast of the main town - Duong Dong town on Phu Quoc Island
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||
</location>
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||
on
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||
<date id="FFBCBE84FFE45E580D7EFAFFFCB70C7E" box="[596,773,1310,1335]" pageId="10" pageNumber="293" value="2015-07-02">
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||
<collectingDate id="EFF8476CFFE45E580D7EFAFFFCB70C7E" box="[596,773,1310,1335]" pageId="10" pageNumber="293" value="2015-07-02">2nd July 2015</collectingDate>
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||
</date>
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||
. Queensland Museum accession number:
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||
<specimenCode id="DBA4303FFFE45E580BD7FAFEFA2E0C7E" box="[1277,1436,1310,1335]" pageId="10" pageNumber="293">QMS 108531</specimenCode>
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||
(deposited in Queensland Museum, Brisbane, Australia).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BBD9844FFE45E580FEDFA87FC630F7A" blockId="10.[151,1437,1238,2019]" pageId="10" pageNumber="293">
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||
<materialsCitation id="3B6A9219FFE45E580FEDFA87FCD60CEA" ID-GBIF-Occurrence="2608822821" collectingDate="2015-07-03" collectionCode="VII" collectorName="Twelve & At & Da Ban Stream" country="Australia" county="Brisbane" elevation="43" latitude="10.2451" location="Suoi Da Ban" longLatPrecision="1" longitude="104.03023" municipality="Museum" pageId="10" pageNumber="293" specimenCount="12" specimenCount-adult="8" specimenCount-female="3" specimenCount-male="2" stateProvince="Queensland" typeStatus="paratype">
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||
<emphasis id="B9764456FFE45E580FEDFA87FEF40C36" bold="true" box="[199,326,1382,1407]" pageId="10" pageNumber="293">
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||
<typeStatus id="54B926E6FFE45E580FEDFA87FEF30C36" box="[199,321,1382,1407]" pageId="10" pageNumber="293" type="paratype">Paratypes</typeStatus>
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||
.
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</emphasis>
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Twelve specimens were collected:
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<specimenCount id="9D0453CDFFE45E580DCAFA86FC9D0C36" box="[736,815,1382,1407]" pageId="10" pageNumber="293" type="adult">3 adult</specimenCount>
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||
males,
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||
<specimenCount id="9D0453CDFFE45E580CA1FA87FC690C36" box="[907,987,1382,1407]" pageId="10" pageNumber="293" type="adult">5 adult</specimenCount>
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||
females (adult stage- stadium VIII, 13 pairs of legs), and
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||
<specimenCount id="9D0453CDFFE45E580E44FA6AFE4A0CEA" box="[366,504,1418,1443]" pageId="10" pageNumber="293">4 sub-adults</specimenCount>
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||
(stadium VII, 12 pairs of legs).
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||
</materialsCitation>
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||
<materialsCitation id="3B6A9219FFE45E580C47FA6AFC7F0F7A" ID-GBIF-Occurrence="2608822822" collectingDate="2015-07-03" collectionCode="VII" collectorName="Twelve & At & Da Ban Stream" country="Australia" county="Brisbane" elevation="43" latitude="10.2451" location="Da Ban Stream (Suoi Da Ban" longLatPrecision="1" longitude="104.03023" municipality="Museum" pageId="10" pageNumber="293" specimenCode="QMS 108532-108548" specimenCount="27" specimenCount-adult="8" specimenCount-female="3" specimenCount-male="2" stateProvince="Queensland" typeStatus="paratype">
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||
At the second location:
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||
<location id="8EDDCE9FFFE45E580B54FA6AFF7A0C8E" LSID="urn:lsid:plazi:treatment:03AB2952FFE45E5D0FBDFB8AFD8F0B51:8EDDCE9FFFE45E580B54FA6AFF7A0C8E" country="Australia" county="Brisbane" latitude="10.2451" longLatPrecision="1" longitude="104.03023" municipality="Museum" name="Da Ban Stream (Suoi Da Ban" pageId="10" pageNumber="293" stateProvince="Queensland">Da Ban Stream (Suoi Da Ban</location>
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||
),
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||
<geoCoordinate id="EE36FE83FFE45E580FF6FA4FFE370C8E" box="[220,389,1454,1479]" degrees="10" direction="north" minutes="14" orientation="latitude" pageId="10" pageNumber="293" precision="1" seconds="42.36" value="10.2451">10°14'42.36"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE36FE83FFE45E580EB8FA4EFDF50C8E" box="[402,583,1455,1479]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="10" pageNumber="293" precision="1" seconds="48.82" value="104.03023">104°01'48.82"E</geoCoordinate>
|
||
,
|
||
<elevation id="002F7F77FFE45E580D79FA4FFD4D0C8E" box="[595,767,1454,1479]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="293" unit="m" value="43.0">
|
||
elevation
|
||
<quantity id="4CFA35A1FFE45E580DE8FA4EFD4D0C8E" box="[706,767,1455,1479]" metricMagnitude="1" metricUnit="m" metricValue="4.3" pageId="10" pageNumber="293" unit="m" value="43.0">43 m</quantity>
|
||
</elevation>
|
||
;
|
||
<location id="8EDDCE9FFFE45E580C29FA4EFA2E0C8E" LSID="urn:lsid:plazi:treatment:03AB2952FFE45E5D0FBDFB8AFD8F0B51:8EDDCE9FFFE45E580C29FA4EFA2E0C8E" box="[771,1436,1454,1479]" country="Australia" county="Brisbane" latitude="10.2451" longLatPrecision="1" longitude="104.03023" municipality="Museum" name="10 km northeast of Duong Dong town on Phu Quoc Island" pageId="10" pageNumber="293" stateProvince="Queensland">
|
||
<quantity id="4CFA35A1FFE45E580C29FA4EFCF90C8E" box="[771,843,1454,1479]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="293" unit="km" value="10.0">10 km</quantity>
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||
northeast of Duong Dong town on Phu Quoc Island
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||
</location>
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||
on
|
||
<date id="FFBCBE84FFE45E580F95FA33FED50CA2" box="[191,359,1490,1515]" pageId="10" pageNumber="293" value="2015-07-03">
|
||
<collectingDate id="EFF8476CFFE45E580F95FA33FED50CA2" box="[191,359,1490,1515]" pageId="10" pageNumber="293" value="2015-07-03">3rd July 2015</collectingDate>
|
||
</date>
|
||
;
|
||
<specimenCount id="9D0453CDFFE45E580E5DFA33FDB50CA2" box="[375,519,1490,1515]" pageId="10" pageNumber="293" type="generic">5 specimens</specimenCount>
|
||
were collected:
|
||
<specimenCount id="9D0453CDFFE45E580DE4FA32FC9B0CA2" box="[718,809,1490,1515]" pageId="10" pageNumber="293" type="male">2 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D0453CDFFE45E580C40FA32FC520CA2" box="[874,992,1490,1515]" pageId="10" pageNumber="293" type="female">3 females</specimenCount>
|
||
. Both locations are in the Ham Ninh Mountains on Phu Quoc Island (
|
||
<figureCitation id="133984C1FFE45E580D01FA17FDC60F46" box="[555,628,1526,1551]" captionStart="FIGURE 1" captionStartId="3.[151,250,1969,1991]" captionTargetBox="[172,1396,196,1923]" captionTargetId="figure@3.[172,1396,195,1923]" captionTargetPageId="3" captionText="FIGURE 1. Phu Quoc Island with the inserted map of Vietnam indicating the location of Phu Quoc Island. Locations of Da Ban stream (·), Tranh stream (.) and Tranh river beach (▲) on Phu Quoc Island (Not to scale)." figureDoi="http://doi.org/10.5281/zenodo.10092666" httpUri="https://zenodo.org/record/10092666/files/figure.png" pageId="10" pageNumber="293">Fig. 1</figureCitation>
|
||
). Queensland Museum accession numbers for
|
||
<specimenCount id="9D0453CDFFE45E580B98FA16FAFF0F46" box="[1202,1357,1527,1551]" pageId="10" pageNumber="293" type="generic" typeStatus="paratypes">17 paratypes</specimenCount>
|
||
:
|
||
<specimenCode id="DBA4303FFFE45E580A4AFA16FEFF0F7A" pageId="10" pageNumber="293">QMS 108532–108548</specimenCode>
|
||
(deposited in Queensland Museum, Brisbane, Australia)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318CBCFFFE45E580FEDF9DFFD340F32" pageId="10" pageNumber="293" type="etymology">
|
||
<paragraph id="8BBD9844FFE45E580FEDF9DFFD340F32" blockId="10.[151,1437,1238,2019]" pageId="10" pageNumber="293">
|
||
<emphasis id="B9764456FFE45E580FEDF9DFFEFC0F1E" bold="true" box="[199,334,1598,1623]" pageId="10" pageNumber="293">Etymology.</emphasis>
|
||
The specific name phuquocensis, noun in apposition, refers to Phu Quoc Island in southwest
|
||
<collectingCountry id="F315D8D4FFE45E580FBDF983FF4B0F32" box="[151,249,1634,1659]" name="Vietnam" pageId="10" pageNumber="293">Vietnam</collectingCountry>
|
||
, where this species was first found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C318CBCFFFE45E590FEDF967FDEA09B1" lastPageId="11" lastPageNumber="294" pageId="10" pageNumber="293" type="diagnosis">
|
||
<paragraph id="8BBD9844FFE45E590FEDF967FDEA09B1" blockId="10.[151,1437,1238,2019]" lastBlockId="11.[151,1436,151,248]" lastPageId="11" lastPageNumber="294" pageId="10" pageNumber="293">
|
||
<emphasis id="B9764456FFE45E580FEDF967FEF30FD6" bold="true" box="[199,321,1670,1695]" pageId="10" pageNumber="293">Diagnosis.</emphasis>
|
||
Colour and trichome patterns of
|
||
<taxonomicName id="4C02E3C7FFE45E580D94F967FC440FD6" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[702,1014,1670,1695]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="phuquocensis" status="sp. nov.">
|
||
<emphasis id="B9764456FFE45E580D94F967FC440FD6" box="[702,1014,1670,1695]" italics="true" pageId="10" pageNumber="293">Monographis phuquocensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE45E580CD5F967FBF20FD6" bold="true" box="[1023,1088,1670,1695]" pageId="10" pageNumber="293">
|
||
<taxonomicNameLabel id="A245F92DFFE45E580CD5F967FBF20FD6" box="[1023,1088,1670,1695]" pageId="10" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
similar in appearance to other described
|
||
<taxonomicName id="4C02E3C7FFE45E580E27F94BFE160F8A" authorityName="Attems" authorityYear="1907" box="[269,420,1706,1731]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9764456FFE45E580E27F94BFE160F8A" box="[269,420,1706,1731]" italics="true" pageId="10" pageNumber="293">Monographis</emphasis>
|
||
</taxonomicName>
|
||
species including
|
||
<taxonomicName id="4C02E3C7FFE45E580DABF94DFCFF0F8A" authorityName="Huynh and Veenstra" authorityYear="2013" box="[641,845,1706,1731]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="queenslandica">
|
||
<emphasis id="B9764456FFE45E580DABF94DFCFF0F8A" box="[641,845,1706,1731]" italics="true" pageId="10" pageNumber="293">M. queenslandica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C02E3C7FFE45E580CA6F94DFBF80F8A" authorityName="Huynh and Veenstra" authorityYear="2015" box="[908,1098,1706,1731]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="dongnaiensis">
|
||
<emphasis id="B9764456FFE45E580CA6F94DFBF80F8A" box="[908,1098,1706,1731]" italics="true" pageId="10" pageNumber="293">M. dongnaiensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF93E5B5FFE45E580B77F94BFA2A0F8A" author="Huynh, C. & Veenstra, A." box="[1117,1432,1706,1731]" pageId="10" pageNumber="293" pagination="573 - 588" refId="ref11479" refString="Huynh, C. & Veenstra, A. (2013) Taxonomy and biology of a new species of Pincushion Millipede of the genus Monographis (Diplopoda: Polyxenida) from Australia. Zootaxa, 3721 (6), 573 - 588. https: // doi. org / 10.11646 / zootaxa. 3721.6.4" type="journal article" year="2013">Huynh and Veenstra, 2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF93E5B5FFE45E580FBDF92FFF660FAE" author="Huynh, C. & Veenstra, A." box="[151,212,1742,1767]" pageId="10" pageNumber="293" pagination="460 - 474" refId="ref11536" refString="Huynh, C. & Veenstra, A. (2015) Description of a new species of Penicillate Millipede from the genus Monographis (Diplopoda: Polyxenidae) found in Vietnam. Zootaxa, 3964 (4), 460 - 474. https: // doi. org / 10.11646 / zootaxa. 3964.4.5" type="journal article" year="2015">2015</bibRefCitation>
|
||
). Characteristics and arrangement of caudal bundles are quite similar among
|
||
<taxonomicName id="4C02E3C7FFE45E580B66F92FFB510FAE" authorityName="Attems" authorityYear="1907" box="[1100,1251,1742,1767]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9764456FFE45E580B66F92FFB510FAE" box="[1100,1251,1742,1767]" italics="true" pageId="10" pageNumber="293">Monographis</emphasis>
|
||
</taxonomicName>
|
||
species (
|
||
<bibRefCitation id="EF93E5B5FFE45E580A64F92FFEC20E42" author="Huynh, C. & Veenstra, A." pageId="10" pageNumber="293" pagination="460 - 474" refId="ref11536" refString="Huynh, C. & Veenstra, A. (2015) Description of a new species of Penicillate Millipede from the genus Monographis (Diplopoda: Polyxenidae) found in Vietnam. Zootaxa, 3964 (4), 460 - 474. https: // doi. org / 10.11646 / zootaxa. 3964.4.5" type="journal article" year="2015">Huynh and Veenstra 2015</bibRefCitation>
|
||
) and this is classified as the caudal bundle
|
||
<typeStatus id="54B926E6FFE45E580C51F912FC190E42" box="[891,939,1779,1803]" pageId="10" pageNumber="293">type</typeStatus>
|
||
II arrangement (Condé and Nguyen Duy- Jacquemin, 2008)Pattern of sensilla on antennal article VII common to all species of
|
||
<taxonomicName id="4C02E3C7FFE45E580B5EF8F7FAB90E66" authorityName="Attems" authorityYear="1907" box="[1140,1291,1814,1839]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9764456FFE45E580B5EF8F7FAB90E66" box="[1140,1291,1814,1839]" italics="true" pageId="10" pageNumber="293">Monographis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF93E5B5FFE45E580A36F8F7FE8E0E1A" author="Huynh, C. & Veenstra, A." pageId="10" pageNumber="293" pagination="573 - 588" refId="ref11479" refString="Huynh, C. & Veenstra, A. (2013) Taxonomy and biology of a new species of Pincushion Millipede of the genus Monographis (Diplopoda: Polyxenida) from Australia. Zootaxa, 3721 (6), 573 - 588. https: // doi. org / 10.11646 / zootaxa. 3721.6.4" type="journal article" year="2013">Huynh and Veenstra 2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF93E5B5FFE45E580E66F8DBFE3A0E1A" author="Huynh, C. & Veenstra, A." box="[332,392,1850,1875]" pageId="10" pageNumber="293" pagination="460 - 474" refId="ref11536" refString="Huynh, C. & Veenstra, A. (2015) Description of a new species of Penicillate Millipede from the genus Monographis (Diplopoda: Polyxenidae) found in Vietnam. Zootaxa, 3964 (4), 460 - 474. https: // doi. org / 10.11646 / zootaxa. 3964.4.5" type="journal article" year="2015">2015</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C02E3C7FFE45E580EB4F8DDFDD20E1A" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[414,608,1850,1875]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="species" species="phuquocensis" status="sp. nov.">
|
||
<emphasis id="B9764456FFE45E580EB4F8DDFDD20E1A" box="[414,608,1850,1875]" italics="true" pageId="10" pageNumber="293">M. phuquocensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE45E580D40F8DBFD1F0E1A" bold="true" box="[618,685,1850,1875]" pageId="10" pageNumber="293">
|
||
<taxonomicNameLabel id="A245F92DFFE45E580D40F8DBFD1F0E1A" box="[618,685,1850,1875]" pageId="10" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
different from other
|
||
<taxonomicName id="4C02E3C7FFE45E580C8DF8DBFB8C0E1A" authorityName="Attems" authorityYear="1907" box="[935,1086,1850,1875]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="10" pageNumber="293" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9764456FFE45E580C8DF8DBFB8C0E1A" box="[935,1086,1850,1875]" italics="true" pageId="10" pageNumber="293">Monographis</emphasis>
|
||
</taxonomicName>
|
||
species in antennal article VI having 15–20 sensilla of various sizes, forming a crescent-shape with the anterior sensilla longer than posterior ones. These sensilla form 4 distinct groups: 3 thick bacilliform sensilla commonly present in a group of 2–4 thin bacilliform sensilla located anteriorly (Group 1), 2–3 medium length thin bacilliform sensilla (Group 2), 1–2 short thin bacilliform sensilla (Group 3), and a group of 6–8 thick short conical sensilla located posteriorly (Group 4). Surface of labrum asetose and smooth, with a row of setae present near anterior edge of lamellae. Telotarsus bearing a posterior process shorter, half the length of claw. Anterior lateral process and lamella process present; setiform process half the length of claw.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7DC8CCFFE55E590FBDFBBFFA8D0D3D" ID-DOI="http://doi.org/10.5281/zenodo.10092680" ID-Zenodo-Dep="10092680" box="[151,1343,1118,1140]" httpUri="https://zenodo.org/record/10092680/files/figure.png" pageId="11" pageNumber="294" startId="11.[151,250,1118,1140]" targetBox="[202,1384,301,1097]" targetPageId="11">
|
||
<paragraph id="8BBD9844FFE55E590FBDFBBFFA8D0D3D" blockId="11.[151,1343,1118,1140]" box="[151,1343,1118,1140]" pageId="11" pageNumber="294">
|
||
<emphasis id="B9764456FFE55E590FBDFBBFFEBF0D3D" bold="true" box="[151,269,1118,1140]" pageId="11" pageNumber="294">FIGURE 8</emphasis>
|
||
. A living specimen of
|
||
<taxonomicName id="4C02E3C7FFE55E590EDEFBBFFCBF0D3D" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[500,781,1118,1140]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="294" phylum="Arthropoda" rank="species" species="phuquocensis" status="sp. nov.">
|
||
<emphasis id="B9764456FFE55E590EDEFBBFFCBF0D3D" box="[500,781,1118,1140]" italics="true" pageId="11" pageNumber="294">Monographis phuquocensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE55E590C39FBBFFCFE0D3D" bold="true" box="[787,844,1118,1140]" pageId="11" pageNumber="294">
|
||
<taxonomicNameLabel id="A245F92DFFE55E590C39FBBFFCFE0D3D" box="[787,844,1118,1140]" pageId="11" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Da Ban stream, Phu Quoc Island, Vietnam.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C318CBCFFFE55E5D0FEDFB7AFD8F0B51" lastPageId="15" lastPageNumber="298" pageId="11" pageNumber="294" type="description">
|
||
<paragraph id="8BBD9844FFE55E590FEDFB7AFC310DB5" blockId="11.[151,1437,1179,2032]" pageId="11" pageNumber="294">
|
||
<emphasis id="B9764456FFE55E590FEDFB7AFEEB0DFD" bold="true" box="[199,345,1179,1204]" pageId="11" pageNumber="294">Description.</emphasis>
|
||
<emphasis id="B9764456FFE55E590E75FB7DFDB30DFA" box="[351,513,1180,1203]" italics="true" pageId="11" pageNumber="294">Measurements</emphasis>
|
||
:
|
||
<typeStatus id="54B926E6FFE55E590D3AFB7AFDCA0DFD" box="[528,632,1179,1204]" pageId="11" pageNumber="294" type="holotype">Holotype</typeStatus>
|
||
male body length 2.0 mm; (
|
||
<typeStatus id="54B926E6FFE55E590C9CFB7DFB970DFD" box="[950,1061,1180,1204]" pageId="11" pageNumber="294" type="paratype">paratypes</typeStatus>
|
||
) males ranging from 1.4–2.0 mm (
|
||
<emphasis id="B9764456FFE55E590F8AFB21FF1C0D9E" box="[160,174,1216,1239]" italics="true" pageId="11" pageNumber="294">n</emphasis>
|
||
= 5) and females (
|
||
<emphasis id="B9764456FFE55E590E55FB21FE3F0D9E" box="[383,397,1216,1239]" italics="true" pageId="11" pageNumber="294">n</emphasis>
|
||
= 8) ranging from
|
||
<quantity id="4CFA35A1FFE55E590D4BFB5EFD580D9E" box="[609,746,1215,1240]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="11" pageNumber="294" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.2–2.5 mm</quantity>
|
||
. Caudal bundle in male slightly narrower in width and shorter (
|
||
<quantity id="4CFA35A1FFE55E590FB4FB02FF430DB5" box="[158,241,1251,1276]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="294" unit="mm" value="0.5">0.5mm</quantity>
|
||
) compared to female, the latter wider and longer (
|
||
<quantity id="4CFA35A1FFE55E590C08FB02FCC50DB5" box="[802,887,1251,1276]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="11" pageNumber="294" unit="mm" value="0.6">0.6mm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE55E590FEDFAE6FF430C0D" blockId="11.[151,1437,1179,2032]" pageId="11" pageNumber="294">
|
||
<emphasis id="B9764456FFE55E590FEDFAE6FEE20C69" box="[199,336,1287,1312]" italics="true" pageId="11" pageNumber="294">Colouration</emphasis>
|
||
: Body yellowish-brown in colour with 2 darker lateral bands, caudal bundle black-silvery colour (
|
||
<figureCitation id="133984C1FFE55E590FB5FACAFF560C0D" box="[159,228,1323,1348]" captionStart="FIGURE 8" captionStartId="11.[151,250,1118,1140]" captionTargetBox="[202,1384,301,1097]" captionTargetId="figure@11.[202,1384,301,1097]" captionTargetPageId="11" captionText="FIGURE 8. A living specimen of Monographis phuquocensis sp. n. from Da Ban stream, Phu Quoc Island, Vietnam." figureDoi="http://doi.org/10.5281/zenodo.10092680" httpUri="https://zenodo.org/record/10092680/files/figure.png" pageId="11" pageNumber="294">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE55E590FEDFAAEFE910FE5" blockId="11.[151,1437,1179,2032]" pageId="11" pageNumber="294">
|
||
<emphasis id="B9764456FFE55E590FEDFAAEFEB60C21" box="[199,260,1359,1384]" italics="true" pageId="11" pageNumber="294">Head</emphasis>
|
||
: Eight ommatidia on each side: 4 dorsal, 4 lateral (1 anterior, 2 medial and 1 posterior). Vertex with 2 posterior trichome groups comprised of 2 rows, with a large gap present between these groups. Anterior, oblique with a variable number of large trichome sockets in centre, gradually reducing in size toward both ends; posterior row with slightly smaller trichome sockets. A narrow space present between anterior and posterior rows.
|
||
<typeStatus id="54B926E6FFE55E590A1EFA5AFA2E0C9D" box="[1332,1436,1467,1492]" pageId="11" pageNumber="294" type="holotype">Holotype</typeStatus>
|
||
male with 11 + 12 trichome sockets in anterior rows and 2 + 2 trichome sockets in posterior rows (
|
||
<figureCitation id="133984C1FFE55E590A1BFA3EFA3D0CB1" box="[1329,1423,1503,1528]" captionStart="FIGURE 9" captionStartId="12.[151,250,1741,1763]" captionTargetBox="[151,1404,201,1696]" captionTargetId="figure@12.[151,1436,193,1719]" captionTargetPageId="12" captionText="FIGURE 9. Holotype male Monographis phuquocensis sp. n., A. Head capsule showing 8 ommatidium on each side, 2 groups of the posterior vertex trichomes (pv) and 3 trichobothria: trichobothrium a, b and c; B. Collum (Co); C. tergite 2 (T2) and D. the last tergite – tergite 10 (T10). E. Right gnathochilarium showing the lateral palp with 11 conical sensilla and the medial palp with 22 conical sensilla (LP: lateral palp, Mp: medial palp). F. Structure of the trichobothria: Trichobothrium a (posterior position), trichobothrium b (lateral position) and trichobothrium c (anterior position). G. Clypeo-labrum showing 10 setae (only 4 setae present and 6 seta sockets) on the posterior margin; a row of 10 slender setae were present on each side of the anterior margin of lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092682" httpUri="https://zenodo.org/record/10092682/files/figure.png" pageId="11" pageNumber="294">Fig. 9A</figureCitation>
|
||
).
|
||
<typeStatus id="54B926E6FFE55E590FBDF9E5FEB60F55" box="[151,260,1540,1564]" pageId="11" pageNumber="294" type="paratype">Paratypes</typeStatus>
|
||
with variation common in this species regardless of sex, ranging from 11–15 (anterior rows) and 2–3 (posterior rows). Trichobothria (trichobothrium
|
||
<emphasis id="B9764456FFE55E590DF8F9C9FD520F76" bold="true" box="[722,736,1576,1599]" italics="true" pageId="11" pageNumber="294">a</emphasis>
|
||
, posterior position, trichobothrium
|
||
<emphasis id="B9764456FFE55E590BBAF9C6FB2C0F09" bold="true" box="[1168,1182,1575,1600]" italics="true" pageId="11" pageNumber="294">b</emphasis>
|
||
, lateral position and trichobothrium
|
||
<emphasis id="B9764456FFE55E590E67F9ADFEEB0F2A" bold="true" box="[333,345,1612,1635]" italics="true" pageId="11" pageNumber="294">c</emphasis>
|
||
, anterior position) typically thin sensory hairs with narrow cylindrical funicles. All these trichobothria (
|
||
<emphasis id="B9764456FFE55E590E12F991FEF40FCE" bold="true" box="[312,326,1648,1671]" italics="true" pageId="11" pageNumber="294">a</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFE55E590E7EF98EFED00FC1" bold="true" box="[340,354,1647,1672]" italics="true" pageId="11" pageNumber="294">b</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE55E590EB2F991FE160FCE" bold="true" box="[408,420,1648,1671]" italics="true" pageId="11" pageNumber="294">c</emphasis>
|
||
) equal in socket size and form an isosceles triangle with an equal distance between
|
||
<emphasis id="B9764456FFE55E590A7AF98EFADE0FC1" bold="true" box="[1360,1388,1647,1672]" italics="true" pageId="11" pageNumber="294">ab</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE55E590FBDF972FF000FE5" bold="true" box="[151,178,1683,1708]" italics="true" pageId="11" pageNumber="294">bc</emphasis>
|
||
(
|
||
<figureCitation id="133984C1FFE55E590FEBF972FEA40FE5" box="[193,278,1683,1708]" captionStart="FIGURE 9" captionStartId="12.[151,250,1741,1763]" captionTargetBox="[151,1404,201,1696]" captionTargetId="figure@12.[151,1436,193,1719]" captionTargetPageId="12" captionText="FIGURE 9. Holotype male Monographis phuquocensis sp. n., A. Head capsule showing 8 ommatidium on each side, 2 groups of the posterior vertex trichomes (pv) and 3 trichobothria: trichobothrium a, b and c; B. Collum (Co); C. tergite 2 (T2) and D. the last tergite – tergite 10 (T10). E. Right gnathochilarium showing the lateral palp with 11 conical sensilla and the medial palp with 22 conical sensilla (LP: lateral palp, Mp: medial palp). F. Structure of the trichobothria: Trichobothrium a (posterior position), trichobothrium b (lateral position) and trichobothrium c (anterior position). G. Clypeo-labrum showing 10 setae (only 4 setae present and 6 seta sockets) on the posterior margin; a row of 10 slender setae were present on each side of the anterior margin of lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092682" httpUri="https://zenodo.org/record/10092682/files/figure.png" pageId="11" pageNumber="294">Fig. 9F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE55E590FEDF959FA940EB9" blockId="11.[151,1437,1179,2032]" pageId="11" pageNumber="294">
|
||
<emphasis id="B9764456FFE55E590FEDF959FE830F86" box="[199,305,1720,1743]" italics="true" pageId="11" pageNumber="294">Antennae</emphasis>
|
||
: Eight articles and 4 sensory cones present, characteristics typical of
|
||
<taxonomicName id="4C02E3C7FFE55E590B66F956FB680F99" authorityName="Lucas" authorityYear="1840" box="[1100,1242,1719,1744]" class="Diplopoda" family="Polyxenidae" kingdom="Animalia" order="Polyxenida" pageId="11" pageNumber="294" phylum="Arthropoda" rank="family">Polyxenidae</taxonomicName>
|
||
. Antennal article VI (
|
||
<figureCitation id="133984C1FFE55E590FEDF93AFE810FBD" box="[199,307,1755,1780]" captionStart="FIGURE 10" captionStartId="13.[151,250,1219,1241]" captionTargetBox="[177,1414,199,1186]" captionTargetId="figure@13.[151,1436,193,1198]" captionTargetPageId="13" captionText="FIGURE 10. Holotype Monographis phuquocensis sp. n., A. The antennal articles V, VI, VII and VIII. B. Arrangement of sensilla on the antennal article VII: A conical sensillum (c) located posteriorly, 2 thick bacilliform sensilla (Tbs) and a setiform sensillum (s). C. Sensilla arrangement on the antennal article VI with 4 groups of sensilla of various lengths: Group 1: 7 bacilliform sensilla, which consists of 3 thick bacilliform sensilla distributed between 1 or 2 thin bacilliform sensilla (Tbs a: Thick bacilliform sensilla located in the anterior position, Tbs i: the thick bacilliform sensilla located in an intermedium position, Tbs p: the thick bacilliform sensilla located in the posterior position); followed by a conical sensillum (c); Group 2: 3 medium length thin bacilliform sensilla (tbs m); Group 3: 2 small thin short bacilliform sensilla (tbs s); Group 4: 8 short thick conical sensilla (Tsc) located posteriorly." figureDoi="http://doi.org/10.5281/zenodo.10092684" httpUri="https://zenodo.org/record/10092684/files/figure.png" pageId="11" pageNumber="294">Figs 10A</figureCitation>
|
||
and
|
||
<figureCitation id="133984C1FFE55E590E58F93DFE130FBD" box="[370,417,1756,1780]" captionStart="FIGURE 11" captionStartId="14.[151,250,1329,1351]" captionTargetBox="[151,1436,301,1308]" captionTargetId="figure@14.[151,1436,301,1308]" captionTargetPageId="14" captionText="FIGURE 11. Scanning Electron Microscope images of antennal articles and the labrum of Monographis phuquocensis sp. n.. A. The antennal articles VI, VII and VIII. B. Sensilla arrangement on the antennal article VI showing the anterior group of 7 bacilliform sensilla which includes 4 thin bacilliform sensilla (tbs) and 3 thicker ones (Tbs a, Tbs i, Tbs p, tbs m and tbs s: refer to Figure 10). C. The arrangement of sensilla on the antennal article VI showing 2 small thin short sensilla (tbs s) and the posterior group of 8 thick short conical sensilla (Tsc). D. Clypeo-labrum: lateral lamella (Ll) and lamella (l), posterior row of setae (s) and the anterior row of slender setae (sa)." figureDoi="http://doi.org/10.5281/zenodo.10092686" httpUri="https://zenodo.org/record/10092686/files/figure.png" pageId="11" pageNumber="294">11A</figureCitation>
|
||
) with 18–20 sensilla.
|
||
<typeStatus id="54B926E6FFE55E590D82F93AFCA20FBD" box="[680,784,1755,1780]" pageId="11" pageNumber="294" type="holotype">Holotype</typeStatus>
|
||
with 20 sensilla forming a crescent-shape with anterior sensilla longer than posterior ones. These sensilla forming 4 groups: Group 1 located anteriorly with 3 thick bacilliform sensilla (Tbs
|
||
<emphasis id="B9764456FFE55E590E96F8C2FE780E75" bold="true" box="[444,458,1827,1852]" pageId="11" pageNumber="294">a</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFE55E590EF7F8C2FE570E75" bold="true" box="[477,485,1827,1852]" pageId="11" pageNumber="294">i</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE55E590D0EF8C2FD810E75" bold="true" box="[548,563,1827,1852]" pageId="11" pageNumber="294">p</emphasis>
|
||
) among 4 thin bacilliform sensilla (tbs), this group followed by a conical sensillum (c); Group 2 with 3 medium length thin bacilliform sensilla (tbs
|
||
<emphasis id="B9764456FFE55E590B0EF8A6FB890E29" bold="true" box="[1060,1083,1863,1888]" pageId="11" pageNumber="294">m</emphasis>
|
||
); Group 3 with 2 short, thin bacilliform sensilla (tbs
|
||
<emphasis id="B9764456FFE55E590E8EF88AFE1D0ECD" bold="true" box="[420,431,1899,1924]" pageId="11" pageNumber="294">s</emphasis>
|
||
); a small gap present between Groups 2 and 3; last group has 8 thick short conical sensilla (Ts
|
||
<emphasis id="B9764456FFE55E590F90F86EFF740EE1" bold="true" box="[186,198,1935,1960]" pageId="11" pageNumber="294">c</emphasis>
|
||
) located posteriorly (
|
||
<figureCitation id="133984C1FFE55E590E9EF86EFD8A0EE1" box="[436,568,1935,1960]" captionStart="FIGURE 10" captionStartId="13.[151,250,1219,1241]" captionTargetBox="[177,1414,199,1186]" captionTargetId="figure@13.[151,1436,193,1198]" captionTargetPageId="13" captionText="FIGURE 10. Holotype Monographis phuquocensis sp. n., A. The antennal articles V, VI, VII and VIII. B. Arrangement of sensilla on the antennal article VII: A conical sensillum (c) located posteriorly, 2 thick bacilliform sensilla (Tbs) and a setiform sensillum (s). C. Sensilla arrangement on the antennal article VI with 4 groups of sensilla of various lengths: Group 1: 7 bacilliform sensilla, which consists of 3 thick bacilliform sensilla distributed between 1 or 2 thin bacilliform sensilla (Tbs a: Thick bacilliform sensilla located in the anterior position, Tbs i: the thick bacilliform sensilla located in an intermedium position, Tbs p: the thick bacilliform sensilla located in the posterior position); followed by a conical sensillum (c); Group 2: 3 medium length thin bacilliform sensilla (tbs m); Group 3: 2 small thin short bacilliform sensilla (tbs s); Group 4: 8 short thick conical sensilla (Tsc) located posteriorly." figureDoi="http://doi.org/10.5281/zenodo.10092684" httpUri="https://zenodo.org/record/10092684/files/figure.png" pageId="11" pageNumber="294">Figure 10C</figureCitation>
|
||
). Different arrangements of sensilla apparent in scanning electron microscope images of other specimens (
|
||
<figureCitation id="133984C1FFE55E590EFBF852FD150E85" box="[465,679,1971,1996]" captionStart="FIGURE 11" captionStartId="14.[151,250,1329,1351]" captionTargetBox="[151,1436,301,1308]" captionTargetId="figure@14.[151,1436,301,1308]" captionTargetPageId="14" captionText="FIGURE 11. Scanning Electron Microscope images of antennal articles and the labrum of Monographis phuquocensis sp. n.. A. The antennal articles VI, VII and VIII. B. Sensilla arrangement on the antennal article VI showing the anterior group of 7 bacilliform sensilla which includes 4 thin bacilliform sensilla (tbs) and 3 thicker ones (Tbs a, Tbs i, Tbs p, tbs m and tbs s: refer to Figure 10). C. The arrangement of sensilla on the antennal article VI showing 2 small thin short sensilla (tbs s) and the posterior group of 8 thick short conical sensilla (Tsc). D. Clypeo-labrum: lateral lamella (Ll) and lamella (l), posterior row of setae (s) and the anterior row of slender setae (sa)." figureDoi="http://doi.org/10.5281/zenodo.10092686" httpUri="https://zenodo.org/record/10092686/files/figure.png" pageId="11" pageNumber="294">Figs 11A, B and C</figureCitation>
|
||
). Antennal article VII with 2 thick bacilliform sensilla (Tbs) with a setiform sensillum (s) between them and a conical sensillum (c) located posteriorly (
|
||
<figureCitation id="133984C1FFE55E590B63F836FB020EB9" box="[1097,1200,2007,2032]" captionStart="FIGURE 10" captionStartId="13.[151,250,1219,1241]" captionTargetBox="[177,1414,199,1186]" captionTargetId="figure@13.[151,1436,193,1198]" captionTargetPageId="13" captionText="FIGURE 10. Holotype Monographis phuquocensis sp. n., A. The antennal articles V, VI, VII and VIII. B. Arrangement of sensilla on the antennal article VII: A conical sensillum (c) located posteriorly, 2 thick bacilliform sensilla (Tbs) and a setiform sensillum (s). C. Sensilla arrangement on the antennal article VI with 4 groups of sensilla of various lengths: Group 1: 7 bacilliform sensilla, which consists of 3 thick bacilliform sensilla distributed between 1 or 2 thin bacilliform sensilla (Tbs a: Thick bacilliform sensilla located in the anterior position, Tbs i: the thick bacilliform sensilla located in an intermedium position, Tbs p: the thick bacilliform sensilla located in the posterior position); followed by a conical sensillum (c); Group 2: 3 medium length thin bacilliform sensilla (tbs m); Group 3: 2 small thin short bacilliform sensilla (tbs s); Group 4: 8 short thick conical sensilla (Tsc) located posteriorly." figureDoi="http://doi.org/10.5281/zenodo.10092684" httpUri="https://zenodo.org/record/10092684/files/figure.png" pageId="11" pageNumber="294">Figs 10B</figureCitation>
|
||
and
|
||
<figureCitation id="133984C1FFE55E590BCCF839FAAB0EB9" box="[1254,1305,2008,2032]" captionStart="FIGURE 11" captionStartId="14.[151,250,1329,1351]" captionTargetBox="[151,1436,301,1308]" captionTargetId="figure@14.[151,1436,301,1308]" captionTargetPageId="14" captionText="FIGURE 11. Scanning Electron Microscope images of antennal articles and the labrum of Monographis phuquocensis sp. n.. A. The antennal articles VI, VII and VIII. B. Sensilla arrangement on the antennal article VI showing the anterior group of 7 bacilliform sensilla which includes 4 thin bacilliform sensilla (tbs) and 3 thicker ones (Tbs a, Tbs i, Tbs p, tbs m and tbs s: refer to Figure 10). C. The arrangement of sensilla on the antennal article VI showing 2 small thin short sensilla (tbs s) and the posterior group of 8 thick short conical sensilla (Tsc). D. Clypeo-labrum: lateral lamella (Ll) and lamella (l), posterior row of setae (s) and the anterior row of slender setae (sa)." figureDoi="http://doi.org/10.5281/zenodo.10092686" httpUri="https://zenodo.org/record/10092686/files/figure.png" pageId="11" pageNumber="294">11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7DC8CCFFE25E5E0FBDF92CFEEF0ED2" ID-DOI="http://doi.org/10.5281/zenodo.10092682" ID-Zenodo-Dep="10092682" httpUri="https://zenodo.org/record/10092682/files/figure.png" pageId="12" pageNumber="295" startId="12.[151,250,1741,1763]" targetBox="[151,1404,201,1696]" targetPageId="12">
|
||
<paragraph id="8BBD9844FFE25E5E0FBDF92CFEEF0ED2" blockId="12.[151,1436,1741,1947]" pageId="12" pageNumber="295">
|
||
<emphasis id="B9764456FFE25E5E0FBDF92CFEBE0FAA" bold="true" box="[151,268,1741,1763]" pageId="12" pageNumber="295">FIGURE 9</emphasis>
|
||
. Holotype male
|
||
<taxonomicName id="4C02E3C7FFE25E5E0E9EF92CFD7F0FAA" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[436,717,1741,1763]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="12" pageNumber="295" phylum="Arthropoda" rank="species" species="phuquocensis" status="sp. nov.">
|
||
<emphasis id="B9764456FFE25E5E0E9EF92CFD7F0FAA" box="[436,717,1741,1763]" italics="true" pageId="12" pageNumber="295">Monographis phuquocensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE25E5E0DF9F92CFCBE0FAA" bold="true" box="[723,780,1741,1763]" pageId="12" pageNumber="295">
|
||
<taxonomicNameLabel id="A245F92DFFE25E5E0DF9F92CFCBE0FAA" box="[723,780,1741,1763]" pageId="12" pageNumber="295" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFE25E5E0C32F92CFC980FAA" bold="true" box="[792,810,1741,1763]" pageId="12" pageNumber="295">A</emphasis>
|
||
. Head capsule showing 8 ommatidium on each side, 2 groups of the posterior vertex trichomes (pv) and 3 trichobothria: trichobothrium
|
||
<emphasis id="B9764456FFE25E5E0CA0F90DFC250E48" bold="true" box="[906,919,1772,1793]" italics="true" pageId="12" pageNumber="295">a</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFE25E5E0C8EF90AFC030E48" bold="true" box="[932,945,1771,1793]" italics="true" pageId="12" pageNumber="295">b</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE25E5E0CC9F90DFC5C0E48" bold="true" box="[995,1006,1772,1793]" italics="true" pageId="12" pageNumber="295">c</emphasis>
|
||
;
|
||
<emphasis id="B9764456FFE25E5E0CD6F90DFBA10E4B" bold="true" box="[1020,1043,1772,1794]" pageId="12" pageNumber="295">B.</emphasis>
|
||
Collum (Co);
|
||
<emphasis id="B9764456FFE25E5E0B80F90DFB700E4B" bold="true" box="[1194,1218,1772,1794]" pageId="12" pageNumber="295">C.</emphasis>
|
||
tergite 2 (T2) and
|
||
<emphasis id="B9764456FFE25E5E0AA9F90DFA2E0E4B" bold="true" box="[1411,1436,1772,1794]" pageId="12" pageNumber="295">D.</emphasis>
|
||
the last tergite – tergite 10 (T10).
|
||
<emphasis id="B9764456FFE25E5E0EC2F8EBFE4B0E69" bold="true" box="[488,505,1802,1824]" pageId="12" pageNumber="295">E</emphasis>
|
||
. Right gnathochilarium showing the lateral palp with 11 conical sensilla and the medial palp with 22 conical sensilla (LP: lateral palp, Mp: medial palp).
|
||
<emphasis id="B9764456FFE25E5E0C31F8C8FC9A0E76" bold="true" box="[795,808,1833,1855]" pageId="12" pageNumber="295">F</emphasis>
|
||
. Structure of the trichobothria: Trichobothrium
|
||
<emphasis id="B9764456FFE25E5E0A09F8C8FA820E77" bold="true" box="[1315,1328,1833,1854]" italics="true" pageId="12" pageNumber="295">a</emphasis>
|
||
(posterior position), trichobothrium
|
||
<emphasis id="B9764456FFE25E5E0EB1F8A6FE1A0E14" bold="true" box="[411,424,1863,1885]" italics="true" pageId="12" pageNumber="295">b</emphasis>
|
||
(lateral position) and trichobothrium
|
||
<emphasis id="B9764456FFE25E5E0C09F8A9FC9C0E14" bold="true" box="[803,814,1864,1885]" italics="true" pageId="12" pageNumber="295">c</emphasis>
|
||
(anterior position).
|
||
<emphasis id="B9764456FFE25E5E0CDDF8A6FBBA0E14" bold="true" box="[1015,1032,1863,1885]" pageId="12" pageNumber="295">G</emphasis>
|
||
. Clypeo-labrum showing 10 setae (only 4 setae present and 6 seta sockets) on the posterior margin; a row of 10 slender setae were present on each side of the anterior margin of lamellae.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7DC8CCFFE35E5F0FBDFB22FD870CF9" ID-DOI="http://doi.org/10.5281/zenodo.10092684" ID-Zenodo-Dep="10092684" httpUri="https://zenodo.org/record/10092684/files/figure.png" pageId="13" pageNumber="296" startId="13.[151,250,1219,1241]" targetBox="[177,1414,199,1186]" targetPageId="13">
|
||
<paragraph id="8BBD9844FFE35E5F0FBDFB22FD870CF9" blockId="13.[151,1436,1219,1456]" pageId="13" pageNumber="296">
|
||
<emphasis id="B9764456FFE35E5F0FBDFB22FEA90D90" bold="true" box="[151,283,1219,1241]" pageId="13" pageNumber="296">FIGURE 10</emphasis>
|
||
. Holotype
|
||
<taxonomicName id="4C02E3C7FFE35E5F0EBBFB22FD1E0D90" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[401,684,1219,1241]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="13" pageNumber="296" phylum="Arthropoda" rank="species" species="phuquocensis" status="sp. nov.">
|
||
<emphasis id="B9764456FFE35E5F0EBBFB22FD1E0D90" box="[401,684,1219,1241]" italics="true" pageId="13" pageNumber="296">Monographis phuquocensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE35E5F0D9EFB22FD5D0D90" bold="true" box="[692,751,1219,1241]" pageId="13" pageNumber="296">
|
||
<taxonomicNameLabel id="A245F92DFFE35E5F0D9EFB22FD5D0D90" box="[692,751,1219,1241]" pageId="13" pageNumber="296" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFE35E5F0DD4FB22FCA20D90" bold="true" box="[766,784,1219,1241]" pageId="13" pageNumber="296">A</emphasis>
|
||
. The antennal articles V, VI, VII and VIII.
|
||
<emphasis id="B9764456FFE35E5F0BF3FB22FB580D90" bold="true" box="[1241,1258,1219,1241]" pageId="13" pageNumber="296">B</emphasis>
|
||
. Arrangement of sensilla on the antennal article VII: A conical sensillum (c) located posteriorly, 2 thick bacilliform sensilla (Tbs) and a setiform sensillum (s).
|
||
<emphasis id="B9764456FFE35E5F0E06FAE1FE8C0C5F" bold="true" box="[300,318,1280,1302]" pageId="13" pageNumber="296">C</emphasis>
|
||
. Sensilla arrangement on the antennal article VI with 4 groups of sensilla of various lengths: Group 1: 7 bacilliform sensilla, which consists of 3 thick bacilliform sensilla distributed between 1 or 2 thin bacilliform sensilla (Tbs
|
||
<emphasis id="B9764456FFE35E5F0AA2FAFEFA270C7C" bold="true" box="[1416,1429,1311,1333]" pageId="13" pageNumber="296">a</emphasis>
|
||
: Thick bacilliform sensilla located in the anterior position, Tbs
|
||
<emphasis id="B9764456FFE35E5F0C1FFADFFC8E0C1D" bold="true" box="[821,828,1342,1364]" pageId="13" pageNumber="296">i</emphasis>
|
||
: the thick bacilliform sensilla located in an intermedium position, Tbs
|
||
<emphasis id="B9764456FFE35E5F0E08FABDFE820C3B" bold="true" box="[290,304,1372,1394]" pageId="13" pageNumber="296">p</emphasis>
|
||
: the thick bacilliform sensilla located in the posterior position); followed by a conical sensillum (c); Group 2: 3 medium length thin bacilliform sensilla (tbs
|
||
<emphasis id="B9764456FFE35E5F0D76FA9AFDC30CD8" bold="true" box="[604,625,1403,1425]" pageId="13" pageNumber="296">m</emphasis>
|
||
); Group 3: 2 small thin short bacilliform sensilla (tbs
|
||
<emphasis id="B9764456FFE35E5F0BB0FA9AFB160CD8" bold="true" box="[1178,1188,1403,1425]" pageId="13" pageNumber="296">s</emphasis>
|
||
); Group 4: 8 short thick conical sensilla (Ts
|
||
<emphasis id="B9764456FFE35E5F0E72FA7BFED10CF9" bold="true" box="[344,355,1434,1456]" pageId="13" pageNumber="296">c</emphasis>
|
||
) located posteriorly.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD9844FFE35E5F0FEDFA3DFDBA0FE0" blockId="13.[151,1436,1500,2029]" pageId="13" pageNumber="296">
|
||
<emphasis id="B9764456FFE35E5F0FEDFA3DFEC30CBC" box="[199,369,1500,1525]" italics="true" pageId="13" pageNumber="296">Clypeo-labrum</emphasis>
|
||
:
|
||
<typeStatus id="54B926E6FFE35E5F0EAAFA3DFE5A0CBC" box="[384,488,1500,1525]" pageId="13" pageNumber="296" type="holotype">Holotype</typeStatus>
|
||
with 10 setae present on posterior margin, these setae shorter than half the width of labrum. Setae on
|
||
<typeStatus id="54B926E6FFE35E5F0E73F9E0FE760F50" box="[345,452,1537,1561]" pageId="13" pageNumber="296" type="paratype">paratypes</typeStatus>
|
||
ranging from
|
||
<quantity id="4CFA35A1FFE35E5F0D49F9E0FD040F50" box="[611,694,1536,1561]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="13" pageNumber="296" unit="in" value="9.5" valueMax="11.0" valueMin="8.0">8–11 in</quantity>
|
||
both sexes. Labral surface smooth and asetose. Anterior margin of labrum with a lateral lamella and single broad lamella on each side of median cleft. A row of minute backwardpointing setae present along posterior margin of labrum, another row of 10 slender setae present on each side of anterior margin of lamellae. Number of these anterior slender setae varying among
|
||
<typeStatus id="54B926E6FFE35E5F0B67F98CFB090FCC" box="[1101,1211,1645,1669]" pageId="13" pageNumber="296" type="paratype">paratypes</typeStatus>
|
||
, ranging from 8–12 on each side. (
|
||
<figureCitation id="133984C1FFE35E5F0E1DF971FE200FE0" box="[311,402,1680,1705]" captionStart="FIGURE 9" captionStartId="12.[151,250,1741,1763]" captionTargetBox="[151,1404,201,1696]" captionTargetId="figure@12.[151,1436,193,1719]" captionTargetPageId="12" captionText="FIGURE 9. Holotype male Monographis phuquocensis sp. n., A. Head capsule showing 8 ommatidium on each side, 2 groups of the posterior vertex trichomes (pv) and 3 trichobothria: trichobothrium a, b and c; B. Collum (Co); C. tergite 2 (T2) and D. the last tergite – tergite 10 (T10). E. Right gnathochilarium showing the lateral palp with 11 conical sensilla and the medial palp with 22 conical sensilla (LP: lateral palp, Mp: medial palp). F. Structure of the trichobothria: Trichobothrium a (posterior position), trichobothrium b (lateral position) and trichobothrium c (anterior position). G. Clypeo-labrum showing 10 setae (only 4 setae present and 6 seta sockets) on the posterior margin; a row of 10 slender setae were present on each side of the anterior margin of lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092682" httpUri="https://zenodo.org/record/10092682/files/figure.png" pageId="13" pageNumber="296">Figs 9G</figureCitation>
|
||
and
|
||
<figureCitation id="133984C1FFE35E5F0EE2F970FE490FE0" box="[456,507,1681,1705]" captionStart="FIGURE 11" captionStartId="14.[151,250,1329,1351]" captionTargetBox="[151,1436,301,1308]" captionTargetId="figure@14.[151,1436,301,1308]" captionTargetPageId="14" captionText="FIGURE 11. Scanning Electron Microscope images of antennal articles and the labrum of Monographis phuquocensis sp. n.. A. The antennal articles VI, VII and VIII. B. Sensilla arrangement on the antennal article VI showing the anterior group of 7 bacilliform sensilla which includes 4 thin bacilliform sensilla (tbs) and 3 thicker ones (Tbs a, Tbs i, Tbs p, tbs m and tbs s: refer to Figure 10). C. The arrangement of sensilla on the antennal article VI showing 2 small thin short sensilla (tbs s) and the posterior group of 8 thick short conical sensilla (Tsc). D. Clypeo-labrum: lateral lamella (Ll) and lamella (l), posterior row of setae (s) and the anterior row of slender setae (sa)." figureDoi="http://doi.org/10.5281/zenodo.10092686" httpUri="https://zenodo.org/record/10092686/files/figure.png" pageId="13" pageNumber="296">11D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE35E5F0FEDF955FC310FB8" blockId="13.[151,1436,1500,2029]" pageId="13" pageNumber="296">
|
||
<emphasis id="B9764456FFE35E5F0FEDF955FD860F84" box="[199,564,1716,1741]" italics="true" pageId="13" pageNumber="296">Lateral palp of gnathochilarium</emphasis>
|
||
: 1.25 times as long as medial palp. 11 conical sensilla on lateral palp and 22 on medial palp observed on both
|
||
<typeStatus id="54B926E6FFE35E5F0D26F939FDDC0FB8" box="[524,622,1752,1777]" pageId="13" pageNumber="296" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54B926E6FFE35E5F0D8FF938FCA20FB8" box="[677,784,1753,1777]" pageId="13" pageNumber="296" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="133984C1FFE35E5F0C35F939FCC40FB8" box="[799,886,1752,1777]" captionStart="FIGURE 9" captionStartId="12.[151,250,1741,1763]" captionTargetBox="[151,1404,201,1696]" captionTargetId="figure@12.[151,1436,193,1719]" captionTargetPageId="12" captionText="FIGURE 9. Holotype male Monographis phuquocensis sp. n., A. Head capsule showing 8 ommatidium on each side, 2 groups of the posterior vertex trichomes (pv) and 3 trichobothria: trichobothrium a, b and c; B. Collum (Co); C. tergite 2 (T2) and D. the last tergite – tergite 10 (T10). E. Right gnathochilarium showing the lateral palp with 11 conical sensilla and the medial palp with 22 conical sensilla (LP: lateral palp, Mp: medial palp). F. Structure of the trichobothria: Trichobothrium a (posterior position), trichobothrium b (lateral position) and trichobothrium c (anterior position). G. Clypeo-labrum showing 10 setae (only 4 setae present and 6 seta sockets) on the posterior margin; a row of 10 slender setae were present on each side of the anterior margin of lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092682" httpUri="https://zenodo.org/record/10092682/files/figure.png" pageId="13" pageNumber="296">Fig. 9E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE35E5C0FEDF91DFB4409B1" blockId="13.[151,1436,1500,2029]" lastBlockId="14.[151,1436,151,248]" lastPageId="14" lastPageNumber="297" pageId="13" pageNumber="296">
|
||
<emphasis id="B9764456FFE35E5F0FEDF91DFEBA0E5C" box="[199,264,1788,1813]" italics="true" pageId="13" pageNumber="296">Trunk</emphasis>
|
||
: Body with 10 segments, 9 pleural projections, excluding telson and caudal bundle; 13 pairs of legs. Collum with lateral protuberances bearing 4–5 trichomes on each side. All other tergites, from 2 to 10, each with a pair of pleural projections located antero-laterally. Tergal trichome socket arrangements typically forming 2 broad oval shapes, slightly enlarged laterally, with posterior row curved slightly upward toward centre with a large gap between these trichome socket rows. Collum with 38 (L) and 38 (R) trichome sockets and 4 (L) and 4 (R) trichome sockets on lateral protuberance of the
|
||
<typeStatus id="54B926E6FFE35E5F0D6EF851FD140E80" box="[580,678,1968,1993]" pageId="13" pageNumber="296" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="133984C1FFE35E5F0D9FF851FCBC0E80" box="[693,782,1968,1993]" captionStart="FIGURE 9" captionStartId="12.[151,250,1741,1763]" captionTargetBox="[151,1404,201,1696]" captionTargetId="figure@12.[151,1436,193,1719]" captionTargetPageId="12" captionText="FIGURE 9. Holotype male Monographis phuquocensis sp. n., A. Head capsule showing 8 ommatidium on each side, 2 groups of the posterior vertex trichomes (pv) and 3 trichobothria: trichobothrium a, b and c; B. Collum (Co); C. tergite 2 (T2) and D. the last tergite – tergite 10 (T10). E. Right gnathochilarium showing the lateral palp with 11 conical sensilla and the medial palp with 22 conical sensilla (LP: lateral palp, Mp: medial palp). F. Structure of the trichobothria: Trichobothrium a (posterior position), trichobothrium b (lateral position) and trichobothrium c (anterior position). G. Clypeo-labrum showing 10 setae (only 4 setae present and 6 seta sockets) on the posterior margin; a row of 10 slender setae were present on each side of the anterior margin of lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092682" httpUri="https://zenodo.org/record/10092682/files/figure.png" pageId="13" pageNumber="296">Fig. 9B</figureCitation>
|
||
). Number of trichome sockets varied in
|
||
<typeStatus id="54B926E6FFE35E5F0BFEF850FAF20E80" box="[1236,1344,1969,1993]" pageId="13" pageNumber="296" type="paratype">paratypes</typeStatus>
|
||
ranging from 36–54 and number of lateral protuberances trichome sockets ranging from 4–6. Tergite 2 with a similar structural pattern with posterior row slightly longer and 39 (L) and 36 (R) trichome sockets (
|
||
<figureCitation id="133984C1FFE05E5C0B84FF76FAB509F9" box="[1198,1287,151,176]" captionStart="FIGURE 9" captionStartId="12.[151,250,1741,1763]" captionTargetBox="[151,1404,201,1696]" captionTargetId="figure@12.[151,1436,193,1719]" captionTargetPageId="12" captionText="FIGURE 9. Holotype male Monographis phuquocensis sp. n., A. Head capsule showing 8 ommatidium on each side, 2 groups of the posterior vertex trichomes (pv) and 3 trichobothria: trichobothrium a, b and c; B. Collum (Co); C. tergite 2 (T2) and D. the last tergite – tergite 10 (T10). E. Right gnathochilarium showing the lateral palp with 11 conical sensilla and the medial palp with 22 conical sensilla (LP: lateral palp, Mp: medial palp). F. Structure of the trichobothria: Trichobothrium a (posterior position), trichobothrium b (lateral position) and trichobothrium c (anterior position). G. Clypeo-labrum showing 10 setae (only 4 setae present and 6 seta sockets) on the posterior margin; a row of 10 slender setae were present on each side of the anterior margin of lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092682" httpUri="https://zenodo.org/record/10092682/files/figure.png" pageId="14" pageNumber="297">Fig. 9C</figureCitation>
|
||
). For tergites 3–9, patterns being similar with characteristic large gaps. Tergite 10 with trichome sockets being smaller and denser, a narrow space present between lateral rosette trichome sockets and posterior row (
|
||
<figureCitation id="133984C1FFE05E5C0BA5FF3EFB5809B1" box="[1167,1258,223,248]" captionStart="FIGURE 9" captionStartId="12.[151,250,1741,1763]" captionTargetBox="[151,1404,201,1696]" captionTargetId="figure@12.[151,1436,193,1719]" captionTargetPageId="12" captionText="FIGURE 9. Holotype male Monographis phuquocensis sp. n., A. Head capsule showing 8 ommatidium on each side, 2 groups of the posterior vertex trichomes (pv) and 3 trichobothria: trichobothrium a, b and c; B. Collum (Co); C. tergite 2 (T2) and D. the last tergite – tergite 10 (T10). E. Right gnathochilarium showing the lateral palp with 11 conical sensilla and the medial palp with 22 conical sensilla (LP: lateral palp, Mp: medial palp). F. Structure of the trichobothria: Trichobothrium a (posterior position), trichobothrium b (lateral position) and trichobothrium c (anterior position). G. Clypeo-labrum showing 10 setae (only 4 setae present and 6 seta sockets) on the posterior margin; a row of 10 slender setae were present on each side of the anterior margin of lamellae." figureDoi="http://doi.org/10.5281/zenodo.10092682" httpUri="https://zenodo.org/record/10092682/files/figure.png" pageId="14" pageNumber="297">Fig. 9D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF7DC8CCFFE05E5C0FBDFAD0FD260CA9" ID-DOI="http://doi.org/10.5281/zenodo.10092686" ID-Zenodo-Dep="10092686" httpUri="https://zenodo.org/record/10092686/files/figure.png" pageId="14" pageNumber="297" startId="14.[151,250,1329,1351]" targetBox="[151,1436,301,1308]" targetPageId="14">
|
||
<paragraph id="8BBD9844FFE05E5C0FBDFAD0FD260CA9" blockId="14.[151,1436,1329,1504]" pageId="14" pageNumber="297">
|
||
<emphasis id="B9764456FFE05E5C0FBDFAD0FEAA0C0E" bold="true" box="[151,280,1329,1351]" pageId="14" pageNumber="297">FIGURE 11</emphasis>
|
||
. Scanning Electron Microscope images of antennal articles and the labrum of
|
||
<taxonomicName id="4C02E3C7FFE05E5C0B11FAD0FAE70C0E" authority="Huynh, 2018" authorityName="Huynh" authorityYear="2018" box="[1083,1365,1329,1351]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="14" pageNumber="297" phylum="Arthropoda" rank="species" species="phuquocensis" status="sp. nov.">
|
||
<emphasis id="B9764456FFE05E5C0B11FAD0FAE70C0E" box="[1083,1365,1329,1351]" italics="true" pageId="14" pageNumber="297">Monographis phuquocensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9764456FFE05E5C0A76FAD0FA240C0E" bold="true" box="[1372,1430,1329,1351]" pageId="14" pageNumber="297">
|
||
<taxonomicNameLabel id="A245F92DFFE05E5C0A76FAD0FA240C0E" box="[1372,1430,1329,1351]" pageId="14" pageNumber="297" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="B9764456FFE05E5C0FBDFAB1FF1B0C2F" bold="true" box="[151,169,1360,1382]" pageId="14" pageNumber="297">A</emphasis>
|
||
. The antennal articles VI, VII and VIII.
|
||
<emphasis id="B9764456FFE05E5C0D66FAB1FDEF0C2F" bold="true" box="[588,605,1360,1382]" pageId="14" pageNumber="297">B</emphasis>
|
||
. Sensilla arrangement on the antennal article VI showing the anterior group of 7 bacilliform sensilla which includes 4 thin bacilliform sensilla (tbs) and 3 thicker ones (Tbs
|
||
<emphasis id="B9764456FFE05E5C0B01FA8FFB8A0CCD" bold="true" box="[1067,1080,1390,1412]" pageId="14" pageNumber="297">a</emphasis>
|
||
, Tbs
|
||
<emphasis id="B9764456FFE05E5C0B5AFA8FFBC50CCD" bold="true" box="[1136,1143,1390,1412]" pageId="14" pageNumber="297">i</emphasis>
|
||
, Tbs
|
||
<emphasis id="B9764456FFE05E5C0B85FA8FFB0F0CCD" bold="true" box="[1199,1213,1390,1412]" pageId="14" pageNumber="297">p</emphasis>
|
||
, tbs
|
||
<emphasis id="B9764456FFE05E5C0BC6FA8FFAB30CCD" bold="true" box="[1260,1281,1390,1412]" pageId="14" pageNumber="297">m</emphasis>
|
||
and tbs
|
||
<emphasis id="B9764456FFE05E5C0A7FFA8FFAED0CCD" bold="true" box="[1365,1375,1390,1412]" pageId="14" pageNumber="297">s</emphasis>
|
||
: refer to Figure 10).
|
||
<emphasis id="B9764456FFE05E5C0E05FA6CFEF30CEA" bold="true" box="[303,321,1421,1443]" pageId="14" pageNumber="297">C</emphasis>
|
||
. The arrangement of sensilla on the antennal article VI showing 2 small thin short sensilla (tbs
|
||
<emphasis id="B9764456FFE05E5C0A1EFA6CFA8C0CEA" bold="true" box="[1332,1342,1421,1443]" pageId="14" pageNumber="297">s</emphasis>
|
||
) and the posterior group of 8 thick short conical sensilla (Ts
|
||
<emphasis id="B9764456FFE05E5C0D89FA4DFD1C0C8B" bold="true" box="[675,686,1452,1474]" pageId="14" pageNumber="297">c</emphasis>
|
||
).
