119 lines
14 KiB
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119 lines
14 KiB
XML
<document id="55D2BD2CBC1FA14AA2EB4ED8B39710C4" ID-CLB-Dataset="21238" ID-DOI="10.11646/zootaxa.4608.1.7" ID-GBIF-Dataset="87dff2eb-0e97-4492-94bd-d8fb5190e35d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3993625" ID-ZooBank="14C9F034-F829-471F-9A64-C0EB88D43B95" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597950951697" checkinUser="felipe" docAuthor="Davis, Adrian L. V., Deschodt, Christian M. & Scholtz, Clarke H." docDate="2019" docId="8C1387A16D5DFFC27AA33DE4E545AD79" docLanguage="en" docName="zootaxa.4608.1.7.pdf" docOrigin="Zootaxa 4608 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Endroedyolini Davis, Deschodt & Scholtz 2019, new tribe" docType="treatment" docUuid="547F8B4A-5820-46AA-98FA-8A1A51402143" docUuidSource="ZooBank" docVersion="6" lastPageNumber="139" masterDocId="702AFFD96D5AFFCA7A343D7DE125AF46" masterDocTitle="Defining new dung beetle tribes to resolve discrepancies between phylogeny and tribal classification in the subfamily Scarabaeinae (Coleoptera: Scarabaeidae)" masterLastPageNumber="144" masterPageNumber="131" pageNumber="138" updateTime="1698845016020" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7E62D573B14B86C7BC7D3186DB3D41B9">Defining new dung beetle tribes to resolve discrepancies between phylogeny and tribal classification in the subfamily Scarabaeinae (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="7997555724D260D4E0CEF05C2C181CA8">Davis, Adrian L. V.</mods:namePart>
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<mods:namePart id="EDAFEE3F672BD7F7E6A788EF3CD4B4D0">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="7F553E9FC81AAC50DBEDED3BAFB06E8A">Scholtz, Clarke H.</mods:namePart>
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<mods:typeOfResource id="C8BCA41AA5FBCE99BC12199385683C83">text</mods:typeOfResource>
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<mods:title id="23B650AF356BF3098037E94DA3777976">Zootaxa</mods:title>
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<mods:part id="9DF164A66B129F927EE0343410371D21">
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<mods:date id="9B3ED7F297066CEAB96FBD16B30393F8">2019</mods:date>
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<mods:number id="F6CB53ED034E40ADF5BFB2EF67810DEE">2019-05-17</mods:number>
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<mods:number id="F61504AC49778C257500612072EB84E0">4608</mods:number>
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<treatment id="8C1387A16D5DFFC27AA33DE4E545AD79" ID-DOI="http://doi.org/10.5281/zenodo.4327857" ID-GBIF-Taxon="167284684" ID-Zenodo-Dep="4327857" LSID="urn:lsid:zoobank.org:act:547F8B4A-5820-46AA-98FA-8A1A51402143" httpUri="http://treatment.plazi.org/id/8C1387A16D5DFFC27AA33DE4E545AD79" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138">
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<subSubSection id="4CA0653C6D5DFFCD7AA33DE4E249AFF5" box="[151,876,153,180]" pageId="7" pageNumber="138" type="nomenclature">
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<paragraph id="040536B76D5DFFCD7AA33DE4E249AFF5" blockId="7.[151,876,153,180]" box="[151,876,153,180]" pageId="7" pageNumber="138">
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<heading id="5F4D81DB6D5DFFCD7AA33DE4E249AFF5" bold="true" box="[151,876,153,180]" fontSize="11" level="1" pageId="7" pageNumber="138" reason="1">
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<emphasis id="36CEEAA56D5DFFCD7AA33DE4E249AFF5" bold="true" box="[151,876,153,180]" pageId="7" pageNumber="138">
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Tribe
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<taxonomicName id="C3BA4D346D5DFFCD7AD63DE4E3CCAFF5" ID-CoL="628D9" authority="Davis, Deschodt & Scholtz" authorityName="Davis, Deschodt & Scholtz" authorityYear="2019" box="[226,745,153,180]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Endroedyolini">Endroedyolini Davis, Deschodt & Scholtz</taxonomicName>
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,
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<taxonomicNameLabel id="2DFD57DE6D5DFFCD78C23DE7E249AFF5" box="[758,876,153,180]" pageId="7" pageNumber="138" rank="tribe">new tribe</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="4CA0653C6D5DFFC27AA33D9FE545AD79" lastPageId="8" lastPageNumber="139" pageId="7" pageNumber="138" type="materials_examined">
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<paragraph id="040536B76D5DFFCD7AA33D9FE50CAFBA" blockId="7.[151,1235,225,324]" box="[151,1065,225,252]" pageId="7" pageNumber="138">
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<typeStatus id="DB0188156D5DFFCD7AA33D9FE1EAAFBA" box="[151,207,226,252]" pageId="7" pageNumber="138">Type</typeStatus>
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genus.
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<taxonomicName id="C3BA4D346D5DFFCD7B113D9CE3C3AFBD" authority="Scholtz & Howden, 1987" authorityName="Scholtz & Howden" authorityYear="1987" box="[293,742,225,251]" class="Insecta" family="Scarabaeidae" genus="Endroedyolus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="genus">
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<emphasis id="36CEEAA56D5DFFCD7B113D9CE0E5AFBD" box="[293,448,225,251]" italics="true" pageId="7" pageNumber="138">Endroedyolus</emphasis>
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Scholtz &
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<bibRefCitation id="602B4B466D5DFFCD780A3D9CE3C3AFBD" author="Howden, H. F. & Scholtz, C. H." box="[574,742,225,251]" pageId="7" pageNumber="138" pagination="155 - 192" refId="ref6380" refString="Howden, H. F. & Scholtz, C. H. (1987) A revision of the African genus Odontoloma Boheman (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa, 50, 155 - 192." type="journal article" year="1987">Howden, 1987</bibRefCitation>
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</taxonomicName>
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, here designated (
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<figureCitation id="9C812A326D5DFFCD799B3D9CE2DBAFBD" box="[943,1022,225,251]" captionStart="FIGURE 3" captionStartId="7.[151,250,1954,1979]" captionTargetBox="[169,1430,368,1806]" captionTargetId="figure@7.[153,1438,343,1853]" captionTargetPageId="7" captionText="FIGURE 3. Endroedyolus paradoxus Scholtz & Howden, 1987, type species of the type genus of the tribe Endroedyolini Davis, Deschodt & Scholtz, new tribe." figureDoi="http://doi.org/10.5281/zenodo.3993631" httpUri="https://zenodo.org/record/3993631/files/figure.png" pageId="7" pageNumber="138">Figs. 3</figureCitation>
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,
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<figureCitation id="9C812A326D5DFFCD7E3F3D9CE53EAFBA" box="[1035,1051,225,252]" captionStart="FIGURE 6" captionStartId="10.[151,250,1282,1307]" captionTargetBox="[151,1436,309,1226]" captionTargetId="figure@10.[151,1436,309,1257]" captionTargetPageId="10" captionText="FIGURE 6. Map showing the distribution of the known species in the tribe, Endroedyolini Davis, Deschodt & Scholtz, new tribe." figureDoi="http://doi.org/10.5281/zenodo.3993637" httpUri="https://zenodo.org/record/3993637/files/figure.png" pageId="7" pageNumber="138">6</figureCitation>
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).
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</paragraph>
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<paragraph id="040536B76D5DFFCD7AF33C78E5F6AE59" blockId="7.[151,1235,225,324]" box="[199,1235,261,288]" pageId="7" pageNumber="138">Head. Clypeus bidentate with a notch to the side of each denticle. Antennae nine segmented.</paragraph>
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<paragraph id="040536B76D5DFFCD7AF33C57E552AE05" blockId="7.[151,1235,225,324]" box="[199,1143,297,324]" pageId="7" pageNumber="138">Pronotum. Very convex with lateral margins explanate, sometimes carinate laterally.</paragraph>
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<caption id="50C5663F6D5DFFCD7AA33ADFE0F3A899" ID-DOI="http://doi.org/10.5281/zenodo.3993631" ID-Zenodo-Dep="3993631" httpUri="https://zenodo.org/record/3993631/files/figure.png" pageId="7" pageNumber="138" startId="7.[151,250,1954,1979]" targetBox="[169,1430,368,1806]" targetPageId="7">
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<paragraph id="040536B76D5DFFCD7AA33ADFE0F3A899" blockId="7.[151,1436,1954,2015]" pageId="7" pageNumber="138">
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<emphasis id="36CEEAA56D5DFFCD7AA33ADFE037A8FD" bold="true" box="[151,274,1954,1979]" pageId="7" pageNumber="138">FIGURE 3.</emphasis>
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<taxonomicName id="C3BA4D346D5DFFCD7B233ADFE23CA8FD" authority="Scholtz & Howden, 1987" authorityName="Scholtz & Howden" authorityYear="1987" box="[279,793,1954,1979]" class="Insecta" family="Scarabaeidae" genus="Endroedyolus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="species" species="paradoxus">
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<emphasis id="36CEEAA56D5DFFCD7B233ADFE336A8FC" box="[279,531,1954,1978]" italics="true" pageId="7" pageNumber="138">Endroedyolus paradoxus</emphasis>
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Scholtz &
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<bibRefCitation id="602B4B466D5DFFCD78B73ADFE23CA8FD" author="Howden, H. F. & Scholtz, C. H." box="[643,793,1954,1979]" pageId="7" pageNumber="138" pagination="155 - 192" refId="ref6380" refString="Howden, H. F. & Scholtz, C. H. (1987) A revision of the African genus Odontoloma Boheman (Coleoptera: Scarabaeidae: Scarabaeinae). Journal of the Entomological Society of Southern Africa, 50, 155 - 192." type="journal article" year="1987">Howden, 1987</bibRefCitation>
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</taxonomicName>
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, type species of the type genus of the tribe
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<taxonomicName id="C3BA4D346D5DFFCD7EF03ADFE043A899" authority="Davis, Deschodt & Scholtz" authorityName="Davis, Deschodt & Scholtz" authorityYear="2019" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="138" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Endroedyolini">Endroedyolini Davis, Deschodt & Scholtz</taxonomicName>
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,
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<taxonomicNameLabel id="2DFD57DE6D5DFFCD7B443ABAE0F7A899" box="[368,466,1990,2015]" pageId="7" pageNumber="138" rank="tribe">new tribe</taxonomicNameLabel>
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.
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</paragraph>
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</caption>
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<paragraph id="040536B76D52FFC27AF33DE4E44AAFF5" blockId="8.[151,1436,153,575]" box="[199,1391,153,179]" pageId="8" pageNumber="139">Elytra. Strongly convex; setae in straight and slightly converging lines; metathoracic wings always absent.</paragraph>
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<paragraph id="040536B76D52FFC27AF33DC0E03EAE59" blockId="8.[151,1436,153,575]" pageId="8" pageNumber="139">Sterna. Smooth to punctate with punctures of varying density and size; prosternum fairly well developed; metasternum smooth to punctate, sometimes with deep fovea extending from anterior to sometimes half way between mesocoxae.</paragraph>
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<paragraph id="040536B76D52FFC27AF33C54E20FAE05" blockId="8.[151,1436,153,575]" box="[199,810,297,323]" pageId="8" pageNumber="139">Protibia. With three small denticles on outside margin.</paragraph>
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<paragraph id="040536B76D52FFC27AF33C30E3E7AE21" blockId="8.[151,1436,153,575]" box="[199,706,333,359]" pageId="8" pageNumber="139">Meso- and metatibia. Unmodified with setae.</paragraph>
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<paragraph id="040536B76D52FFC27AF33C0CE56CAECD" blockId="8.[151,1436,153,575]" box="[199,1097,369,395]" pageId="8" pageNumber="139">Pygidium. With small setiferous punctures; with or without depression or fovea.</paragraph>
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<paragraph id="040536B76D52FFC27AF33CE8E374AEE9" blockId="8.[151,1436,153,575]" box="[199,593,405,431]" pageId="8" pageNumber="139">Aedeagus. Parameres symmetrical.</paragraph>
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<paragraph id="040536B76D52FFC27AF33CC4E3DEAE95" blockId="8.[151,1436,153,575]" box="[199,763,440,467]" pageId="8" pageNumber="139">
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Body size. Very small to small:
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to
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<quantity id="C3429B526D52FFC278AA3CC4E3D2AE95" box="[670,759,441,467]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="139" unit="mm" value="4.8">4.8 mm</quantity>
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.
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</paragraph>
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<paragraph id="040536B76D52FFC27AF33CA0E545AD79" blockId="8.[151,1436,153,575]" pageId="8" pageNumber="139">
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Diagnosis and known distribution. The combination of extreme small size, very convex body, unique clypeal shape and explanate lateral pronotal margins, distinguishes members of this tribe from any other. They are restricted to forest patches of south east
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<collectingCountry id="7CAD76276D52FFC27BDE3F58E35CAD79" box="[490,633,549,575]" name="South Africa" pageId="8" pageNumber="139">South Africa</collectingCountry>
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where they are associated with forest litter.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |