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<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e101441" ID-Pensoft-Pub="1864-8312-81-475" ID-Pensoft-UUID="92502718827453528F8B0D342C6EF134" ID-ZooBank="5992057016FD4897BAA55272443A29F3" ModsDocID="1864-8312-81-475" checkinTime="1685027893856" checkinUser="pensoft" docAuthor="Rohacek, Jindrich, Hammel, Joerg U. &amp; Baranov, Viktor" docDate="2023" docId="829AD03B700B5604BCA72E0C1E3AB65F" docLanguage="en" docName="ArthropodSystPhyl 81: 475-498" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-05-25" docSource="http://dx.doi.org/10.3897/asp.81.e101441" docTitle="Christelenka Roháček &amp; Hammel &amp; Baranov 2023, fam. nov." docType="treatment" docUuid="AA3CA70D-2918-4836-AA9E-2CFE288B06DE" docUuidSource="ZooBank" docVersion="1" id="92502718827453528F8B0D342C6EF134" lastPageNumber="475" masterDocId="92502718827453528F8B0D342C6EF134" masterDocTitle="Christelenkidae, a new extinct family based on a new taxon from Eocene Baltic amber (Diptera: Acalyptratae), with X-ray synchrotron microtomography imaging of its structures" masterLastPageNumber="498" masterPageNumber="475" pageNumber="475" updateTime="1685027893856" updateUser="pensoft">
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<mods:title id="D44C3F91AF729260F1CAEFD70E7BAED3">Christelenkidae, a new extinct family based on a new taxon from Eocene Baltic amber (Diptera: Acalyptratae), with X-ray synchrotron microtomography imaging of its structures</mods:title>
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<mods:namePart id="E5BF5BF566ECB95E50513A8DB4D443C8">Rohacek, Jindrich</mods:namePart>
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<mods:affiliation id="8AC3506552DADAB9320BE8B1294DA357">Slezske zemske muzeum, Nadrazni okruh 31, CZ- 746 01 Opava, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="6E6BE1D7C98669C0A57D3EB817F136B4" type="email">rohacek@szm.cz</mods:nameIdentifier>
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<mods:namePart id="74EA5BA82E319FDBBAEC3761DF973CFC">Hammel, Joerg U.</mods:namePart>
<mods:affiliation id="D64AE6E9336285433575E6CEE9749FEB">Helmholtz-Zentrum Geesthacht, Max-Planck-Strasse 1, 21502 Geesthacht, Germany</mods:affiliation>
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<mods:namePart id="73457AE87662B01AE415651B4D6601A7">Baranov, Viktor</mods:namePart>
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<mods:affiliation id="8E74CFE476E01B270CE9A2C843B9D7CD">Estacion Biologica de Donana-CSIC, Avd. Americo Vespucio 26, 41092 Sevilla, Spain</mods:affiliation>
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<mods:date id="5C5915115AA23A190245FF0EE0350FA2">2023</mods:date>
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<mods:number id="CF1CE68FD99512AF767E0084D61854F2">81</mods:number>
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<paragraph id="C06A0AF567FCEF34DB8802EB142A514B" pageId="0" pageNumber="475">
3.1.
<taxonomicName id="BA10210374DE22083442A4D0B1D32FAF" LSID="http://zoobank.org/AA3CA70D-2918-4836-AA9E-2CFE288B06DE" authority="Roháček" authorityName="Roháček &amp; Hammel &amp; Baranov" authorityYear="2023" family="Christelenkidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="475" rank="family" status="fam. nov.">
Christelenkidae
<normalizedToken id="1FFD59477858AA14AEC87341DE7D7BFB" originalValue="Roháček">Rohacek</normalizedToken>
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<taxonomicNameLabel id="A7B719471E7DEC6DFDEE8DD414A3D674" pageId="0" pageNumber="475">fam. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="2AA735212724DCB45F2D5665E8D5AD5C" pageId="0" pageNumber="475" type="type genus">
<paragraph id="78BD8CC6ADC913ED2E87463E60483E6C" pageId="0" pageNumber="475">Type genus.</paragraph>
<paragraph id="52841717D0F51ABCE9A7A6B5D94481DF" pageId="0" pageNumber="475">
<taxonomicName id="AD2B0BB1B69FC5FA1CEDE37F93A5D124" authorityName="Roháček &amp; Hammel &amp; Baranov" authorityYear="2023" family="Christelenkidae" genus="Christelenka" kingdom="Animalia" lsidName="Christelenka" order="Diptera" pageId="0" pageNumber="475" rank="genus">
<emphasis id="93C1708D074032BE68E5DFC2687E63F7" italics="true" pageId="0" pageNumber="475">Christelenka</emphasis>
</taxonomicName>
gen. nov., designated here.
</paragraph>
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<subSubSection id="86ACE830FD040DD03072608E076CAA4B" pageId="0" pageNumber="475" type="diagnosis">
<paragraph id="E107286221C8EB0223D95CED3462B170" pageId="0" pageNumber="475">Diagnosis.</paragraph>
<paragraph id="D0B5535D870B1389C7A854096FFBD444" pageId="0" pageNumber="475">
Body: relatively slender and elongate, particularly the abdomen, ca 3 mm long (Figs
<figureCitation id="EC4A360D5A2F9B329BE1A430F0F53C02" captionStart="Figures 1, 2" captionStartId="F1" captionText="Figures 1, 2. Christelenka multiplex sp. nov., male holotype. 1 specimen in the whole amber preparatum, left laterally, with blue background; 2 whole specimen dorsally, with blue background. Scale bars: 1.0 mm. Photos by J. Rohacek." figureDoi="10.3897/asp.81.e101441.figures1-2" httpUri="https://binary.pensoft.net/fig/854546" pageId="0" pageNumber="475">1</figureCitation>
-
<figureCitation id="D5B7BF0BD2D46CC3C9CA41F0D52E9FAF" captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Christelenka multiplex sp. nov., male holotype. 3 whole specimen, left laterally; 4 ditto, right laterally. Scale bars: 1.0 mm. Photos by J. Rohacek." figureDoi="10.3897/asp.81.e101441.figures3-4" httpUri="https://binary.pensoft.net/fig/854547" pageId="0" pageNumber="475">4</figureCitation>
). Head: Antenna exclinate (Figs
<figureCitation id="0C157F88A21A7C395269E3F47D322871" captionStart="Figures 57" captionStartId="F3" captionText="Figures 5 - 7. Christelenka multiplex sp. nov., male holotype. 5 head left laterally; 6 ditto, frontally; 7 ditto, laterocaudally. Scale bar: 0.5 mm. Photos by J. Rohacek." figureDoi="10.3897/asp.81.e101441.figures5-7" httpUri="https://binary.pensoft.net/fig/854548" pageId="0" pageNumber="475">6</figureCitation>
,
<figureCitation id="3C0D0B54D09448A205ED7885FBC134D1" captionStart="Figs 810" captionStartId="F4" captionText="Figs 8 - 10. Christelenka multiplex sp. nov., male holotype, cephalic chaetotaxy. 8 head left laterally; 9 ditto, frontally; 10 ditto, laterocaudally. - Scale bar: 0.5 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figs8-10" httpUri="https://binary.pensoft.net/fig/854549" pageId="0" pageNumber="475">9</figureCitation>
) and distinctly geniculate between pedicel and 1st flagellomere, the latter strongly decumbent (Figs
<figureCitation id="579E17C6D0035B8625E6B6F36C6B6C66" captionStart="Figures 57" captionStartId="F3" captionText="Figures 5 - 7. Christelenka multiplex sp. nov., male holotype. 5 head left laterally; 6 ditto, frontally; 7 ditto, laterocaudally. Scale bar: 0.5 mm. Photos by J. Rohacek." figureDoi="10.3897/asp.81.e101441.figures5-7" httpUri="https://binary.pensoft.net/fig/854548" pageId="0" pageNumber="475">5</figureCitation>
,
<figureCitation id="6229A881641094E23C56C2B286A28AA2" captionStart="Figs 810" captionStartId="F4" captionText="Figs 8 - 10. Christelenka multiplex sp. nov., male holotype, cephalic chaetotaxy. 8 head left laterally; 9 ditto, frontally; 10 ditto, laterocaudally. - Scale bar: 0.5 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figs8-10" httpUri="https://binary.pensoft.net/fig/854549" pageId="0" pageNumber="475">8</figureCitation>
), discoidal but slightly elongate (somewhat longer than scape + pedicel); arista extremely dorsobasal; pedicel cap-like, with distal margin simple. Frontal triangle not developed; lunule absent (Figs
<figureCitation id="C9F9B0796B811C2421F021054CF0A4B9" captionStart="Figures 57" captionStartId="F3" captionText="Figures 5 - 7. Christelenka multiplex sp. nov., male holotype. 5 head left laterally; 6 ditto, frontally; 7 ditto, laterocaudally. Scale bar: 0.5 mm. Photos by J. Rohacek." figureDoi="10.3897/asp.81.e101441.figures5-7" httpUri="https://binary.pensoft.net/fig/854548" pageId="0" pageNumber="475">6</figureCitation>
,
<figureCitation id="5CC04CAB81C1BFB36614B55B4A2373E4" captionStart="Figs 810" captionStartId="F4" captionText="Figs 8 - 10. Christelenka multiplex sp. nov., male holotype, cephalic chaetotaxy. 8 head left laterally; 9 ditto, frontally; 10 ditto, laterocaudally. - Scale bar: 0.5 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figs8-10" httpUri="https://binary.pensoft.net/fig/854549" pageId="0" pageNumber="475">9</figureCitation>
). Cephalic setae long but only present on posterior third of frons and on vertex (Figs
<figureCitation id="60A92564E69D65E54B5D13D768975A72" captionStart="Figs 810" captionStartId="F4" captionText="Figs 8 - 10. Christelenka multiplex sp. nov., male holotype, cephalic chaetotaxy. 8 head left laterally; 9 ditto, frontally; 10 ditto, laterocaudally. - Scale bar: 0.5 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figs8-10" httpUri="https://binary.pensoft.net/fig/854549" pageId="0" pageNumber="475">8</figureCitation>
-
<figureCitation id="20049210189BF37453D73CA6120F6F5D" captionStart="Figs 810" captionStartId="F4" captionText="Figs 8 - 10. Christelenka multiplex sp. nov., male holotype, cephalic chaetotaxy. 8 head left laterally; 9 ditto, frontally; 10 ditto, laterocaudally. - Scale bar: 0.5 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figs8-10" httpUri="https://binary.pensoft.net/fig/854549" pageId="0" pageNumber="475">10</figureCitation>
); pvt small, convergent (crossed); vti by far the longest cephalic seta, upright and reclinate; vte strongly exclinate; oc upright to proclinate and divergent, longer than pvt; only 1 reclinate and slightly exclinate ors; frons in front of ors bare; vibrissa small but distinct, strongly medially curved; no genal seta or setulae. Thorax: with 1 postsutural dc, 1 prs, 1 sa, 1 pa, 2 sc (apical crossed, laterobasal the longest thoracic seta); 0 mspl, 1 stpl, 0 ppl. Mesonotum convex, somewhat hump-like; prosternum bare; mesopleuron with posterodorsal ridge developed (Fig.
<figureCitation id="41B3D2ED144CB4DEE81FBE08F09FD96D" captionStart="Figures 13, 14" captionStartId="F6" captionText="Figures 13, 14. Christelenka multiplex sp. nov., male holotype. 13 whole specimen, left laterally; 14 ditto, right laterally. Scale bar: 1.0 mm. Microtomograph images by V. Baranov." figureDoi="10.3897/asp.81.e101441.figures13-14" httpUri="https://binary.pensoft.net/fig/854551" pageId="0" pageNumber="475">14</figureCitation>
); subscutellum distinctly protruding (Fig.
<figureCitation id="9CD3131C3977E934FD174E3E5BAA2814" captionStart="Figure 17" captionStartId="F8" captionText="Figure 17. Christelenka multiplex sp. nov., male holotype, anterior part of body: sagittal section right laterally. Scale bar: 0.5 mm. For abbreviations see 2.2.4. Morphological terminology. Microtomograph image by V. Baranov." figureDoi="10.3897/asp.81.e101441.figure17" httpUri="https://binary.pensoft.net/fig/854553" pageId="0" pageNumber="475">17</figureCitation>
), metasternal area bare. Wing: with apex at end of R4+5, somewhat pointed; membrane not distinctly patterned; C extending to apex of M, with 2 breaks (humeral and subcostal) proximally, and with uniform dark setulae (none spine-like) ending between apices of R2+3 and R4+5 (see Fig.
<figureCitation id="D23EECE16D088640BBE729C336012584" captionStart="Figures 18, 19" captionStartId="F9" captionText="Figures 18, 19. Christelenka multiplex sp. nov., male holotype, wings. 18 right wing, dorsally; 19 left wing, dorsally. Scale bar: 1.0 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figures18-19" httpUri="https://binary.pensoft.net/fig/854554" pageId="0" pageNumber="475">18</figureCitation>
); Sc basally distinct but distally fused with R1, the latter without a preapical kink; no distinct humeral crossvein; cells dm, bm and cup complete; alula (not clearly visible) small and anal lobe well developed. Legs: cx1 elongate (Fig.
<figureCitation id="DAFAD9232D1DAF171874E57DC48CD2B3" captionStart="Figures 13, 14" captionStartId="F6" captionText="Figures 13, 14. Christelenka multiplex sp. nov., male holotype. 13 whole specimen, left laterally; 14 ditto, right laterally. Scale bar: 1.0 mm. Microtomograph images by V. Baranov." figureDoi="10.3897/asp.81.e101441.figures13-14" httpUri="https://binary.pensoft.net/fig/854551" pageId="0" pageNumber="475">13</figureCitation>
), and with 1 ventral seta in middle (Fig.
<figureCitation id="E6A74A66F2939F77F28AEFC9CF21DB55" captionStart="Figures 15, 16" captionStartId="F7" captionText="Figures 15, 16. Christelenka multiplex sp. nov., male holotype, thoracic chaetotaxy. 15 thorax, left laterally; 16 ditto, dorsally. Scale bar: 0.5 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figures15-16" httpUri="https://binary.pensoft.net/fig/854552" pageId="0" pageNumber="475">15</figureCitation>
); f1 with 1 dorsal seta and an anteroventral ctenidium-like row of small blunt spines (Fig.
<figureCitation id="597132D569FBFF80D660AE982F66D4EF" captionStart="Figures 2023" captionStartId="F10" captionText="Figures 20 - 23. Christelenka multiplex sp. nov., male holotype, legs. 20 left fore femur, dorsally; 21 right fore femur, anteriorly; 22 right mid femur and tibia, anteriorly; 23 left hind femur, tibia and tarsus, anteriorly. Scale bar: 0.3 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figures20-23" httpUri="https://binary.pensoft.net/fig/854555" pageId="0" pageNumber="475">21</figureCitation>
); f3 with 1 dorsal and 1 anteroventral seta (Fig.
<figureCitation id="752D5C27C3D8ACABB13003F4CBE20999" captionStart="Figures 2023" captionStartId="F10" captionText="Figures 20 - 23. Christelenka multiplex sp. nov., male holotype, legs. 20 left fore femur, dorsally; 21 right fore femur, anteriorly; 22 right mid femur and tibia, anteriorly; 23 left hind femur, tibia and tarsus, anteriorly. Scale bar: 0.3 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figures20-23" httpUri="https://binary.pensoft.net/fig/854555" pageId="0" pageNumber="475">23</figureCitation>
); all tibiae with 1 dorsal preapical seta. Abdomen (of male) with unusual preabdominal sclerites: segments 1-4 with both terga and sterna short, while T5 and S5 are unusually strongly elongate (Figs
<figureCitation id="3D9F475F9A7A0619D70E05B3DFA6640D" captionStart="Figures 24, 25" captionStartId="F11" captionText="Figures 24, 25. Christelenka multiplex sp. nov., male holotype. 24 abdomen, left laterally; 25 ditto, with some structures coloured. Scale bar: 0.5 mm. For abbreviations see 2.2.4. Morphological terminology. Microtomograph image by V. Baranov." figureDoi="10.3897/asp.81.e101441.figures24-25" httpUri="https://binary.pensoft.net/fig/854556" pageId="0" pageNumber="475">25</figureCitation>
,
<figureCitation id="740F41B6B1F7B38B873CE024591F5012" captionStart="Figures 2830" captionStartId="F13" captionText="Figures 28 - 30. Christelenka multiplex sp. nov., male holotype, abdominal structures. 28 abdomen, left laterally; 29 postabdomen with terminalia, right laterally; 30 ditto, left laterally. Scale bars: 28 - 0.5 mm; others - 0.2 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figures28-30" httpUri="https://binary.pensoft.net/fig/854558" pageId="0" pageNumber="475">28</figureCitation>
). Male postabdomen: T6 reduced (transversely band-like) but distinct and slightly asymmetrical; S6 of moderate length and symmetrical; T7 absent; S7+S8 fused to form a somewhat asymmetrical dorsal synsclerite with a pair of strong posterior setae. Male genitalia: epandrium arch-shaped, slightly asymmetrical and open ventrally (Figs
<figureCitation id="1AA3FB4324F7AFACB49D2EF40D070776" captionStart="Figures 2830" captionStartId="F13" captionText="Figures 28 - 30. Christelenka multiplex sp. nov., male holotype, abdominal structures. 28 abdomen, left laterally; 29 postabdomen with terminalia, right laterally; 30 ditto, left laterally. Scale bars: 28 - 0.5 mm; others - 0.2 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figures28-30" httpUri="https://binary.pensoft.net/fig/854558" pageId="0" pageNumber="475">29</figureCitation>
,
<figureCitation id="CC383318676C1773380592AD7489D108" captionStart="Figures 2830" captionStartId="F13" captionText="Figures 28 - 30. Christelenka multiplex sp. nov., male holotype, abdominal structures. 28 abdomen, left laterally; 29 postabdomen with terminalia, right laterally; 30 ditto, left laterally. Scale bars: 28 - 0.5 mm; others - 0.2 mm. For abbreviations see 2.2.4. Morphological terminology." figureDoi="10.3897/asp.81.e101441.figures28-30" httpUri="https://binary.pensoft.net/fig/854558" pageId="0" pageNumber="475">30</figureCitation>
); cerci separate, large, inserted below large anal fissure; gonostyli somewhat asymmetrical, simple, elongate, tapered towards blunt apex, articulated with epandrium; hypandrium, short, obviously frame-shaped and symmetrical (Fig.
<figureCitation id="599CFBEAE65FE1CFBA92EC7DAFFD2C62" captionStart="Figures 3135" captionStartId="F14" captionText="Figures 31 - 35. Christelenka multiplex sp. nov., male holotype, abdominal structures. 31 abdomen (without 1 st segment), sagittal section right laterally, with some structures coloured; 32 posterior end of abdomen, right ventrolaterally, with structures coloured; 33 ditto, without colours; 34 posterior end of abdomen, ventrally; 35 ditto, enlarged and coloured. Scale bars: 0.2 mm. For abbreviations see 2.2.4. Morphological terminology. Microtomograph images by V. Baranov." figureDoi="10.3897/asp.81.e101441.figures31-35" httpUri="https://binary.pensoft.net/fig/854559" pageId="0" pageNumber="475">35</figureCitation>
); postgonites simple, elongate and pointed, symmetrical; distal part of aedeagus (considered to be distiphallus) short, broad distally (Figs
<figureCitation id="896DB390F2642BE09648752C91E7C25E" captionStart="Figures 3135" captionStartId="F14" captionText="Figures 31 - 35. Christelenka multiplex sp. nov., male holotype, abdominal structures. 31 abdomen (without 1 st segment), sagittal section right laterally, with some structures coloured; 32 posterior end of abdomen, right ventrolaterally, with structures coloured; 33 ditto, without colours; 34 posterior end of abdomen, ventrally; 35 ditto, enlarged and coloured. Scale bars: 0.2 mm. For abbreviations see 2.2.4. Morphological terminology. Microtomograph images by V. Baranov." figureDoi="10.3897/asp.81.e101441.figures31-35" httpUri="https://binary.pensoft.net/fig/854559" pageId="0" pageNumber="475">32</figureCitation>
,
<figureCitation id="8BFD06DB10763766A38FD76888083794" captionStart="Figures 3135" captionStartId="F14" captionText="Figures 31 - 35. Christelenka multiplex sp. nov., male holotype, abdominal structures. 31 abdomen (without 1 st segment), sagittal section right laterally, with some structures coloured; 32 posterior end of abdomen, right ventrolaterally, with structures coloured; 33 ditto, without colours; 34 posterior end of abdomen, ventrally; 35 ditto, enlarged and coloured. Scale bars: 0.2 mm. For abbreviations see 2.2.4. Morphological terminology. Microtomograph images by V. Baranov." figureDoi="10.3897/asp.81.e101441.figures31-35" httpUri="https://binary.pensoft.net/fig/854559" pageId="0" pageNumber="475">35</figureCitation>
).
</paragraph>
<caption id="30DDC1B75CB91EF5764D20AC9CECC231" doi="10.3897/asp.81.e101441.figures3-4" httpUri="https://binary.pensoft.net/fig/854547" pageId="0" pageNumber="475" start="Figures 3, 4" startId="F2">
<paragraph id="B0FE4296C330923C6A6590DB52BCAB3A" pageId="0" pageNumber="475">
<emphasis id="67FE800A6FD8DFFD9C5CF554B14F9B3F" bold="true" pageId="0" pageNumber="475">Figures 3, 4.</emphasis>
<taxonomicName id="51F6F891CDDE4A171A9086C6401AE861" authorityName="Rohacek" authorityYear="2023" family="Christelenkidae" genus="Christelenka" kingdom="Animalia" lsidName="Christelenka multiplex" order="Diptera" pageId="0" pageNumber="475" rank="species" species="multiplex">
<emphasis id="B652B7E004EB77CCDAE09EFEBBCAEE9D" italics="true" pageId="0" pageNumber="475">Christelenka multiplex</emphasis>
</taxonomicName>
<emphasis id="FA66E234D4337D4771F2A81F40BAB2C1" bold="true" pageId="0" pageNumber="475">sp. nov.</emphasis>
, male holotype.
<emphasis id="1158B90DEEC3B6A06F1F69B4847D1A3A" bold="true" pageId="0" pageNumber="475">3</emphasis>
whole specimen, left laterally;
<emphasis id="086C08A20FF59687625573211C54CEC6" bold="true" pageId="0" pageNumber="475">4</emphasis>
ditto, right laterally. Scale bars: 1.0 mm. Photos by J.
<normalizedToken id="4C62C5B088A9BBF07353107FA1E7D556" originalValue="Roháček">Rohacek</normalizedToken>
.
</paragraph>
</caption>
<paragraph id="05FB00804B8C1612E5C196C7044ED6C4" pageId="0" pageNumber="475">
For more detailed description see below under
<taxonomicName id="C5168430ABFD1815F0115898E65D19BF" authorityName="Roháček &amp; Hammel &amp; Baranov" authorityYear="2023" family="Christelenkidae" genus="Christelenka" kingdom="Animalia" lsidName="Christelenka" order="Diptera" pageId="0" pageNumber="475" rank="genus">
<emphasis id="4C9F221DB992FC6364B2F9FC5A6F378C" italics="true" pageId="0" pageNumber="475">Christelenka</emphasis>
</taxonomicName>
gen.n.
</paragraph>
</subSubSection>
</treatment>
</document>