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<document id="31B6DA5454D5AF15EA607A0C498935C6" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D9C92DAFDFBFE4ACD202E5A" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius yamdena Telnov 2020, sp. nov." docType="treatment" docUuid="E3C00D89-AD86-4615-A09A-CA0CD01E1DC4" docUuidSource="ZooBank" docVersion="8" lastPageNumber="130" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="129" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="3BD99395F9756F2348B83C298DCAC253" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="04EA08DEC06E511AECF7A69AD4844935">
<mods:title id="865A81CE6604934CACF01DDAB3571C00">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="69ABCE0A1ACE4337F04E0DA9BF4E99F9" type="personal">
<mods:role id="D4E0304B7D43FFB5284A28FEB7D1290A">
<mods:roleTerm id="0605B533CFE76CA4F5C4F0D0E0336A49">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1328C28A4B5B4D4C1D9A544CC2D8628D">Telnov, Dmitry</mods:namePart>
</mods:name>
<mods:typeOfResource id="C347F9913D00D31EBE782F6A3E262BAA">text</mods:typeOfResource>
<mods:relatedItem id="BBF45D735BF60EA0B68366EBC8BDE76B" type="host">
<mods:titleInfo id="2ED00D1A5FA09893CA8463444FB21959">
<mods:title id="C90DC5E46BF5799DEB8FB5F0EECB40D9">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="E691D2FE594F0F21EB34C73107EE8942">
<mods:date id="F1820367367A2BFD09C38444EC3AA9F0">2020</mods:date>
<mods:detail id="663DBED4A52FFC4F0D3028815E979C70" type="pubDate">
<mods:number id="2B11580ADA4A438FFA9E65F1B4A2A34A">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="CA35495C1C5E648F1F9A645A4035380C" type="volume">
<mods:number id="A3D11E5C947B4335D39347C7F910DB98">721</mods:number>
</mods:detail>
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<mods:start id="A8FE5E8BCD7F7DAD71F3FC764CA9B647">1</mods:start>
<mods:end id="054613AEB16C797F7BF13ADD3FBECB2A">210</mods:end>
</mods:extent>
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<mods:classification id="300E2BAAB57DFF19284E6FFCF9DFF91F">journal article</mods:classification>
<mods:identifier id="E684BCF3D2CFB8E409DEC1B6650F0D82" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="F6E81A19ECFF2E51C242E282D54069EB" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="CE11A8F3272BF9A3791B596B2CF56B83" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="7058600DF6F89EF263C7CB989AD46528" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="3CE65D9554BCDE96633A6CDCE6D6558A" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
</mods:mods>
<treatment id="1A3787F69D9C92DAFDFBFE4ACD202E5A" ID-GBIF-Taxon="168648119" LSID="urn:lsid:zoobank.org:act:E3C00D89-AD86-4615-A09A-CA0CD01E1DC4" httpUri="http://treatment.plazi.org/id/1A3787F69D9C92DAFDFBFE4ACD202E5A" lastPageId="129" lastPageNumber="130" pageId="128" pageNumber="129">
<subSubSection id="DA84656B9D9C92DBFDFBFE4ACB9A2A6D" box="[616,972,500,527]" pageId="128" pageNumber="129" type="nomenclature">
<paragraph id="922136E09D9C92DBFDFBFE4ACB9A2A6D" blockId="128.[354,1234,500,598]" box="[616,972,500,527]" pageId="128" pageNumber="129">
<heading id="C969818C9D9C92DBFDFBFE4ACB9A2A6D" box="[616,972,500,527]" centered="true" fontSize="11" level="2" pageId="128" pageNumber="129" reason="2">
<taxonomicName id="559E4D639D9C92DBFDFBFE4ACB3C2A6D" ID-CoL="7X3XZ" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[616,874,500,527]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="yamdena" status="sp. nov.">
<emphasis id="A0EAEAF29D9C92DBFDFBFE4ACB3C2A6D" bold="true" box="[616,874,500,527]" italics="true" pageId="128" pageNumber="129">Maechidius yamdena</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D9C92DBFCE2FE4BCB9A2A6D" box="[881,972,501,527]" pageId="128" pageNumber="129" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D9C92DBFEF1FDA6CC952A34" pageId="128" pageNumber="129" type="description">
<paragraph id="922136E09D9C92DBFEF1FDA6CC842A51" blockId="128.[354,1234,500,598]" box="[354,1234,536,563]" pageId="128" pageNumber="129">
<uri id="E60F3AE29D9C92DBFEF1FDA6CC842A51" box="[354,1234,536,563]" pageId="128" pageNumber="129">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D9C92DBFD11FDA6CC842A51" box="[642,1234,536,563]" pageId="128" pageNumber="129">E3C00D89-AD86-4615-A09A-CA0CD01E1DC4</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D9C92DBFEE2FD85CC952A34" blockId="128.[354,1234,500,598]" box="[369,1219,571,598]" pageId="128" pageNumber="129">
<figureCitation id="0AA52A659D9C92DBFEE2FD85C99D2A37" box="[369,459,571,597]" captionStart="Figs 94–96" captionStartId="126.[189,241,1928,1954]" captionTargetBox="[346,1191,267,1904]" captionTargetId="figure-55@126.[339,1249,265,1909]" captionTargetPageId="126" captionText="Figs 94–96. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 94. M. weigeli sp. nov., holotype, ♂ (NME). 95. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 96. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122171" httpUri="https://zenodo.org/record/4122171/files/figure.png" pageId="128" pageNumber="129">Figs 96</figureCitation>
,
<figureCitation id="0AA52A659D9C92DBFE44FD85CA162A34" box="[471,576,571,598]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="128" pageNumber="129">195–196</figureCitation>
,
<figureCitation id="0AA52A659D9C92DBFDDFFD85CA2D2A34" box="[588,635,571,598]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="128" pageNumber="129">265</figureCitation>
,
<figureCitation id="0AA52A659D9C92DBFD14FD85CAE02A34" box="[647,694,571,598]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="128" pageNumber="129">356</figureCitation>
,
<figureCitation id="0AA52A659D9C92DBFD52FD85CAA62A37" box="[705,752,571,597]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="128" pageNumber="129">428</figureCitation>
,
<figureCitation id="0AA52A659D9C92DBFD6FFD85CB8D2A37" box="[764,987,571,597]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="128" pageNumber="129">441–442, 447–448</figureCitation>
, 534, 547, 745–747
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D9C92DBFF2EFDC2C9272B6E" pageId="128" pageNumber="129" type="diagnosis">
<paragraph id="922136E09D9C92DBFF2EFDC2C99F2AF4" blockId="128.[189,457,636,662]" box="[189,457,636,662]" pageId="128" pageNumber="129">
<heading id="C969818C9D9C92DBFF2EFDC2C99F2AF4" bold="true" box="[189,457,636,662]" fontSize="11" level="3" pageId="128" pageNumber="129" reason="3">
<emphasis id="A0EAEAF29D9C92DBFF2EFDC2C99F2AF4" bold="true" box="[189,457,636,662]" pageId="128" pageNumber="129">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D9C92DBFF2EFD15C9272B6E" blockId="128.[189,1398,683,780]" pageId="128" pageNumber="129">This species is peculiar due to the combination of the following characters: structure of labroclypeus, shape of lateral margins of pronotum, rugulose punctures of elytra, very short dorsal setae and shape of male aedeagus.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D9C92DBFF2EFC8AC90B2BFD" pageId="128" pageNumber="129" type="etymology">
<paragraph id="922136E09D9C92DBFF2EFC8AC9122B2C" blockId="128.[189,324,820,846]" box="[189,324,820,846]" pageId="128" pageNumber="129">
<heading id="C969818C9D9C92DBFF2EFC8AC9122B2C" bold="true" box="[189,324,820,846]" fontSize="11" level="3" pageId="128" pageNumber="129" reason="3">
<emphasis id="A0EAEAF29D9C92DBFF2EFC8AC9122B2C" bold="true" box="[189,324,820,846]" pageId="128" pageNumber="129">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D9C92DBFF2EFCDCC90B2BFD" blockId="128.[189,1398,866,927]" pageId="128" pageNumber="129">Named after the Yamdena, the main tribe and language of the main island in the Tanimbar group. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D9C92DBFF2EFC79C8A92CBD" pageId="128" pageNumber="129" type="materials_examined">
<paragraph id="922136E09D9C92DBFF2EFC79C93A2B83" blockId="128.[189,364,967,993]" box="[189,364,967,993]" pageId="128" pageNumber="129">
<heading id="C969818C9D9C92DBFF2EFC79C93A2B83" bold="true" box="[189,364,967,993]" fontSize="11" level="3" pageId="128" pageNumber="129" reason="3">
<emphasis id="A0EAEAF29D9C92DBFF2EFC79C93A2B83" bold="true" box="[189,364,967,993]" pageId="128" pageNumber="129">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D9C92DBFF79FC48C90A2C72" blockId="128.[189,1399,1014,1111]" box="[234,348,1014,1040]" pageId="128" pageNumber="129">
<emphasis id="A0EAEAF29D9C92DBFF79FC48C90A2C72" bold="true" box="[234,348,1014,1040]" pageId="128" pageNumber="129">
<typeStatus id="4D2588429D9C92DBFF79FC48C90A2C72" box="[234,348,1014,1040]" pageId="128" pageNumber="129" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D9C92DBFF2EFBA7C94B2C35" blockId="128.[189,1399,1014,1111]" pageId="128" pageNumber="129">
<materialsCitation id="22F63CBD9D9C92DBFF2EFBA7C94B2C35" ID-GBIF-Occurrence="2898878370" collectingDate="2009-01" collectingDateMax="2077" collectingDateMin="2009-01" collectionCode="BMNH" country="Indonesia" location="Tanimbar Islands" pageId="128" pageNumber="129" specimenCount="1" typeStatus="holotype">
<collectingCountry id="EA8976709D9C92DBFF2EFBA7C90D2C51" box="[189,347,1049,1075]" name="Indonesia" pageId="128" pageNumber="129">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D9C92DBFEE9FBA4C9C42C56" box="[378,402,1050,1076]" pageId="128" pageNumber="129">♂</specimenCount>
; “
<collectingCountry id="EA8976709D9C92DBFE3EFBA7CA182C51" box="[429,590,1049,1075]" name="Indonesia" pageId="128" pageNumber="129">INDONESIA</collectingCountry>
<location id="9741603B9D9C92DBFDCBFBA7CB702C51" LSID="urn:lsid:plazi:treatment:1A3787F69D9C92DAFDFBFE4ACD202E5A:9741603B9D9C92DBFDCBFBA7CB702C51" box="[600,806,1049,1075]" country="Indonesia" name="Tanimbar Islands" pageId="128" pageNumber="129">Tanimbar Islands</location>
Yamdena Is.
<date id="E62010209D9C92DBFC5DFBA4CC392C51" box="[974,1135,1049,1076]" pageId="128" pageNumber="129" value="2009-01">
<collectingDate id="F664E9C89D9C92DBFC5DFBA4CC392C51" box="[974,1135,1049,1076]" pageId="128" pageNumber="129" value="2009-01">January 2009</collectingDate>
</date>
<collectionCode id="F48FAE259D9C92DBFBEAFBA7CC8D2C56" box="[1145,1243,1049,1076]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="128" pageNumber="129">BMNH</collectionCode>
{E}
<date id="E62010209D9C92DBFA93FBA7CD312C56" box="[1280,1383,1049,1076]" pageId="128" pageNumber="129" value="2015" valueMax="2077">
<collectingDate id="F664E9C89D9C92DBFA93FBA7CD312C56" box="[1280,1383,1049,1076]" pageId="128" pageNumber="129" value="2015" valueMax="2077">2015-77</collectingDate>
</date>
”;
<collectionCode id="F48FAE259D9C92DBFF2EFB83C94F2C35" box="[189,281,1085,1111]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="128" pageNumber="129">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D9C92DBFF79FBC0CA742CFB" blockId="128.[189,1398,1150,1247]" box="[234,546,1150,1177]" pageId="128" pageNumber="129">
<emphasis id="A0EAEAF29D9C92DBFF79FBC0C9312CFA" bold="true" box="[234,359,1150,1176]" pageId="128" pageNumber="129">
<typeStatus id="4D2588429D9C92DBFF79FBC0C9312CFA" box="[234,359,1150,1176]" pageId="128" pageNumber="129" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="8498FD699D9C92DBFEEBFBC0CA4C2CFB" box="[376,538,1150,1177]" pageId="128" pageNumber="129" type="generic">28 specimens</specimenCount>
)
</paragraph>
<paragraph id="922136E09D9C92DBFF2EFB1FC8A92CBD" blockId="128.[189,1398,1150,1247]" pageId="128" pageNumber="129">
<materialsCitation id="22F63CBD9D9C92DBFF2EFB1FCBC02CDE" ID-GBIF-Occurrence="2898878501" box="[189,918,1185,1212]" collectionCode="BMNH" country="Indonesia" location="INDONESIA" pageId="128" pageNumber="129" specimenCount="1" typeStatus="holotype">
<collectingCountry id="EA8976709D9C92DBFF2EFB1FC90D2CD9" box="[189,347,1185,1211]" name="Indonesia" pageId="128" pageNumber="129">INDONESIA</collectingCountry>
• 26 specs; same label as for
<typeStatus id="4D2588429D9C92DBFD55FB1FCB662CD9" box="[710,816,1185,1211]" pageId="128" pageNumber="129" type="holotype">holotype</typeStatus>
;
<collectionCode id="F48FAE259D9C92DBFCADFB1CCBC02CDE" box="[830,918,1186,1212]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="128" pageNumber="129">BMNH</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D9C92DBFC20FB1FC8AD2CBD" ID-GBIF-Occurrence="2898878537" collectionCode="DTC" country="Indonesia" pageId="128" pageNumber="129" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="8498FD699D9C92DBFC20FB1FCBB42CDE" box="[947,994,1185,1212]" pageId="128" pageNumber="129" type="male">1 ♂</specimenCount>
,
<specimenCount id="8498FD699D9C92DBFC61FB1FCC772CDE" box="[1010,1057,1185,1212]" pageId="128" pageNumber="129" type="female">1 ♀</specimenCount>
; same label as for
<typeStatus id="4D2588429D9C92DBFA9BFB1FCD242CD9" box="[1288,1394,1185,1211]" pageId="128" pageNumber="129" type="holotype">holotype</typeStatus>
;
<collectionCode id="F48FAE259D9C92DBFF2EFB7BC8AD2CBD" box="[189,251,1221,1247]" collectionName="Dmitrijs Telnovs" pageId="128" pageNumber="129">DTC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D9C92DAFF2EFAB8C98E2D34" lastPageId="129" lastPageNumber="130" pageId="128" pageNumber="129" type="description">
<paragraph id="922136E09D9C92DBFF2EFAB8C9182D42" blockId="128.[189,334,1286,1312]" box="[189,334,1286,1312]" pageId="128" pageNumber="129">
<heading id="C969818C9D9C92DBFF2EFAB8C9182D42" bold="true" box="[189,334,1286,1312]" fontSize="11" level="3" pageId="128" pageNumber="129" reason="3">
<emphasis id="A0EAEAF29D9C92DBFF2EFAB8C9182D42" bold="true" box="[189,334,1286,1312]" pageId="128" pageNumber="129">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D9C92DBFF2EFA8BCACA2DF7" blockId="128.[189,1399,1332,1430]" pageId="128" pageNumber="129">
MEASUREMENTS.
<typeStatus id="4D2588429D9C92DBFEEFFA8BC9BA2D2D" box="[380,492,1333,1359]" pageId="128" pageNumber="129" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="55669B059D9C92DBFD54FA8ACB652D2D" box="[711,819,1332,1359]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" pageId="128" pageNumber="129" unit="mm" value="8.05">8.05 mm</quantity>
. Head
<quantity id="55669B059D9C92DBFC16FA8ACBB82D2D" box="[901,1006,1332,1359]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="128" pageNumber="129" unit="mm" value="1.45">1.45 mm</quantity>
long, across eyes
<quantity id="55669B059D9C92DBFB55FA8BCD792D2D" box="[1222,1327,1333,1359]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="128" pageNumber="129" unit="mm" value="1.9">1.90 mm</quantity>
wide. Pronotum
<quantity id="55669B059D9C92DBFEA4FAE6C9C92D10" box="[311,415,1368,1394]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="128" pageNumber="129" unit="mm" value="1.8">1.80 mm</quantity>
long, maximum width
<quantity id="55669B059D9C92DBFD3EFAE6CB412D10" box="[685,791,1368,1394]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="128" pageNumber="129" unit="mm" value="2.8">2.80 mm</quantity>
. Elytral length
<quantity id="55669B059D9C92DBFC54FAE6CC642D10" box="[967,1074,1368,1394]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="128" pageNumber="129" unit="mm" value="4.8">4.80 mm</quantity>
, maximum combined width
<quantity id="55669B059D9C92DBFF2EFAC5C97F2DF4" box="[189,297,1403,1430]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="128" pageNumber="129" unit="mm" value="3.8">3.80 mm</quantity>
.
<typeStatus id="4D2588429D9C92DBFEA7FAC2C9F02DF4" box="[308,422,1404,1430]" pageId="128" pageNumber="129" type="paratype">Paratypes</typeStatus>
up to
<quantity id="55669B059D9C92DBFE63FAC5CA0F2DF4" box="[496,601,1403,1430]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="128" pageNumber="129" unit="mm" value="9.2">9.20 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D9C92DAFF2EFA03CA0C2C19" blockId="128.[189,1399,1469,2025]" lastBlockId="129.[189,1399,265,1147]" lastPageId="129" lastPageNumber="130" pageId="128" pageNumber="129">
Dorsum and venter uniformly black-brown with reddish brown labroclypeus, mouthparts, antennae, tibiae and tarsi. Head transverse, glossy dorsally and ventrally, somewhat convex between eyes and vaguely impressed on labroclypeus anterior to each compound eye. Compound eye large, occupying about half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D9C92DBFD20F999CB432E23" box="[691,789,1574,1601]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="128" pageNumber="129">Fig. 195</figureCitation>
) broadly and moderately deep emarginate anteriorly, its lateral margins sinuous in both dorsal and lateral views, anterolateral angles moderately protruding, to slightly acute in dorsal view, bent up at ~90° to frons in lateral view. Female labroclypeus (
<figureCitation id="0AA52A659D9C92DBFA99F9D3CD382EEA" box="[1290,1390,1645,1672]" captionStart="Figs 193–207" captionStartId="145.[189,241,1716,1742]" captionTargetBox="[385,1203,267,1681]" captionTargetId="figure-159@145.[385,1203,265,1694]" captionTargetPageId="145" captionText="Figs 193–207. Papuan and Wallacean Maechidius Macleay, 1819, head (193–196) and pronotum (197– 207), dorsal view. 193. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 194. Same, ♂ (SNSD). 195. M. yamdena sp. nov., paratype, ♂ (BMNH). 196. Same, paratype, ♀ (BMNH). 197. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 198. M. aiyura sp. nov., holotype, ♂ (BMNH). 199. M. alesbezdeki sp. nov., holotype, ♂ (NME). 200. M. angusticeps Arrow, 1941, holotype, ♂ (BMNH). 201. M. aroae Heller, 1914, lectotype, ♀ (SNSD). 202. M. awu sp. nov., holotype, ♂ (IECA). 203. M. babyrousa sp. nov., holotype, ♂ (BMNH). 204. M. bintang sp. nov., holotype, ♂ (NME). 205. M. boessnecki sp. nov., holotype, ♂ (SNSD). 206. M. brocki sp. nov., holotype, ♂ (BMNH). 207. M. crypticus sp. nov., paratype, ♀ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122189" httpUri="https://zenodo.org/record/4122189/files/figure.png" pageId="128" pageNumber="129">Fig. 196</figureCitation>
) somewhat shallower emarginate anteriorly, with anteriorly less protruding anterolateral angles. Upper- and underside of labroclypeus with sparse long setae along anterior and lateral margins. Canthus obtusely angulate in dorsal view. Anterior and lateral margins of labroclypeus smooth. Head punctures irregularly shaped, moderately deep, variably large. Intervening spaces glossy and glabrous, variably large, generally smaller than punctures. Moderately long suberect seta rises from anterior margin of each puncture, surpassing its length. Antenna 8-segmented, club 3-lamellate. Scape large, with bulbous predistal projection on dorsal side, provided with two very long setae near distal margin. Antennomere 2 bulbous, subspherical. Pronotum glossy dorsally and laterally. Anterior margin of pronotum slightly sinuous with anterolateral angles (stronger) and mesal portion (inconspicuously) protruding anteriad. Basal margin of pronotum very broadly rounded. Lateral margin of pronotum evenly rounded, shallowly emarginate in front of posterolateral angles (
<figureCitation id="0AA52A659D9D92DAFD47FEB7CB6B2946" box="[724,829,265,292]" captionStart="Figs 259–265" captionStartId="149.[189,241,1372,1398]" captionTargetBox="[189,1379,265,1308]" captionTargetId="figure-88@149.[189,1398,265,1311]" captionTargetPageId="149" captionText="Figs 259–265. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 259. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 260. M. trivialis sp. nov., holotype, ♂ (BMNH). 261. M. ursus sp. nov., holotype, ♀ (NME). 262. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 263. M. weigeli sp. nov., holotype, ♂ (NME). 264. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 265. M. yamdena sp. nov., paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122198" httpUri="https://zenodo.org/record/4122198/files/figure.png" pageId="129" pageNumber="130">Fig. 265</figureCitation>
). Crenulae of lateral margin delicate, short seta present between every two crenulae. Lateral margin of pronotum slightly sinuous in lateral view. Pronotal punctures ovoid (more circular along lateral sides), horseshoe-shaped, variably large, deep. Intervening spaces glossy and glabrous, variably large but generally much smaller than punctures. Pronotal setae inconspicuous, suberect, shorter than those on head, not or hardly surpassing length of corresponding punctures, appressed to median (raised) part of corresponding horseshoe-shaped puncture. Hypomeron slightly curved and very long setose on anterior margin which is moderately raised. Antennal pocket moderately deep. Median anterior process of prosternum long brushy setose, broad, moderately raised. Scutellar shield rounded apically. Elytra slightly widened in posterior half, maximum width across apical third, slightly glossy dorsally, with distinct humeri. Inconspicuous obtuse transverse hump at apex of each elytron. Tracks of four to five inconspicuous flat glabrous longitudinal carinae on each elytron (including one sutural and two lateral). Elytral disc in part wrinkled, punctures sinuous (elongate and narrow), incision-shaped, moderately deep (
<figureCitation id="0AA52A659D9D92DAFD52FD0BCB7F2AB2" box="[705,809,693,720]" captionStart="Figs 349–358" captionStartId="156.[189,241,1786,1812]" captionTargetBox="[287,1299,265,1743]" captionTargetId="figure-123@156.[287,1301,265,1760]" captionTargetPageId="156" captionText="Figs 349–358. Papuan and Wallacean Maechidius Macleay, 1819, elytron (349–356) and pronotal setation (357–358), dorsal view. 349. M. suwawa sp. nov., paratype, ♀ (BMNH). 350. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 351. M. trivialis sp. nov., holotype, ♂ (BMNH). 352. M. ursus sp. nov., holotype, ♀ (NME). 353. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 354. M. weigeli sp. nov., holotype, ♂ (NME). 355. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 356. M. yamdena sp. nov., paratype, ♂ (BMNH). 357. M. leucopsar sp. nov., holotype, ♂ (IECA). 358. M. ursus sp. nov., holotype, ♀ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122214" httpUri="https://zenodo.org/record/4122214/files/figure.png" pageId="129" pageNumber="130">Fig. 356</figureCitation>
). Elytral setae inconspicuous, appressed except on apices; seta rises from anterior margin of each puncture, not surpassing length of corresponding incision. Upper carina of epipleuron all along with short suberect setae. Male and female pygidium flat to slightly convex, with large irregularly shaped shallow annular punctures and short almost erect setae, which rise from centre of each puncture (Fig. 534). Venter covered with sparse large shallow annular punctures, each provided with short seta. Legs long and slender. Protibia nearly straight on external margin, with delicate dorsal carina. Three obtuse external teeth on protibia: distal narrower, median somewhat larger and broader, basal one less prominent (
<figureCitation id="0AA52A659D9D92DAFDECFC10CAB22BAA" box="[639,740,942,968]" captionStart="Figs 424–435" captionStartId="161.[189,241,1394,1420]" captionTargetBox="[238,1349,265,1318]" captionTargetId="figure-136@161.[217,1354,265,1322]" captionTargetPageId="161" captionText="Figs 424–435. Papuan and Wallacean Maechidius Macleay, 1819, protibia, dorsal view (424–428), and abdominal ventrites (429–435), ventral and lateral views. 424. M. ursus sp. nov., holotype, ♀ (NME). 425. M. vicinus Heller, 1914, lectotype, ♀ (SNSD). 426. M. weigeli sp. nov., holotype, ♂ (NME). 427. M. woodlarkianus Heller, 1914, lectotype, ♀ (SNSD). 428. M. yamdena sp. nov., paratype, ♂ (BMNH). 429. M.dani sp. nov., holotype, ♂ (IECA). 430. Same, different aspect (IECA). 431. M. hamatus sp. nov., holotype, ♂ (IECA). 432. Same, different aspect (IECA). 433. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 434. M. subcostatus Heller, 1895, lectotype, ♂ (SNSD). 435. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122216" httpUri="https://zenodo.org/record/4122216/files/figure.png" pageId="129" pageNumber="130">Fig. 428</figureCitation>
), nearly disappearing in some males. Basal protarsomere lengthened, asymmetrical, distinctly widened distally, concave ventrally, with short acute ventro-lateral denticle at inner distal margin which is larger in male than in female (
<figureCitation id="0AA52A659D9D92DAFC61FC4BCC0C2C72" box="[1010,1114,1013,1040]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="129" pageNumber="130">Fig. 447</figureCitation>
). Protibial terminal spur short, straight, acute (
<figureCitation id="0AA52A659D9D92DAFE2BFBA7CA082C51" box="[440,606,1049,1075]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="129" pageNumber="130">Figs 447–448</figureCitation>
). Metatibial terminal spurs paired, acute in male (
<figureCitation id="0AA52A659D9D92DAFB0EFBA7CD552C51" box="[1181,1283,1049,1075]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="129" pageNumber="130">Fig. 442</figureCitation>
), rounded in female (
<figureCitation id="0AA52A659D9D92DAFEACFB83C9FF2C35" box="[319,425,1085,1111]" captionStart="Figs 436–450" captionStartId="162.[189,241,1619,1645]" captionTargetBox="[205,1349,265,1554]" captionTargetId="figure-184@162.[205,1377,265,1557]" captionTargetPageId="162" captionText="Figs 436–450. Papuan and Wallacean Maechidius Macleay, 1819, legs, various views. 436. M. hirtipes Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 437. M. tarsalis Arrow, 1941, lectotype, ♂, basal metatarsomere (BMNH). 438. M. babyrousa sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 439. Same, paratype, ♂ (BMNH). 440. Same, different aspect (BMNH). 441. M. yamdena sp. nov., paratype, ♀, metatibial terminal spurs (BMNH). 442. Same, paratype, ♂ (BMNH). 443. M. longipes sp. nov., holotype, ♂, metatibial terminal spurs (BMNH). 444. M. ursus sp. nov., holotype, ♀, left metatibia and basal metatarsomere (NME). 445. M. dani sp. nov., holotype, ♂, apical portion of metatibia (IECA). 446. M. skalei sp. nov., holotype, ♂, apical portion of metatibia and basal metatarsomere (NME). 447. M. yamdena sp. nov., paratype, ♂, protibial terminal spur (BMNH). 448. Same, paratype, ♀ (BMNH). 449. M. babyrousa sp. nov., paratype, ♀, protibial terminal spur (BMNH). 450. Same, paratype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122218" httpUri="https://zenodo.org/record/4122218/files/figure.png" pageId="129" pageNumber="130">Fig. 441</figureCitation>
). Tarsal claws in both sexes with large pulvilli. Spiculum gastrale as in
<figureCitation id="0AA52A659D9D92DAFA98FB83CD252C3A" box="[1291,1395,1085,1112]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="129" pageNumber="130">Fig. 547</figureCitation>
. Male aedeagus as in
<figureCitation id="0AA52A659D9D92DAFE22FBDECA032C19" box="[433,597,1120,1147]" captionStart="Figs 739–750" captionStartId="181.[189,241,1892,1918]" captionTargetBox="[360,1207,271,1867]" captionTargetId="figure-66@181.[347,1240,265,1871]" captionTargetPageId="181" captionText="Figs 739–750. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 739– 741. M. weigeli sp. nov., paratype, ♂ (NME). 742–744. M. woodlarkianus Heller, 1914, ♂ (SNSD). 745–747. M. yamdena sp. nov., paratype, ♂ (BMNH). 748–750. M. paupianus Heller, 1910 (paratype ♂ of M. arrowi Frey, 1969) (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122256" httpUri="https://zenodo.org/record/4122256/files/figure.png" pageId="129" pageNumber="130">Figs 745–747</figureCitation>
.
</paragraph>
<paragraph id="922136E09D9D92DAFF2EFB1CC9E62CDE" blockId="129.[189,432,1186,1212]" box="[189,432,1186,1212]" pageId="129" pageNumber="130">
<heading id="C969818C9D9D92DAFF2EFB1CC9E62CDE" bold="true" box="[189,432,1186,1212]" fontSize="11" level="3" pageId="129" pageNumber="130" reason="3">
<emphasis id="A0EAEAF29D9D92DAFF2EFB1CC9E62CDE" bold="true" box="[189,432,1186,1212]" pageId="129" pageNumber="130">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D9D92DAFF2EFB6FC98E2D34" blockId="129.[189,1399,1233,1366]" pageId="129" pageNumber="130">Poorly indicated. Female generally more robust than male, with comparatively smaller lamellae of antennal club, anterior margin of labroclypeus shallower emarginate, somewhat longer protibial terminal spur and dorsally flat pygidium. The ventro-lateral denticle of the basal protarsomere is larger in the male than in the female.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D9D92DAFF2EFAC0C9C42DA5" box="[189,402,1406,1480]" pageId="129" pageNumber="130" type="biology_ecology">
<paragraph id="922136E09D9D92DAFF2EFAC0C9762DFA" blockId="129.[189,288,1406,1432]" box="[189,288,1406,1432]" pageId="129" pageNumber="130">
<heading id="C969818C9D9D92DAFF2EFAC0C9762DFA" bold="true" box="[189,288,1406,1432]" fontSize="11" level="3" pageId="129" pageNumber="130" reason="3">
<emphasis id="A0EAEAF29D9D92DAFF2EFAC0C9762DFA" bold="true" box="[189,288,1406,1432]" pageId="129" pageNumber="130">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D9D92DAFF2EFA10C9C42DA5" blockId="129.[189,402,1453,1480]" box="[189,402,1453,1480]" pageId="129" pageNumber="130">No data available.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D9D92DAFF2EFA51CD202E5A" box="[189,1398,1519,1592]" pageId="129" pageNumber="130" type="distribution">
<paragraph id="922136E09D9D92DAFF2EFA51C9002E6B" blockId="129.[189,342,1519,1545]" box="[189,342,1519,1545]" pageId="129" pageNumber="130">
<heading id="C969818C9D9D92DAFF2EFA51C9002E6B" bold="true" box="[189,342,1519,1545]" fontSize="11" level="3" pageId="129" pageNumber="130" reason="3">
<emphasis id="A0EAEAF29D9D92DAFF2EFA51C9002E6B" bold="true" box="[189,342,1519,1545]" pageId="129" pageNumber="130">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D9D92DAFF2EF9A0CD202E5A" blockId="129.[189,1398,1566,1592]" box="[189,1398,1566,1592]" pageId="129" pageNumber="130">
Hitherto only known from Yamdena, the main island of the Tanimbar Islands, SE
<collectingCountry id="EA8976709D9D92DAFBEFF9A0CCA72E5A" box="[1148,1265,1566,1592]" name="Indonesia" pageId="129" pageNumber="130">Moluccas</collectingCountry>
,
<collectingCountry id="EA8976709D9D92DAFB6FF9A0CD242E5A" box="[1276,1394,1566,1592]" name="Indonesia" pageId="129" pageNumber="130">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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