|
||
<emphasis id="B9764456FFE05E5C0DEEFA4DFD640C8B" bold="true" box="[708,726,1452,1474]" pageId="14" pageNumber="297">D</emphasis>
|
||
. Clypeo-labrum: lateral lamella (Ll) and lamella (l), posterior row of setae (s) and the anterior row of slender setae (sa).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBD9844FFE05E5C0FEDF9E8FBF90E54" blockId="14.[151,1436,1544,2037]" pageId="14" pageNumber="297">
|
||
<emphasis id="B9764456FFE05E5C0FEDF9E8FF4E0F69" box="[199,252,1545,1568]" italics="true" pageId="14" pageNumber="297">Legs</emphasis>
|
||
: Leg segments are named following
|
||
<bibRefCitation id="EF93E5B5FFE05E5C0DB6F9E8FCFA0F68" author="Manton, S. M." box="[668,840,1544,1569]" pageId="14" pageNumber="297" pagination="153 - 187" refId="ref11810" refString="Manton, S. M. (1956) The Evolution of Arthropodan Locomotory Mechanisms-Part 5. The Structure, Habits and Evolution of the Pselaphognatha (Diplopoda). Journal of the Linnean Society London, 43, 153 - 187. https: // doi. org / 10.1111 / j. 1096 - 3642.1957. tb 02516. x" type="journal article" year="1956">Manton (1956)</bibRefCitation>
|
||
. Legs 1 and 2 are without trochanter, leg 1 also lacks tarsus 1. Chaetotaxy: coxa 1 and 2: 2 setae, coxae 3–13: 2–3 setae; prefemur, post-femur, tibia with 1 seta, except femur with 1–4 setae, tarsus 1 and tarsus 2 with a spine (
|
||
<figureCitation id="133984C1FFE05E5C0C06F9B1FC250F21" box="[812,919,1616,1641]" captionStart="FIGURE 12" captionStartId="15.[151,250,1832,1854]" captionTargetBox="[160,1435,611,1801]" captionTargetId="figure@15.[151,1436,591,1810]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (p: penis, co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2). B. The biarticulated seta with a ridged funicle present on the coxa, prefemur, femur and the last sternite. C. A similar smaller form of B present in the posterior region of the femur, sex organs - penis or vulva. D. Setiform seta in post-femur and tibia. E & F: Spines found on tarsus 1 and 2. G. Dorsal ornamental trichome sockets: sockets of trichomes a: a1 to a12; socket of trichome b; sockets of trichomes c: c1, c2 and c3; circular indentation d. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: setiform process." figureDoi="http://doi.org/10.5281/zenodo.10092688" httpUri="https://zenodo.org/record/10092688/files/figure.png" pageId="14" pageNumber="297">Figs 12A</figureCitation>
|
||
and
|
||
<figureCitation id="133984C1FFE05E5C0CF9F9B1FBB50F21" box="[979,1031,1616,1640]" captionStart="FIGURE 13" captionStartId="16.[151,250,1287,1309]" captionTargetBox="[151,1436,193,1266]" captionTargetId="figure@16.[151,1436,193,1266]" captionTargetPageId="16" captionText="FIGURE 13. Scanning Electron Microscope images of the leg segments, claws and the caudal bundle of Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (refer to figure 12). B. Claw structure indicating that the posterior process and the setiform process were half the length of the claw. C. Caudal bundle in the male (cb) showing the ornamental trichome sockets (ots) which hold trichomes a, b and c, the caudal bundle trichome sockets (cbts) showing the uniform size sockets of the caudal trichomes (ct). D. Caudal bundle in the female showing a similar structure as the male but there are 2 groups of sockets of nest trichomes (nts) present, located ventral to the caudal bundle where the nest trichomes (nt) are carried." figureDoi="http://doi.org/10.5281/zenodo.10092690" httpUri="https://zenodo.org/record/10092690/files/figure.png" pageId="14" pageNumber="297">13A</figureCitation>
|
||
). Coxa and prefemur, femur with a biarticulated seta with a ridged funicle (
|
||
<figureCitation id="133984C1FFE05E5C0D70F995FD700FC5" box="[602,706,1652,1677]" captionStart="FIGURE 12" captionStartId="15.[151,250,1832,1854]" captionTargetBox="[160,1435,611,1801]" captionTargetId="figure@15.[151,1436,591,1810]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (p: penis, co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2). B. The biarticulated seta with a ridged funicle present on the coxa, prefemur, femur and the last sternite. C. A similar smaller form of B present in the posterior region of the femur, sex organs - penis or vulva. D. Setiform seta in post-femur and tibia. E & F: Spines found on tarsus 1 and 2. G. Dorsal ornamental trichome sockets: sockets of trichomes a: a1 to a12; socket of trichome b; sockets of trichomes c: c1, c2 and c3; circular indentation d. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: setiform process." figureDoi="http://doi.org/10.5281/zenodo.10092688" httpUri="https://zenodo.org/record/10092688/files/figure.png" pageId="14" pageNumber="297">Fig. 12B</figureCitation>
|
||
). 1–3 smaller biarticulated setae with the ridged funicles present on posterior position of femur and some also on penis (
|
||
<figureCitation id="133984C1FFE05E5C0DD6F979FC7F0FF9" box="[764,973,1688,1713]" captionStart="FIGURE 12" captionStartId="15.[151,250,1832,1854]" captionTargetBox="[160,1435,611,1801]" captionTargetId="figure@15.[151,1436,591,1810]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (p: penis, co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2). B. The biarticulated seta with a ridged funicle present on the coxa, prefemur, femur and the last sternite. C. A similar smaller form of B present in the posterior region of the femur, sex organs - penis or vulva. D. Setiform seta in post-femur and tibia. E & F: Spines found on tarsus 1 and 2. G. Dorsal ornamental trichome sockets: sockets of trichomes a: a1 to a12; socket of trichome b; sockets of trichomes c: c1, c2 and c3; circular indentation d. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: setiform process." figureDoi="http://doi.org/10.5281/zenodo.10092688" httpUri="https://zenodo.org/record/10092688/files/figure.png" pageId="14" pageNumber="297">Figs 12A and 12C</figureCitation>
|
||
). At distal edge of post-femur and tibia, a setiform seta present (
|
||
<figureCitation id="133984C1FFE05E5C0EA6F95DFE460F9D" box="[396,500,1724,1749]" captionStart="FIGURE 12" captionStartId="15.[151,250,1832,1854]" captionTargetBox="[160,1435,611,1801]" captionTargetId="figure@15.[151,1436,591,1810]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (p: penis, co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2). B. The biarticulated seta with a ridged funicle present on the coxa, prefemur, femur and the last sternite. C. A similar smaller form of B present in the posterior region of the femur, sex organs - penis or vulva. D. Setiform seta in post-femur and tibia. E & F: Spines found on tarsus 1 and 2. G. Dorsal ornamental trichome sockets: sockets of trichomes a: a1 to a12; socket of trichome b; sockets of trichomes c: c1, c2 and c3; circular indentation d. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: setiform process." figureDoi="http://doi.org/10.5281/zenodo.10092688" httpUri="https://zenodo.org/record/10092688/files/figure.png" pageId="14" pageNumber="297">Fig. 12D</figureCitation>
|
||
). A spine on tarsus 2 sharply pointed and twice as long as the spine (
|
||
<figureCitation id="133984C1FFE05E5C0BC4F95DFAFD0F9D" box="[1262,1359,1724,1749]" captionStart="FIGURE 12" captionStartId="15.[151,250,1832,1854]" captionTargetBox="[160,1435,611,1801]" captionTargetId="figure@15.[151,1436,591,1810]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (p: penis, co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2). B. The biarticulated seta with a ridged funicle present on the coxa, prefemur, femur and the last sternite. C. A similar smaller form of B present in the posterior region of the femur, sex organs - penis or vulva. D. Setiform seta in post-femur and tibia. E & F: Spines found on tarsus 1 and 2. G. Dorsal ornamental trichome sockets: sockets of trichomes a: a1 to a12; socket of trichome b; sockets of trichomes c: c1, c2 and c3; circular indentation d. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: setiform process." figureDoi="http://doi.org/10.5281/zenodo.10092688" httpUri="https://zenodo.org/record/10092688/files/figure.png" pageId="14" pageNumber="297">Fig. 12F</figureCitation>
|
||
) in the anterosternal position of tarsus 1 (
|
||
<figureCitation id="133984C1FFE05E5C0D33F901FD320FB1" box="[537,640,1760,1785]" captionStart="FIGURE 12" captionStartId="15.[151,250,1832,1854]" captionTargetBox="[160,1435,611,1801]" captionTargetId="figure@15.[151,1436,591,1810]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (p: penis, co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2). B. The biarticulated seta with a ridged funicle present on the coxa, prefemur, femur and the last sternite. C. A similar smaller form of B present in the posterior region of the femur, sex organs - penis or vulva. D. Setiform seta in post-femur and tibia. E & F: Spines found on tarsus 1 and 2. G. Dorsal ornamental trichome sockets: sockets of trichomes a: a1 to a12; socket of trichome b; sockets of trichomes c: c1, c2 and c3; circular indentation d. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: setiform process." figureDoi="http://doi.org/10.5281/zenodo.10092688" httpUri="https://zenodo.org/record/10092688/files/figure.png" pageId="14" pageNumber="297">Fig. 12E</figureCitation>
|
||
). Posterior edge of last sternite with 2–4 setae similar to those present on coxa, number of these setae varying; 2 present on
|
||
<typeStatus id="54B926E6FFE05E5C0DC1F8E5FCFF0E54" box="[747,845,1796,1821]" pageId="14" pageNumber="297" type="holotype">holotype</typeStatus>
|
||
and 2–4 on
|
||
<typeStatus id="54B926E6FFE05E5C0CF2F8E4FBF50E54" box="[984,1095,1797,1821]" pageId="14" pageNumber="297" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE05E5C0FEDF8C9FA250E2D" blockId="14.[151,1436,1544,2037]" pageId="14" pageNumber="297">
|
||
<emphasis id="B9764456FFE05E5C0FEDF8C9FECD0E08" box="[199,383,1832,1857]" italics="true" pageId="14" pageNumber="297">Telotarsus-Claw</emphasis>
|
||
: A slender structure with an anterior lateral process, a posterior process less than half the length of claw. Anterior setiform process half of the length of claw; a lamella process present (
|
||
<figureCitation id="133984C1FFE05E5C0B93F8ADFA900E2D" box="[1209,1314,1868,1893]" captionStart="FIGURE 12" captionStartId="15.[151,250,1832,1854]" captionTargetBox="[160,1435,611,1801]" captionTargetId="figure@15.[151,1436,591,1810]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (p: penis, co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2). B. The biarticulated seta with a ridged funicle present on the coxa, prefemur, femur and the last sternite. C. A similar smaller form of B present in the posterior region of the femur, sex organs - penis or vulva. D. Setiform seta in post-femur and tibia. E & F: Spines found on tarsus 1 and 2. G. Dorsal ornamental trichome sockets: sockets of trichomes a: a1 to a12; socket of trichome b; sockets of trichomes c: c1, c2 and c3; circular indentation d. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: setiform process." figureDoi="http://doi.org/10.5281/zenodo.10092688" httpUri="https://zenodo.org/record/10092688/files/figure.png" pageId="14" pageNumber="297">Figs 12H</figureCitation>
|
||
and
|
||
<figureCitation id="133984C1FFE05E5C0A72F8ADFA390E2D" box="[1368,1419,1868,1892]" captionStart="FIGURE 13" captionStartId="16.[151,250,1287,1309]" captionTargetBox="[151,1436,193,1266]" captionTargetId="figure@16.[151,1436,193,1266]" captionTargetPageId="16" captionText="FIGURE 13. Scanning Electron Microscope images of the leg segments, claws and the caudal bundle of Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (refer to figure 12). B. Claw structure indicating that the posterior process and the setiform process were half the length of the claw. C. Caudal bundle in the male (cb) showing the ornamental trichome sockets (ots) which hold trichomes a, b and c, the caudal bundle trichome sockets (cbts) showing the uniform size sockets of the caudal trichomes (ct). D. Caudal bundle in the female showing a similar structure as the male but there are 2 groups of sockets of nest trichomes (nts) present, located ventral to the caudal bundle where the nest trichomes (nt) are carried." figureDoi="http://doi.org/10.5281/zenodo.10092690" httpUri="https://zenodo.org/record/10092690/files/figure.png" pageId="14" pageNumber="297">13B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE05E5C0FEDF890FEF00EE5" blockId="14.[151,1436,1544,2037]" pageId="14" pageNumber="297">
|
||
<emphasis id="B9764456FFE05E5C0FEDF890FE740EC0" box="[199,454,1904,1929]" italics="true" pageId="14" pageNumber="297">Sex organs in the male</emphasis>
|
||
: A pair of penes present on coxae 2 and 2 pairs of coxal glands located on coxal plates of legs 8 and 9.
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE05E5D0FEDF859FD430809" blockId="14.[151,1436,1544,2037]" lastBlockId="15.[151,1436,151,536]" lastPageId="15" lastPageNumber="298" pageId="14" pageNumber="297">
|
||
<emphasis id="B9764456FFE05E5C0FEDF859FEBF0E98" box="[199,269,1976,2001]" italics="true" pageId="14" pageNumber="297">Telson</emphasis>
|
||
: Dorsal ornamental trichome sockets arranged almost symmetrically with 8 (L) and 12 (R) sockets of trichomes
|
||
<emphasis id="B9764456FFE05E5C0E25F83DFEAF0EBA" bold="true" box="[271,285,2012,2035]" italics="true" pageId="14" pageNumber="297">a</emphasis>
|
||
on each side of telson in
|
||
<typeStatus id="54B926E6FFE05E5C0D64F83DFD010EBC" box="[590,691,2012,2037]" pageId="14" pageNumber="297" type="holotype">holotype</typeStatus>
|
||
, 7–14 sockets trichomes
|
||
<emphasis id="B9764456FFE05E5C0CFDF83DFC570EBA" bold="true" box="[983,997,2012,2035]" italics="true" pageId="14" pageNumber="297">a</emphasis>
|
||
on each side in
|
||
<typeStatus id="54B926E6FFE05E5C0B82F83CFAA50EBC" box="[1192,1303,2013,2037]" pageId="14" pageNumber="297" type="paratype">paratypes</typeStatus>
|
||
. Sockets of trichomes
|
||
<emphasis id="B9764456FFE15E5D0E26FF79FEA809E6" bold="true" box="[268,282,152,175]" italics="true" pageId="15" pageNumber="298">a</emphasis>
|
||
being more numerous on telsons of
|
||
<typeStatus id="54B926E6FFE15E5D0D84FF79FCAB09F9" box="[686,793,152,176]" pageId="15" pageNumber="298" type="paratype">paratypes</typeStatus>
|
||
in 12 legs stage (Stadium VII) with these sockets ranging from 14–18 on each side of telson. Sockets of trichomes
|
||
<emphasis id="B9764456FFE15E5D0C27FF5DFCA9099A" bold="true" box="[781,795,188,211]" italics="true" pageId="15" pageNumber="298">a</emphasis>
|
||
forming 2 rows, a top row with small sockets and bottom row with larger sockets. The 2 largest sockets located close to the socket of trichome
|
||
<emphasis id="B9764456FFE15E5D0B7BFF3EFBD409B1" box="[1105,1126,223,248]" italics="true" pageId="15" pageNumber="298">
|
||
<emphasis id="B9764456FFE15E5D0B7BFF3EFBED09B1" bold="true" box="[1105,1119,223,248]" italics="true" pageId="15" pageNumber="298">b</emphasis>
|
||
.
|
||
</emphasis>
|
||
3 large sockets trichomes
|
||
<emphasis id="B9764456FFE15E5D0AA5FF01FA2909BE" bold="true" box="[1423,1435,224,247]" italics="true" pageId="15" pageNumber="298">c</emphasis>
|
||
with protruding base:
|
||
<emphasis id="B9764456FFE15E5D0EBCFEE5FE030855" bold="true" box="[406,433,260,284]" italics="true" pageId="15" pageNumber="298">c1</emphasis>
|
||
,
|
||
<emphasis id="B9764456FFE15E5D0EE8FEE5FE6E0855" bold="true" box="[450,476,260,284]" italics="true" pageId="15" pageNumber="298">c2</emphasis>
|
||
and
|
||
<emphasis id="B9764456FFE15E5D0D33FEE5FD810855" bold="true" box="[537,563,260,284]" italics="true" pageId="15" pageNumber="298">c3</emphasis>
|
||
, forming a triangular shape. Circular indentation
|
||
<emphasis id="B9764456FFE15E5D0B5EFEE2FB300855" bold="true" box="[1140,1154,259,284]" italics="true" pageId="15" pageNumber="298">d</emphasis>
|
||
apparent each side near exterior side of trichomes
|
||
<emphasis id="B9764456FFE15E5D0E96FEC9FE7A0876" bold="true" box="[444,456,296,319]" italics="true" pageId="15" pageNumber="298">c</emphasis>
|
||
(
|
||
<figureCitation id="133984C1FFE15E5D0EF2FEC6FDF20809" box="[472,576,295,320]" captionStart="FIGURE 12" captionStartId="15.[151,250,1832,1854]" captionTargetBox="[160,1435,611,1801]" captionTargetId="figure@15.[151,1436,591,1810]" captionTargetPageId="15" captionText="FIGURE 12. Holotype Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (p: penis, co: coxa, pre.fe: prefemur, fe: femur, post.fe: post-femur, tib: tibia, T1: tarsus 1, T2: tarsus 2). B. The biarticulated seta with a ridged funicle present on the coxa, prefemur, femur and the last sternite. C. A similar smaller form of B present in the posterior region of the femur, sex organs - penis or vulva. D. Setiform seta in post-femur and tibia. E & F: Spines found on tarsus 1 and 2. G. Dorsal ornamental trichome sockets: sockets of trichomes a: a1 to a12; socket of trichome b; sockets of trichomes c: c1, c2 and c3; circular indentation d. H. Telotarsus structure with processes: c: claw, l: lamella process, lp: lateral process, p: posterior process, s: setiform process." figureDoi="http://doi.org/10.5281/zenodo.10092688" httpUri="https://zenodo.org/record/10092688/files/figure.png" pageId="15" pageNumber="298">Figs 12G</figureCitation>
|
||
,
|
||
<figureCitation id="133984C1FFE15E5D0D66FEC9FD560809" box="[588,740,295,320]" captionStart="FIGURE 13" captionStartId="16.[151,250,1287,1309]" captionTargetBox="[151,1436,193,1266]" captionTargetId="figure@16.[151,1436,193,1266]" captionTargetPageId="16" captionText="FIGURE 13. Scanning Electron Microscope images of the leg segments, claws and the caudal bundle of Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (refer to figure 12). B. Claw structure indicating that the posterior process and the setiform process were half the length of the claw. C. Caudal bundle in the male (cb) showing the ornamental trichome sockets (ots) which hold trichomes a, b and c, the caudal bundle trichome sockets (cbts) showing the uniform size sockets of the caudal trichomes (ct). D. Caudal bundle in the female showing a similar structure as the male but there are 2 groups of sockets of nest trichomes (nts) present, located ventral to the caudal bundle where the nest trichomes (nt) are carried." figureDoi="http://doi.org/10.5281/zenodo.10092690" httpUri="https://zenodo.org/record/10092690/files/figure.png" pageId="15" pageNumber="298">13C and 13D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBD9844FFE15E5D0FEDFEADFD8F0B51" blockId="15.[151,1436,151,536]" pageId="15" pageNumber="298">
|
||
<emphasis id="B9764456FFE15E5D0FEDFEADFECA082C" box="[199,376,332,357]" italics="true" pageId="15" pageNumber="298">Caudal bundles</emphasis>
|
||
: In males, caudal structure comprised of a group of caudal trichome sockets forming a single structure of uniform size (
|
||
<figureCitation id="133984C1FFE15E5D0EE4FE8EFD8B08C1" box="[462,569,367,392]" captionStart="FIGURE 13" captionStartId="16.[151,250,1287,1309]" captionTargetBox="[151,1436,193,1266]" captionTargetId="figure@16.[151,1436,193,1266]" captionTargetPageId="16" captionText="FIGURE 13. Scanning Electron Microscope images of the leg segments, claws and the caudal bundle of Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (refer to figure 12). B. Claw structure indicating that the posterior process and the setiform process were half the length of the claw. C. Caudal bundle in the male (cb) showing the ornamental trichome sockets (ots) which hold trichomes a, b and c, the caudal bundle trichome sockets (cbts) showing the uniform size sockets of the caudal trichomes (ct). D. Caudal bundle in the female showing a similar structure as the male but there are 2 groups of sockets of nest trichomes (nts) present, located ventral to the caudal bundle where the nest trichomes (nt) are carried." figureDoi="http://doi.org/10.5281/zenodo.10092690" httpUri="https://zenodo.org/record/10092690/files/figure.png" pageId="15" pageNumber="298">Fig. 13C</figureCitation>
|
||
). In females, caudal bundle with 2 distinguishing features: a main dorsal structure, carrying caudal trichomes, similar to that of male caudal bundle; and 2 latero-sternal bundles of nest trichomes, smaller sockets compared to caudal trichome sockets. Nest trichomes using for nest building and protecting egg clusters (Huynh and Veenstra, 2014). Caudal bundle structures well defined with borders between caudal and nest trichomes (
|
||
<figureCitation id="133984C1FFE15E5D0EE2FE1EFD820B51" box="[456,560,511,536]" captionStart="FIGURE 13" captionStartId="16.[151,250,1287,1309]" captionTargetBox="[151,1436,193,1266]" captionTargetId="figure@16.[151,1436,193,1266]" captionTargetPageId="16" captionText="FIGURE 13. Scanning Electron Microscope images of the leg segments, claws and the caudal bundle of Monographis phuquocensis sp. n., A. Leg segments and the chaetotaxy of the second leg and penis (refer to figure 12). B. Claw structure indicating that the posterior process and the setiform process were half the length of the claw. C. Caudal bundle in the male (cb) showing the ornamental trichome sockets (ots) which hold trichomes a, b and c, the caudal bundle trichome sockets (cbts) showing the uniform size sockets of the caudal trichomes (ct). D. Caudal bundle in the female showing a similar structure as the male but there are 2 groups of sockets of nest trichomes (nts) present, located ventral to the caudal bundle where the nest trichomes (nt) are carried." figureDoi="http://doi.org/10.5281/zenodo.10092690" httpUri="https://zenodo.org/record/10092690/files/figure.png" pageId="15" pageNumber="298">Fig. 13D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |