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<document id="6E9EE10EE2975BB850ADD7DAA16D52A7" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D569217FDE9FC0FCAE12EC5" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius maleo Telnov 2020, sp. nov." docType="treatment" docUuid="5C492B03-9C41-49A3-9D56-663F3C42574A" docUuidSource="ZooBank" docVersion="10" lastPageNumber="77" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="75" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="F24A58A6AC883FF3FF2EEB561EC08442" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="693535F215D2064CC3C3449741CED514">
<mods:title id="DE312CB8A238C4DFD5BEABB1547C2D82">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="A4FACD9592F099AEFE7F5624E646968C" type="personal">
<mods:role id="F212F14EB70311A519121CAA55A33D26">
<mods:roleTerm id="69412E59C08AB8001DCEE1C5C3746CAB">Author</mods:roleTerm>
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<mods:namePart id="A640D650F678F69F2C218100E7B06021">Telnov, Dmitry</mods:namePart>
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<mods:title id="6DE3CACD5AB21F8907B3FA8338CD0AA5">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="D65AE0EC04BB2A484A1B0CA5112B61AB">
<mods:date id="EAA2051D026526D66306D956536CB837">2020</mods:date>
<mods:detail id="04357D2A9656D8487F37763AAF075263" type="pubDate">
<mods:number id="5956A5A375110B7D52E79CD725B86950">2020-10-19</mods:number>
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<mods:detail id="CCD31944739A2ABD84A755E8BB24E94A" type="volume">
<mods:number id="921A99AE735FEDC8EED227F4AEBB9567">721</mods:number>
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<mods:classification id="0A3B9E5A9D77055FDEE8D2B1A98302B5">journal article</mods:classification>
<mods:identifier id="BE80FFF7F66C706D8A1DE867C9FB3D04" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="F8EDFA7E8BAE8BDD530D32E52D2FDC46" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="136174FDAC7527318A8B3F063314E0EB" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
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<mods:identifier id="1D4DCA19D218C743AE80829D953BBF8C" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
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<treatment id="1A3787F69D569217FDE9FC0FCAE12EC5" ID-DOI="http://doi.org/10.5281/zenodo.4344289" ID-GBIF-Taxon="168648164" ID-Zenodo-Dep="4344289" LSID="urn:lsid:zoobank.org:act:5C492B03-9C41-49A3-9D56-663F3C42574A" httpUri="http://treatment.plazi.org/id/1A3787F69D569217FDE9FC0FCAE12EC5" lastPageId="76" lastPageNumber="77" pageId="74" pageNumber="75">
<subSubSection id="DA84656B9D569211FDE9FC0FCBEC2BAE" box="[634,954,945,972]" pageId="74" pageNumber="75" type="nomenclature">
<paragraph id="922136E09D569211FDE9FC0FCBEC2BAE" blockId="74.[373,1215,945,1044]" box="[634,954,945,972]" pageId="74" pageNumber="75">
<heading id="C969818C9D569211FDE9FC0FCBEC2BAE" box="[634,954,945,972]" centered="true" fontSize="11" level="2" pageId="74" pageNumber="75" reason="2">
<taxonomicName id="559E4D639D569211FDE9FC0FCB0E2BAE" ID-CoL="7X3X5" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[634,856,945,972]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="maleo" status="sp. nov.">
<emphasis id="A0EAEAF29D569211FDE9FC0FCB0E2BAE" bold="true" box="[634,856,945,972]" italics="true" pageId="74" pageNumber="75">Maechidius maleo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D569211FCF3FC0CCBEC2BAE" box="[864,954,946,972]" pageId="74" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D569211FEE6FC6BCC662C71" pageId="74" pageNumber="75" type="description">
<paragraph id="922136E09D569211FEE6FC6BCCE92B92" blockId="74.[373,1215,945,1044]" box="[373,1215,981,1008]" pageId="74" pageNumber="75">
<uri id="E60F3AE29D569211FEE6FC6BCCE92B92" box="[373,1215,981,1008]" pageId="74" pageNumber="75">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D569211FD0BFC6BCCE92B92" box="[664,1215,981,1008]" pageId="74" pageNumber="75">5C492B03-9C41-49A3-9D56-663F3C42574A</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D569211FD90FC47CC662C71" blockId="74.[373,1215,945,1044]" box="[515,1072,1017,1044]" pageId="74" pageNumber="75">
<figureCitation id="0AA52A659D569211FD90FC47CA0B2C76" box="[515,605,1017,1044]" captionStart="Figs 58–61" captionStartId="75.[189,241,1928,1954]" captionTargetBox="[336,1249,266,1914]" captionTargetId="figure-62@75.[333,1249,257,1923]" captionTargetPageId="75" captionText="Figs 58–61. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 58. M. maleo sp. nov., holotype, ♂ (IRSN). 59. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 60. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 61. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122152" httpUri="https://zenodo.org/record/4122152/files/figure.png" pageId="74" pageNumber="75">Figs 58</figureCitation>
,
<figureCitation id="0AA52A659D569211FDFAFC47CACE2C71" box="[617,664,1017,1043]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="74" pageNumber="75">144</figureCitation>
,
<figureCitation id="0AA52A659D569211FD37FC47CA852C71" box="[676,723,1017,1043]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="74" pageNumber="75">231</figureCitation>
,
<figureCitation id="0AA52A659D569211FD4CFC47CB582C71" box="[735,782,1017,1043]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="74" pageNumber="75">322</figureCitation>
, 395, 491, 553, 661–663
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D569211FF2EFB85CD212CE7" pageId="74" pageNumber="75" type="diagnosis">
<paragraph id="922136E09D569211FF2EFB85C99F2C37" blockId="74.[189,457,1083,1109]" box="[189,457,1083,1109]" pageId="74" pageNumber="75">
<heading id="C969818C9D569211FF2EFB85C99F2C37" bold="true" box="[189,457,1083,1109]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A0EAEAF29D569211FF2EFB85C99F2C37" bold="true" box="[189,457,1083,1109]" pageId="74" pageNumber="75">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D569211FF2EFBD5CD212CE7" blockId="74.[189,1399,1131,1157]" box="[189,1399,1131,1157]" pageId="74" pageNumber="75">
<taxonomicName id="559E4D639D569211FF2EFBD5C9C32CE7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[189,405,1131,1157]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="maleo" status="sp. nov.">
<emphasis id="A0EAEAF29D569211FF2EFBD5C9C32CE7" box="[189,405,1131,1157]" italics="true" pageId="74" pageNumber="75">Maechidius maleo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D569211FE08FBD5C9A32CE7" box="[411,501,1131,1157]" pageId="74" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
is peculiar among all congeners primarily in the shape of the male aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D569211FF2EFB13CB5F2D78" pageId="74" pageNumber="75" type="etymology">
<paragraph id="922136E09D569211FF2EFB13C9122CA5" blockId="74.[189,324,1197,1223]" box="[189,324,1197,1223]" pageId="74" pageNumber="75">
<heading id="C969818C9D569211FF2EFB13C9122CA5" bold="true" box="[189,324,1197,1223]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A0EAEAF29D569211FF2EFB13C9122CA5" bold="true" box="[189,324,1197,1223]" pageId="74" pageNumber="75">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D569211FF2EFB62CB5F2D78" blockId="74.[189,1398,1244,1307]" pageId="74" pageNumber="75">
Named after the Maleo (
<taxonomicName id="559E4D639D569211FE71FB62CBE62C95" authority="S. Muller, 1846" authorityName="S. Muller" authorityYear="1846" box="[482,944,1244,1271]" class="Aves" family="Megapodiidae" genus="Macrocephalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galliformes" pageId="74" pageNumber="75" phylum="Chordata" rank="species" species="maleo">
<emphasis id="A0EAEAF29D569211FE71FB62CABD2C94" box="[482,747,1244,1270]" italics="true" pageId="74" pageNumber="75">Macrocephalon maleo</emphasis>
S. Müller, 1846
</taxonomicName>
), the enigmatic and endangered
<collectingRegion id="505AF8029D569211FAA1FB62C9712D78" country="Indonesia" name="Sulawesi Utara" pageId="74" pageNumber="75">North Sulawesi</collectingRegion>
endemic megapode. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D569211FF2EFAFCCD382E3F" pageId="74" pageNumber="75" type="materials_examined">
<paragraph id="922136E09D569211FF2EFAFCC93A2D3E" blockId="74.[189,364,1346,1372]" box="[189,364,1346,1372]" pageId="74" pageNumber="75">
<heading id="C969818C9D569211FF2EFAFCC93A2D3E" bold="true" box="[189,364,1346,1372]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A0EAEAF29D569211FF2EFAFCC93A2D3E" bold="true" box="[189,364,1346,1372]" pageId="74" pageNumber="75">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D569211FF79FACCCBB82DB6" ID-GBIF-Occurrence="2898878366" collectingDate="2008-02-18" collectionCode="NME" collectorName="A. Skale &amp; Coll. A. Skale Hof" country="Indonesia" elevation="260" latitude="1.3825" location="INDONESIA" pageId="74" pageNumber="75" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<paragraph id="922136E09D569211FF79FACCC90A2DEE" blockId="74.[189,1398,1394,1492]" box="[234,348,1394,1420]" pageId="74" pageNumber="75">
<emphasis id="A0EAEAF29D569211FF79FACCC90A2DEE" bold="true" box="[234,348,1394,1420]" pageId="74" pageNumber="75">
<typeStatus id="4D2588429D569211FF79FACCC90A2DEE" box="[234,348,1394,1420]" pageId="74" pageNumber="75" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D569211FF2EFA2BCBB82DB6" blockId="74.[189,1398,1394,1492]" pageId="74" pageNumber="75">
<collectingCountry id="EA8976709D569211FF2EFA2BC90D2DCD" box="[189,347,1429,1455]" name="Indonesia" pageId="74" pageNumber="75">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D569211FEE2FA28C9DF2DD2" box="[369,393,1430,1456]" pageId="74" pageNumber="75" type="male">♂</specimenCount>
; “
<collectingCountry id="EA8976709D569211FE0CFA2BCA162DCD" box="[415,576,1429,1455]" name="Indonesia" pageId="74" pageNumber="75">INDONESIA</collectingCountry>
<locationDeviation id="83932FC29D569211FDD4FA2BCBE92DCD" box="[583,959,1429,1455]" pageId="74" pageNumber="75">
N-Sulawesi
<quantity id="55669B059D569211FD46FA2BCB7E2DCD" box="[725,808,1429,1455]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="74" pageNumber="75" unit="km" value="1.5" valueMax="2.0" valueMin="1.0">1-2 km</quantity>
S Airmadidi
</locationDeviation>
,
<quantity id="55669B059D569211FC5AFA2BCC5D2DCD" box="[969,1035,1429,1455]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="74" pageNumber="75" unit="m" value="260.0">
<elevation id="19B3D1D39D569211FC5AFA2BCC5D2DCD" box="[969,1035,1429,1455]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="74" pageNumber="75" unit="m" value="260.0">260m</elevation>
</quantity>
<geoCoordinate id="F7AA50279D569211FB82FA28CCCC2DD2" box="[1041,1178,1429,1456]" degrees="1" direction="north" minutes="22" orientation="latitude" pageId="74" pageNumber="75" precision="15" seconds="57" value="1.3825">N 1°22’57”</geoCoordinate>
,
<geoCoordinate id="F7AA50279D569211FB35FA28CD1F2DD2" box="[1190,1353,1429,1456]" pageId="74" pageNumber="75">E 124°59’76”</geoCoordinate>
, LF
<date id="E62010209D569211FF2EFA07C9612DB1" box="[189,311,1465,1491]" pageId="74" pageNumber="75" value="2008-02-18">
<collectingDate id="F664E9C89D569211FF2EFA07C9612DB1" box="[189,311,1465,1491]" pageId="74" pageNumber="75" value="2008-02-18">18.II.2008</collectingDate>
</date>
leg.
<collectorName id="3F6B53369D569211FEFCFA07C9822DB1" box="[367,468,1465,1491]" pageId="74" pageNumber="75">A. Skale</collectorName>
(004) //
<collectorName id="3F6B53369D569211FDA0FA07CB462DB1" box="[563,784,1465,1491]" pageId="74" pageNumber="75">Coll. A. Skale Hof</collectorName>
,
<collectingCountry id="EA8976709D569211FC88FA07CBD92DB1" box="[795,911,1465,1491]" name="Germany" pageId="74" pageNumber="75">Germany</collectingCountry>
”;
<collectionCode id="F48FAE259D569211FC36FA04CBBC2DB6" box="[933,1002,1466,1492]" country="Germany" httpUri="http://grbio.org/cool/87iw-xebj" name="Sammlung des Naturkundemseum Erfurt" pageId="74" pageNumber="75">NME</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="922136E09D569211FF79FA42C90A2E74" blockId="74.[189,1398,1532,1630]" box="[234,348,1532,1558]" pageId="74" pageNumber="75">
<emphasis id="A0EAEAF29D569211FF79FA42C90A2E74" bold="true" box="[234,348,1532,1558]" pageId="74" pageNumber="75">
<typeStatus id="4D2588429D569211FF79FA42C90A2E74" box="[234,348,1532,1558]" pageId="74" pageNumber="75" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D569211FF2EF9A1CD382E3F" blockId="74.[189,1398,1532,1630]" pageId="74" pageNumber="75">
<materialsCitation id="22F63CBD9D569211FF2EF9A1CD3C2E3F" ID-GBIF-Occurrence="2898878304" collectingDate="1985-10" collectionCode="IRSN" collectorName="Leg. J. Van Stalle" country="Indonesia" elevation="1600" location="Utara - Dumoga Bone Nat. Park - Poniki Trail" pageId="74" pageNumber="75" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi" typeStatus="paratype">
<collectingCountry id="EA8976709D569211FF2EF9A1C90D2E5B" box="[189,347,1567,1593]" name="Indonesia" pageId="74" pageNumber="75">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D569211FEEDF9A1C9E62E58" box="[382,432,1567,1594]" pageId="74" pageNumber="75" type="male">1 ♂</specimenCount>
; “Coll. I.R.Sc.N.B.
<collectingRegion id="505AF8029D569211FD39F9A1CB4F2E5B" box="[682,793,1567,1593]" country="Indonesia" name="Sulawesi Utara" pageId="74" pageNumber="75">Sulawesi</collectingRegion>
:
<location id="9741603B9D569211FCBBF99ECD202E5B" LSID="urn:lsid:plazi:treatment:1A3787F69D569217FDE9FC0FCAE12EC5:9741603B9D569211FCBBF99ECD202E5B" box="[808,1398,1567,1594]" country="Indonesia" name="Utara - Dumoga Bone Nat. Park - Poniki Trail" pageId="74" pageNumber="75" stateProvince="Sulawesi">Utara – Dumoga Bone Nat. Park – Poniki Trail</location>
(
<quantity id="55669B059D569211FF56F9FDC94F2E3C" box="[197,281,1603,1630]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="74" pageNumber="75" unit="m" value="1600.0">
<elevation id="19B3D1D39D569211FF56F9FDC94F2E3C" box="[197,281,1603,1630]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="74" pageNumber="75" unit="m" value="1600.0">1600m</elevation>
</quantity>
) Station 029
<date id="E62010209D569211FE24F9FDCA582E3C" box="[439,526,1603,1630]" pageId="74" pageNumber="75" value="1985-10">
<collectingDate id="F664E9C89D569211FE24F9FDCA582E3C" box="[439,526,1603,1630]" pageId="74" pageNumber="75" value="1985-10">X.1985</collectingDate>
</date>
<collectorName id="3F6B53369D569211FD85F9FACAB52E3F" box="[534,739,1603,1630]" pageId="74" pageNumber="75">Leg. J. Van Stalle</collectorName>
// Coll. I.R.Sc.N.B. Don. O. Missa I.G.:33-444”;
<collectionCode id="F48FAE259D569211FAB0F9FDCD3C2E3F" box="[1315,1386,1603,1629]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/wzbt-fxwr" name="Institut Royal des Sciences Naturelles de Belgique" pageId="74" pageNumber="75">IRSN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D569217FF2EF93BC9FD2DDE" lastPageId="76" lastPageNumber="77" pageId="74" pageNumber="75" type="description">
<paragraph id="922136E09D569211FF2EF93BC9182EFD" blockId="74.[189,334,1669,1695]" box="[189,334,1669,1695]" pageId="74" pageNumber="75">
<heading id="C969818C9D569211FF2EF93BC9182EFD" bold="true" box="[189,334,1669,1695]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A0EAEAF29D569211FF2EF93BC9182EFD" bold="true" box="[189,334,1669,1695]" pageId="74" pageNumber="75">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D569211FF2EF90BCAE42F74" blockId="74.[189,1399,1716,1815]" pageId="74" pageNumber="75">
MEASUREMENTS. Male
<typeStatus id="4D2588429D569211FE52F90BCA7D2EAD" box="[449,555,1717,1743]" pageId="74" pageNumber="75" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D569211FC9AF90ACB202EAD" box="[777,886,1716,1743]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" pageId="74" pageNumber="75" unit="mm" value="6.35">6.35 mm</quantity>
. Head
<quantity id="55669B059D569211FC5AF90ACC652EAD" box="[969,1075,1716,1743]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="74" pageNumber="75" unit="mm" value="1.05">1.05 mm</quantity>
long, across eyes
<quantity id="55669B059D569211FA9EF90BCD212EAD" box="[1293,1399,1717,1743]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="74" pageNumber="75" unit="mm" value="1.6">1.60 mm</quantity>
wide. Pronotum
<quantity id="55669B059D569211FEEDF966C9B02E91" box="[382,486,1752,1779]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="74" pageNumber="75" unit="mm" value="1.6">1.60 mm</quantity>
long, maximum width
<quantity id="55669B059D569211FD66F966CB092E91" box="[757,863,1752,1779]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="74" pageNumber="75" unit="mm" value="2.6">2.60 mm</quantity>
. Elytral length
<quantity id="55669B059D569211FB83F967CC2D2E91" box="[1040,1147,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="74" pageNumber="75" unit="mm" value="3.7">3.70 mm</quantity>
, maximum combined width
<quantity id="55669B059D569211FE94F943C9252F75" box="[263,371,1789,1815]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="74" pageNumber="75" unit="mm" value="3.1">3.10 mm</quantity>
.
<typeStatus id="4D2588429D569211FEEDF943C9B32F75" box="[382,485,1789,1815]" pageId="74" pageNumber="75" type="paratype">Paratype</typeStatus>
is
<quantity id="55669B059D569211FD94F942CA262F75" box="[519,624,1788,1815]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="74" pageNumber="75" unit="mm" value="6.8">6.80 mm</quantity>
long.
</paragraph>
<paragraph id="922136E09D569217FF2EF881CD5A2D25" blockId="74.[189,1399,1855,2025]" lastBlockId="76.[189,1399,265,1352]" lastPageId="76" lastPageNumber="77" pageId="74" pageNumber="75">
Dorsum and venter uniformly black-brown with brown labroclypeus, antennae and legs. Head transverse, slightly convex dorsally between eyes, subopaque dorsally and ventrally, with large not prominent eyes occupying less than half side of head. Male labroclypeus (
<figureCitation id="0AA52A659D569211FCE8F838CBB72FC3" box="[891,993,1926,1953]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="74" pageNumber="75">Fig. 144</figureCitation>
) broadly V-shaped emarginate on anterior margin, with lateral margins smooth, sinuous in both dorsal and lateral views. Anterolateral angles of labroclypeus protruding anteriad, nearly right-angular. Head punctures irregularly ovoid, very deep and dense. Each puncture encircled with microscopical velvety pubescence. Intervening spaces glossy, generally much smaller than punctures. Head setae dirty yellowish, generally sparse, long and erect, with some scattered minute setae not surpassing length of their corresponding punctures. Labroclypeus laterally and frontally with long sparse setae. Male antenna 9-segmented, club 3-lamellate. Scape moderately large, bulbous dorsally in distal half, provided with few long erect setae on its posterior and distal margin. Antennomere 2 short, slightly transverse. Pronotum transverse, glossy dorsally and laterally. Anterior margin of pronotum slightly sinuous with anterolateral angles (stronger) and mesal portion (slightly) protruding anteriad. Basal margin of pronotum broadly rounded. Lateral margin of pronotum broadly rounded with maximum width posterior to midlength, crenulate all along (
<figureCitation id="0AA52A659D509217FA91FD86CD3F2A30" box="[1282,1385,568,594]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="76" pageNumber="77">Fig. 231</figureCitation>
). Long curved erect seta present between every two crenulae. Lateral margin of pronotum nearly straight in lateral view. Pronotal punctures ovoid, very deep and dense. Intervening spaces glossy and glabrous, much smaller than punctures. Setae similar to those on head, stronger curved and comparatively shorter. Lateral and basal margins, antero- and posterolateral angles partly covered with microscopical velvety pubescence. Hypomeron smooth, sparsely covered by circular punctures, flange-like, with large emargination opposite to compound eye (to receive canthus separating compound eye), provided with long setae. Antennal pocket deep. Scutellar shield pointed apically. Elytra cylindrical, maximum width in median third, subopaque, with rounded humeri and slightly elevated omoplates. Indistinct tracks of two flat longitudinal carinae on each elytron. Sutural carina not indicated. Elytral disc and lateral sides of elytra with dense and deep ovoid punctures; perimeters of many punctures completely or partly encircled with microscopical velvety pubescence (
<figureCitation id="0AA52A659D509217FCC5FC66CBEB2B90" box="[854,957,984,1010]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="76" pageNumber="77">Fig. 322</figureCitation>
). Intervening spaces glossy, generally smaller than punctures. Setae of two kinds on elytral disc. Very long, erect, in part clavate setae arranged in irregular longitudinal rows present on each elytron. Inconspicuous short setae rising from anterior margin of corresponding punctures and not surpassing their length positioned between longitudinal rows of longer setae. Male pygidium flattened dorsally, with deep ovoid punctures (
<figureCitation id="0AA52A659D509217FBFCFBCECC8E2CE8" box="[1135,1240,1136,1162]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="76" pageNumber="77">Fig. 491</figureCitation>
). Intervening spaces variably large, covered with dense microscopical pubescence. Setae of pygidium erect, long, in part clavate. Male protibia with two inconspicuous distal teeth on external margin (Fig. 395). Male lower meso- and metatibial terminal spur strongly curved. Distal margin of male metatibia with projection over basal metatarsomere. Male basal metatarsomere widened and dorso-ventrally flattened. Claws with pulvilli (in males only?). Spiculum gastrale as in
<figureCitation id="0AA52A659D509217FD6CFA93CB322D2A" box="[767,868,1325,1352]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="76" pageNumber="77">Fig. 553</figureCitation>
. Male aedeagus as in
<figureCitation id="0AA52A659D509217FBF7FA93CD5E2D25" box="[1124,1288,1325,1351]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="76" pageNumber="77">Figs 661–663</figureCitation>
.
</paragraph>
<caption id="C6E166689D579210FF2EF836CD7C2F8A" ID-DOI="http://doi.org/10.5281/zenodo.4122152" ID-Zenodo-Dep="4122152" httpUri="https://zenodo.org/record/4122152/files/figure.png" pageId="75" pageNumber="76" startId="75.[189,241,1928,1954]" targetBox="[336,1249,266,1914]" targetPageId="75">
<paragraph id="922136E09D579210FF2EF836CD7C2F8A" blockId="75.[189,1399,1928,2025]" pageId="75" pageNumber="76">
<emphasis id="A0EAEAF29D579210FF2EF836C91B2FC0" bold="true" box="[189,333,1928,1954]" pageId="75" pageNumber="76">Figs 58–61.</emphasis>
Papuan and Wallacean
<taxonomicName id="559E4D639D579210FDE5F836CBEB2FC0" authority="Macleay, 1819" authorityName="Macleay" authorityYear="1819" box="[630,957,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="A0EAEAF29D579210FDE5F836CAA92FC0" box="[630,767,1928,1954]" italics="true" pageId="75" pageNumber="76">Maechidius</emphasis>
<bibRefCitation id="F60F4B119D579210FC99F836CBEB2FC0" author="Macleay W. S." box="[778,957,1928,1954]" pageId="75" pageNumber="76" refId="ref98319" refString="Macleay W. S. 1819. Horae Entomologicae: or Essays on the Annulose Animals. Volume I, Part I. Containing General Observations on the Geography, Manners, and the Affinities of the Insects which compose the Genus Scarabaeus of Linnaeus; to which are added a few incidental Remarks on the Genera Lucanus and Hister of the same Author. With Appendix and Plates. R. and A. Taylor, London." type="book" year="1819">Macleay, 1819</bibRefCitation>
</taxonomicName>
, habitus, dorsal view.
<emphasis id="A0EAEAF29D579210FB40F836CCA62FC0" bold="true" box="[1235,1264,1928,1954]" pageId="75" pageNumber="76">58</emphasis>
.
<taxonomicName id="559E4D639D579210FA90F837CD212FC0" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[1283,1399,1928,1954]" class="Aves" family="Megapodiidae" genus="Macrocephalon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Galliformes" pageId="75" pageNumber="76" phylum="Chordata" rank="species" species="maleo" status="sp. nov.">
<emphasis id="A0EAEAF29D579210FA90F837CD212FC0" box="[1283,1399,1928,1954]" italics="true" pageId="75" pageNumber="76">M. maleo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D579210FF2EF812C94F2FA4" box="[189,281,1964,1990]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (IRSN).
<emphasis id="A0EAEAF29D579210FD8AF815CA602FA7" bold="true" box="[537,566,1963,1989]" pageId="75" pageNumber="76">59</emphasis>
.
<taxonomicName id="559E4D639D579210FDD7F812CB632FA7" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[580,821,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="miklouhomaclayi" status="sp. nov.">
<emphasis id="A0EAEAF29D579210FDD7F812CB632FA7" box="[580,821,1963,1989]" italics="true" pageId="75" pageNumber="76">M. miklouhomaclayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D579210FCA8F812CBC12FA4" box="[827,919,1964,1990]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (NHMB).
<emphasis id="A0EAEAF29D579210FB3FF815CC9F2FA7" bold="true" box="[1196,1225,1963,1989]" pageId="75" pageNumber="76">60</emphasis>
.
<taxonomicName id="559E4D639D579210FB4BF812C9072F8A" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="milneanus">
<emphasis id="A0EAEAF29D579210FB4BF812CD212FA7" box="[1240,1399,1963,1989]" italics="true" pageId="75" pageNumber="76">M. milneanus</emphasis>
<bibRefCitation id="F60F4B119D579210FF2EF870C9072F8A" author="Heller K. M." box="[189,337,1998,2024]" pageId="75" pageNumber="76" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
, lectotype, ♀ (SNSD).
<emphasis id="A0EAEAF29D579210FDF1F870CA292F8A" bold="true" box="[610,639,1998,2024]" pageId="75" pageNumber="76">61</emphasis>
.
<taxonomicName id="559E4D639D579210FD1DF871CB4F2F8A" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[654,793,1998,2024]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="merdeka" status="sp. nov.">
<emphasis id="A0EAEAF29D579210FD1DF871CB4F2F8A" box="[654,793,1998,2024]" italics="true" pageId="75" pageNumber="76">M. merdeka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D579210FCB3F871CB2B2F8B" box="[800,893,1999,2025]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (BMNH). Not to scale.
</paragraph>
</caption>
<paragraph id="922136E09D509217FF2EFACFC9E62DE9" blockId="76.[189,432,1393,1419]" box="[189,432,1393,1419]" pageId="76" pageNumber="77">
<heading id="C969818C9D509217FF2EFACFC9E62DE9" bold="true" box="[189,432,1393,1419]" fontSize="11" level="3" pageId="76" pageNumber="77" reason="3">
<emphasis id="A0EAEAF29D509217FF2EFACFC9E62DE9" bold="true" box="[189,432,1393,1419]" pageId="76" pageNumber="77">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D509217FF2EFA1CC9FD2DDE" blockId="76.[189,427,1442,1468]" box="[189,427,1442,1468]" pageId="76" pageNumber="77">Female is unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D509217FF2EFA58CD3A2E50" box="[189,1388,1510,1586]" pageId="76" pageNumber="77" type="biology_ecology">
<paragraph id="922136E09D509217FF2EFA58C9762E62" blockId="76.[189,288,1510,1536]" box="[189,288,1510,1536]" pageId="76" pageNumber="77">
<heading id="C969818C9D509217FF2EFA58C9762E62" bold="true" box="[189,288,1510,1536]" fontSize="11" level="3" pageId="76" pageNumber="77" reason="3">
<emphasis id="A0EAEAF29D509217FF2EFA58C9762E62" bold="true" box="[189,288,1510,1536]" pageId="76" pageNumber="77">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D509217FF2EF9A6CD3A2E50" blockId="76.[189,1388,1559,1586]" box="[189,1388,1559,1586]" pageId="76" pageNumber="77">
Occurs in lowland up to mid-montane rainforests. Possibly nocturnal (
<typeStatus id="4D2588429D509217FC66F9A9CC082E53" box="[1013,1118,1559,1585]" pageId="76" pageNumber="77" type="holotype">holotype</typeStatus>
was attracted to light).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D509217FF2EF9E5CAE12EC5" box="[189,695,1627,1703]" pageId="76" pageNumber="77" type="distribution">
<paragraph id="922136E09D509217FF2EF9E5C9002E17" blockId="76.[189,342,1627,1653]" box="[189,342,1627,1653]" pageId="76" pageNumber="77">
<heading id="C969818C9D509217FF2EF9E5C9002E17" bold="true" box="[189,342,1627,1653]" fontSize="11" level="3" pageId="76" pageNumber="77" reason="3">
<emphasis id="A0EAEAF29D509217FF2EF9E5C9002E17" bold="true" box="[189,342,1627,1653]" pageId="76" pageNumber="77">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D509217FF2EF933CAE12EC5" blockId="76.[189,695,1677,1703]" box="[189,695,1677,1703]" pageId="76" pageNumber="77">
Hitherto only known from
<collectingRegion id="505AF8029D509217FE6AF933CAE52EC5" box="[505,691,1677,1703]" country="Indonesia" name="Sulawesi Utara" pageId="76" pageNumber="77">North Sulawesi</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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