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<document id="97DEF8D7F5FF50AD27ED2BE0665464A1" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D5F921FFDBBFC73C9F52BC7" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius lineatopunctatus Frey 1969" docType="treatment" docVersion="10" lastPageNumber="69" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="68" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="65248229FD6025F3B9BC4364D7433D32" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E8ADA85CBA9AFB22A424291AE3AAA9CA">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="3C906034D803595956D34BBAA20F2F1E">Telnov, Dmitry</mods:namePart>
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<mods:title id="B42396FE3D4C11602B9A9E37D267C5A2">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="65F66492284DC070916A34015BEB6B27">
<mods:date id="716367D9A356436FC46554DFC6120E9A">2020</mods:date>
<mods:detail id="71A2E6672478B185D97C83781B464BA8" type="pubDate">
<mods:number id="733E7758226EE4AD77A4142F2036FA1F">2020-10-19</mods:number>
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<mods:number id="D833D31773CBE1E2F93EDAF836D5420F">721</mods:number>
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<mods:classification id="9DE421A2921DE26670F32B876A75B65A">journal article</mods:classification>
<mods:identifier id="2D6D16A84FC19AA5D2A0BE280C61B2C0" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="C0B7B6900BDAEB61839827B9A1CA5055" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
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<treatment id="1A3787F69D5F921FFDBBFC73C9F52BC7" ID-DOI="http://doi.org/10.5281/zenodo.4344473" ID-GBIF-Taxon="168648175" ID-Zenodo-Dep="4344473" LSID="urn:lsid:plazi:treatment:1A3787F69D5F921FFDBBFC73C9F52BC7" httpUri="http://treatment.plazi.org/id/1A3787F69D5F921FFDBBFC73C9F52BC7" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
<subSubSection id="DA84656B9D5F9218FDBBFC73CC5A2B8A" box="[552,1036,973,1000]" pageId="67" pageNumber="68" type="nomenclature">
<paragraph id="922136E09D5F9218FDBBFC73CC5A2B8A" blockId="67.[515,1072,973,1036]" box="[552,1036,973,1000]" pageId="67" pageNumber="68">
<heading id="C969818C9D5F9218FDBBFC73CC5A2B8A" box="[552,1036,973,1000]" centered="true" fontSize="11" level="2" pageId="67" pageNumber="68" reason="2">
<taxonomicName id="559E4D639D5F9218FDBBFC73CC5A2B8A" ID-CoL="3XFJX" authority="Frey, 1969" authorityName="Frey" authorityYear="1969" box="[552,1036,973,1000]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lineatopunctatus">
<emphasis id="A0EAEAF29D5F9218FDBBFC73CBD12B8A" bold="true" box="[552,903,973,1000]" italics="true" pageId="67" pageNumber="68">Maechidius lineatopunctatus</emphasis>
<bibRefCitation id="F60F4B119D5F9218FC1DFC70CC5A2B8A" author="Frey G." box="[910,1036,974,1000]" pageId="67" pageNumber="68" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey, 1969</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5F9218FD90FC4FCC662C69" box="[515,1072,1009,1036]" pageId="67" pageNumber="68" type="description">
<paragraph id="922136E09D5F9218FD90FC4FCC662C69" blockId="67.[515,1072,973,1036]" box="[515,1072,1009,1036]" pageId="67" pageNumber="68">
<figureCitation id="0AA52A659D5F9218FD90FC4FCA0B2C6E" box="[515,605,1009,1036]" captionStart="Figs 50–53" captionStartId="62.[189,241,1927,1953]" captionTargetBox="[326,1248,265,1865]" captionTargetId="figure-65@62.[324,1254,249,1883]" captionTargetPageId="62" captionText="Figs 50–53. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 50. M. lapsus sp. nov., holotype, ♂ (BMNH). 51. M. legalovi sp. nov., holotype, ♂ (BMNH). 52. M. leucopsar sp. nov., holotype, ♂ (IECA). 53. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122148" httpUri="https://zenodo.org/record/4122148/files/figure.png" pageId="67" pageNumber="68">Figs 53</figureCitation>
,
<figureCitation id="0AA52A659D5F9218FDFAFC4FCAC12C6E" box="[617,663,1009,1036]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="67" pageNumber="68">135</figureCitation>
,
<figureCitation id="0AA52A659D5F9218FD37FC4FCA852C69" box="[676,723,1009,1035]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="67" pageNumber="68">226</figureCitation>
,
<figureCitation id="0AA52A659D5F9218FD4CFC4FCB582C69" box="[735,782,1009,1035]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="67" pageNumber="68">270</figureCitation>
,
<figureCitation id="0AA52A659D5F9218FC8AFC4FCB1E2C69" box="[793,840,1009,1035]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="67" pageNumber="68">317</figureCitation>
, 390, 486, 646–648
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5F9218FF2EFB8CCB602D2A" pageId="67" pageNumber="68" type="materials_examined">
<paragraph id="922136E09D5F9218FF2EFB8CC93A2C2E" blockId="67.[189,364,1074,1100]" box="[189,364,1074,1100]" pageId="67" pageNumber="68">
<heading id="C969818C9D5F9218FF2EFB8CC93A2C2E" bold="true" box="[189,364,1074,1100]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="A0EAEAF29D5F9218FF2EFB8CC93A2C2E" bold="true" box="[189,364,1074,1100]" pageId="67" pageNumber="68">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5F9218FF79FBDEC90A2C18" blockId="67.[189,1398,1120,1217]" box="[234,348,1120,1146]" pageId="67" pageNumber="68">
<emphasis id="A0EAEAF29D5F9218FF79FBDEC90A2C18" bold="true" box="[234,348,1120,1146]" pageId="67" pageNumber="68">
<typeStatus id="4D2588429D5F9218FF79FBDEC90A2C18" box="[234,348,1120,1146]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D5F9218FF2EFB3DCC542CA3" blockId="67.[189,1398,1120,1217]" pageId="67" pageNumber="68">
<materialsCitation id="22F63CBD9D5F9218FF2EFB3DCC542CA3" ID-GBIF-Occurrence="2898878323" collectionCode="NHMB" collectorName="New-Guinea P. Voss" country="Papua New Guinea" determinerName="G. Frey" location="Port Moresby" pageId="67" pageNumber="68" specimenCount="1" stateProvince="Port Moresby" typeStatus="type">
<collectingCountry id="EA8976709D5F9218FF2EFB3DC9B52CFF" box="[189,483,1155,1182]" name="Papua New Guinea" pageId="67" pageNumber="68">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D5F9218FD95FB3ACA482CFC" box="[518,542,1156,1182]" pageId="67" pageNumber="68" type="male">♂</specimenCount>
; “
<collectingRegion id="505AF8029D5F9218FDA9FB3DCAB52CFF" box="[570,739,1155,1181]" country="Papua New Guinea" name="National Capital" pageId="67" pageNumber="68">Port Moresby</collectingRegion>
<collectorName id="3F6B53369D5F9218FD7CFB3DCBBF2CFC" box="[751,1001,1155,1182]" pageId="67" pageNumber="68">New-Guinea P. Voss</collectorName>
[h] //
<typeStatus id="4D2588429D5F9218FBBCFB3ACC2E2CFC" box="[1071,1144,1156,1182]" pageId="67" pageNumber="68">TYPE</typeStatus>
[p, red label] //
<typeStatus id="4D2588429D5F9218FAAFFB3ACD202CFC" box="[1340,1398,1156,1182]" pageId="67" pageNumber="68">Type</typeStatus>
<taxonomicName id="559E4D639D5F9218FF2EFB18CA472CA2" authorityName="Frey" authorityYear="1969" box="[189,529,1190,1216]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lineatopunctatus">
<emphasis id="A0EAEAF29D5F9218FF2EFB18CA472CA2" box="[189,529,1190,1216]" italics="true" pageId="67" pageNumber="68">Maechidius lineatopunctatus</emphasis>
</taxonomicName>
n sp [h] det.
<determinerName id="67FDDA249D5F9218FD30FB18CAA02CA2" box="[675,758,1190,1216]" pageId="67" pageNumber="68">G.Frey</determinerName>
,1967/68 [p]”;
<collectionCode id="F48FAE259D5F9218FC31FB19CBA82CA3" box="[930,1022,1191,1217]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="67" pageNumber="68">NHMB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D5F9218FF79FB56C9312D60" blockId="67.[189,1399,1256,1352]" box="[234,359,1256,1282]" pageId="67" pageNumber="68">
<emphasis id="A0EAEAF29D5F9218FF79FB56C9312D60" bold="true" box="[234,359,1256,1282]" pageId="67" pageNumber="68">
<typeStatus id="4D2588429D5F9218FF79FB56C9312D60" box="[234,359,1256,1282]" pageId="67" pageNumber="68" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D5F9218FF2EFAB4CB602D2A" blockId="67.[189,1399,1256,1352]" pageId="67" pageNumber="68">
According to the original description, there are 17 ♂
<specimenCount id="8498FD699D5F9218FCB7FAB5CB6E2D47" box="[804,824,1291,1317]" pageId="67" pageNumber="68" type="female">♀</specimenCount>
at
<collectionCode id="F48FAE259D5F9218FCC8FAB5CBE12D47" box="[859,951,1291,1317]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="67" pageNumber="68">NHMB</collectionCode>
. I had the opportunity to examine four of them which are referred below to
<taxonomicName id="559E4D639D5F9218FDF9FA90CA822D25" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[618,724,1325,1351]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="mailu" status="sp. nov.">
<emphasis id="A0EAEAF29D5F9218FDF9FA90CA822D25" box="[618,724,1325,1351]" italics="true" pageId="67" pageNumber="68">M. mailu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D5F9218FD48FA90CB602D2A" box="[731,822,1326,1352]" pageId="67" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5F9218FF2EFAD1CCF62DD5" box="[189,1184,1391,1464]" pageId="67" pageNumber="68" type="discussion">
<paragraph id="922136E09D5F9218FF2EFAD1C9792DEB" blockId="67.[189,303,1391,1417]" box="[189,303,1391,1417]" pageId="67" pageNumber="68">
<heading id="C969818C9D5F9218FF2EFAD1C9792DEB" bold="true" box="[189,303,1391,1417]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="A0EAEAF29D5F9218FF2EFAD1C9792DEB" bold="true" box="[189,303,1391,1417]" pageId="67" pageNumber="68">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5F9218FF2EFA23CCF62DD5" blockId="67.[189,1184,1437,1464]" box="[189,1184,1437,1464]" pageId="67" pageNumber="68">
<bibRefCitation id="F60F4B119D5F9218FF2EFA23C9D22DDA" author="Frey G." box="[189,388,1437,1464]" pageId="67" pageNumber="68" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey (1969: 502)</bibRefCitation>
designated the single specimen from
<collectingRegion id="505AF8029D5F9218FCACFA23CB892DD5" box="[831,991,1437,1463]" country="Papua New Guinea" name="National Capital" pageId="67" pageNumber="68">Port Moresby</collectingRegion>
as the
<typeStatus id="4D2588429D5F9218FBA1FA23CCCB2DD5" box="[1074,1181,1437,1463]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D5F921FFF2EFA60C9F52BC7" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68" type="description">
<paragraph id="922136E09D5F9218FF2EFA60C9182D9A" blockId="67.[189,334,1502,1528]" box="[189,334,1502,1528]" pageId="67" pageNumber="68">
<heading id="C969818C9D5F9218FF2EFA60C9182D9A" bold="true" box="[189,334,1502,1528]" fontSize="11" level="3" pageId="67" pageNumber="68" reason="3">
<emphasis id="A0EAEAF29D5F9218FF2EFA60C9182D9A" bold="true" box="[189,334,1502,1528]" pageId="67" pageNumber="68">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D5F9218FF2EF9B3CA8D2E44" blockId="67.[189,731,1548,1575]" box="[189,731,1548,1575]" pageId="67" pageNumber="68">
MEASUREMENTS.
<typeStatus id="4D2588429D5F9218FEE8F9B2C9BE2E44" box="[379,488,1548,1574]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
male
<quantity id="55669B059D5F9218FDA3F9B2CACF2E45" box="[560,665,1548,1575]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="67" pageNumber="68" unit="mm" value="6.7">6.70 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D5F921FFF2EF9F0CA202B56" blockId="67.[189,1399,1614,2025]" lastBlockId="68.[189,1399,265,821]" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
Dorsum and venter uniformly black-brown, appendages and labroclypeus castaneous brown. Head flattened dorsally on frons, glossy dorsally and ventrally. Male labroclypeus (
<figureCitation id="0AA52A659D5F9218FBE0F9CFCC8D2EE9" box="[1139,1243,1648,1675]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="67" pageNumber="68">Fig. 135</figureCitation>
) subtruncate anteriorly, its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles broadly rounded, not protruding. Upper- and underside of labroclypeus with sparse inconspicuous setae along anterior and lateral margins. Canthus very broadly rounded in dorsal view. Punctures of frons circular to ovoid, moderately deep and dense. Intervening spaces glossy, variably large. Head setae minute, appressed to suberect, not surpassing length of corresponding punctures. Antenna 9-segmented, club 3-lamellate. Pronotum strongly transverse, glossy dorsally and laterally. Anterior margin of pronotum sinuous with slightly protruding anterolateral angles. Basal margin of pronotum broadly rounded. Lateral margin of pronotum in dorsal view nearly straight, stronger constricted anteriad and straight at base, crenulate all along (
<figureCitation id="0AA52A659D5F9218FE7CF815CA0F2FA4" box="[495,601,1963,1990]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="67" pageNumber="68">Fig. 226</figureCitation>
). An inconspicuous short seta present between every two crenulae. Lateral margin of pronotum hardly sinuous in lateral view. Hypomeron separated from prosternum by moderately high straight carina which is shortly flange-like protruding in posterior half, obtusely angulate medially, with long setae on its anterolateral margin opposite to compound eye (
<figureCitation id="0AA52A659D58921FFB6CFE92CD3C2924" box="[1279,1386,300,326]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="68" pageNumber="69">Fig. 270</figureCitation>
). Antennal pocket moderately deep. Punctures of pronotal disc ovoid, moderately deep and dense, filled with microscopical velvety pubescence, in part horseshoe-shaped along lateral margins of pronotum. Intervening spaces glossy, variably large. Setae as those on head, very inconspicuous. Base of pronotum with very narrow area of microscopical velvety pubescence. Scutellar shield triangular, pointed apically, covered with microscopical velvety pubescence. Elytron glossy, with vague tracks of three longitudinal carinae, including sutural one. Base of elytra with narrow area covered with microscopical velvety pubescence. Punctures of elytral disc linear (long and narrow), rather deep (
<figureCitation id="0AA52A659D58921FFBB6FD9DCCDC2A5F" box="[1061,1162,547,573]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="68" pageNumber="69">Fig. 317</figureCitation>
). Intervening spaces larger than punctures, glossy. Elytral setae inconspicuous, appressed; each seta rises from anterior margin of each puncture, not surpassing length of incision. Male pygidium slightly convex dorsally, with large and shallow annular punctures (
<figureCitation id="0AA52A659D58921FFDA5FD33CAC82ACA" box="[566,670,653,680]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="68" pageNumber="69">Fig. 486</figureCitation>
). Intervening spaces glossy, generally smaller than punctures. Setae of pygidium suberect to erect, short in anterior, longer in apical half. Male protibia with two distal teeth of which basal one very inconspicuous (Fig. 390). Male protibial terminal spur straight, pointed. Male lower meso- and metatibial terminal spurs strongly curved. Tarsal claws with pulvilli (in male only?). Aedeagus as in
<figureCitation id="0AA52A659D58921FFE5DFCA4CA242B56" box="[462,626,794,820]" captionStart="Figs 632–651" captionStartId="174.[189,241,1822,1848]" captionTargetBox="[447,1137,271,1796]" captionTargetId="figure-96@174.[442,1145,265,1796]" captionTargetPageId="174" captionText="Figs 632–651. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, aedeagus, different aspects. 632–634. M. lapsus sp. nov., paratype, ♂, from Cyclops Mts. (BMNH). 635–637. Same, paratype, ♂, from Yapen Island (BMNH). 638–639. Same, paratype, ♂, from Waigeo Island (BMNH). 640– 642. M. legalovi sp. nov., holotype, ♂ (BMNH). 643–645. M. leucopsar sp. nov., paratype, ♂ (IECA). 646–648. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 649–651. M. lobaticeps Frey, 1969, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122242" httpUri="https://zenodo.org/record/4122242/files/figure.png" pageId="68" pageNumber="69">Figs 646–648</figureCitation>
.
</paragraph>
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<heading id="C969818C9D58921FFF2EFCE2C9E62B14" bold="true" box="[189,432,860,886]" fontSize="11" level="3" pageId="68" pageNumber="69" reason="3">
<emphasis id="A0EAEAF29D58921FFF2EFCE2C9E62B14" bold="true" box="[189,432,860,886]" pageId="68" pageNumber="69">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D58921FFF2EFC35C9F52BC7" blockId="68.[189,419,907,933]" box="[189,419,907,933]" pageId="68" pageNumber="69">Female is unknown</paragraph>
</subSubSection>
</treatment>
</document>
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