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<document id="74E135B0C6B0D2A823D9AFF0283233E3" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D559211FDE8FEB6CBDD2B11" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius mailu Telnov 2020, sp. nov." docType="treatment" docUuid="65367219-2F03-4D5A-B717-CA6A980839F6" docUuidSource="ZooBank" docVersion="10" lastPageNumber="75" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="74" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="923C6162CF6F3B1F3E614E8FE9D9F124" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE168A4B55F35E0840A0F729345338F2">
<mods:title id="BF3F3AAAB8605D2C96019596DBC82782">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="BB82180A0D960854CD5BA9C86F60D9AD" type="personal">
<mods:role id="406C664499CCB51DE29FC1B172B1AF18">
<mods:roleTerm id="AE981FE62BC09DF4EC98E8CFAE0DC4D8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D5498F0E3E644009DF046687152B8B98">Telnov, Dmitry</mods:namePart>
</mods:name>
<mods:typeOfResource id="7B61EFF29C0D984AF4D366A078DC0866">text</mods:typeOfResource>
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<mods:titleInfo id="DC136FAB30DEDFF11AB6EC6597A5A5B8">
<mods:title id="5E7901D69C0177B5EAC632AF14B085A6">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="F952A4F01282F24E77583E56DC40F966">
<mods:date id="F53E247214AB81509D853C782E5F0CCF">2020</mods:date>
<mods:detail id="AC28A298812AC3604F63DFE51933DF84" type="pubDate">
<mods:number id="9DC5FEEFAAFDE17C1E68BC804595CEEE">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="6EB391B546B8F4D6C03D78CB7180BD75" type="volume">
<mods:number id="108E3D6A9F8067A126D82D597461BC65">721</mods:number>
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<mods:start id="712ED6BCBFBC1F83CC3F82CDF1F49B22">1</mods:start>
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<mods:classification id="EBA122AEFA7A9EE2C1F13187F9F5C58E">journal article</mods:classification>
<mods:identifier id="57B3BFD779F2E5A34ECB0422118913C6" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="81C92E56C15737DB1A7FB326827AD692" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="B6AFF745C91BDDB85426342CA7E680BD" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
<mods:identifier id="7E291F9934A9C9104D8567A88D868B8F" type="Zenodo-Dep">4122118</mods:identifier>
<mods:identifier id="FADE65295AE832327A5C0FC7A9ECB317" type="ZooBank">89E62EF8-2E45-4C59-94B7-6A5603E8939B</mods:identifier>
</mods:mods>
<treatment id="1A3787F69D559211FDE8FEB6CBDD2B11" ID-DOI="http://doi.org/10.5281/zenodo.4344287" ID-GBIF-Taxon="168648165" ID-Zenodo-Dep="4344287" LSID="urn:lsid:zoobank.org:act:65367219-2F03-4D5A-B717-CA6A980839F6" httpUri="http://treatment.plazi.org/id/1A3787F69D559211FDE8FEB6CBDD2B11" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
<subSubSection id="DA84656B9D559212FDE8FEB6CBEF2941" box="[635,953,264,291]" pageId="73" pageNumber="74" type="nomenclature">
<paragraph id="922136E09D559212FDE8FEB6CBEF2941" blockId="73.[373,1214,264,363]" box="[635,953,264,291]" pageId="73" pageNumber="74">
<heading id="C969818C9D559212FDE8FEB6CBEF2941" box="[635,953,264,291]" centered="true" fontSize="11" level="2" pageId="73" pageNumber="74" reason="2">
<taxonomicName id="559E4D639D559212FDE8FEB6CB002941" ID-CoL="7X3X4" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[635,854,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="mailu" status="sp. nov.">
<emphasis id="A0EAEAF29D559212FDE8FEB6CB002941" bold="true" box="[635,854,264,291]" italics="true" pageId="73" pageNumber="74">Maechidius mailu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D559212FCCDFEB7CBEF2941" box="[862,953,265,291]" pageId="73" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D559212FEE6FE92CC662909" pageId="73" pageNumber="74" type="description">
<paragraph id="922136E09D559212FEE6FE92CCE82925" blockId="73.[373,1214,264,363]" box="[373,1214,300,327]" pageId="73" pageNumber="74">
<uri id="E60F3AE29D559212FEE6FE92CCE82925" box="[373,1214,300,327]" pageId="73" pageNumber="74">
urn:lsid:zoobank.org:act:
<uuid id="E6380C359D559212FD01FE92CCE82925" box="[658,1214,300,327]" pageId="73" pageNumber="74">65367219-2F03-4D5A-B717-CA6A980839F6</uuid>
</uri>
</paragraph>
<paragraph id="922136E09D559212FD90FEEECC662909" blockId="73.[373,1214,264,363]" box="[515,1072,336,363]" pageId="73" pageNumber="74">
<figureCitation id="0AA52A659D559212FD90FEEECA0B2909" box="[515,605,336,363]" captionStart="Figs 54–57" captionStartId="69.[189,241,1925,1951]" captionTargetBox="[359,1240,274,1913]" captionTargetId="figure-59@69.[346,1240,265,1925]" captionTargetPageId="69" captionText="Figs 54–57. Papuan and Wallacean Maechidius Macleay, 1819, habitus, dorsal view. 54. M. lobaticeps Frey, 1969, ♂ (DTC). 55. M. longipes sp. nov., holotype, ♂ (BMNH). 56. M. mailu sp. nov., holotype, ♂ (NHMB). 57. M. luniceps Fairmaire, 1883, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122150" httpUri="https://zenodo.org/record/4122150/files/figure.png" pageId="73" pageNumber="74">Figs 56</figureCitation>
,
<figureCitation id="0AA52A659D559212FDFAFEEFCAC12909" box="[617,663,337,363]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="73" pageNumber="74">141</figureCitation>
,
<figureCitation id="0AA52A659D559212FD37FEEFCA852909" box="[676,723,337,363]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="73" pageNumber="74">230</figureCitation>
,
<figureCitation id="0AA52A659D559212FD4CFEEFCB582909" box="[735,782,337,363]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="73" pageNumber="74">271</figureCitation>
,
<figureCitation id="0AA52A659D559212FC8AFEEFCB1E2909" box="[793,840,337,363]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="73" pageNumber="74">321</figureCitation>
, 394, 490, 658–660
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D559212FF2EFE2CCD39299D" pageId="73" pageNumber="74" type="diagnosis">
<paragraph id="922136E09D559212FF2EFE2CC99F29CE" blockId="73.[189,457,402,428]" box="[189,457,402,428]" pageId="73" pageNumber="74">
<heading id="C969818C9D559212FF2EFE2CC99F29CE" bold="true" box="[189,457,402,428]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="A0EAEAF29D559212FF2EFE2CC99F29CE" bold="true" box="[189,457,402,428]" pageId="73" pageNumber="74">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D559212FF2EFE7FCD39299D" blockId="73.[189,1398,449,512]" pageId="73" pageNumber="74">
<taxonomicName id="559E4D639D559212FF2EFE7FC9C229B9" authority="Telnov, 2020" authorityName="Telnov" authorityYear="2020" box="[189,404,449,475]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="mailu" status="sp. nov.">
<emphasis id="A0EAEAF29D559212FF2EFE7FC9C229B9" box="[189,404,449,475]" italics="true" pageId="73" pageNumber="74">Maechidius mailu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BBD957899D559212FE33FE7CC9A929BE" box="[416,511,450,476]" pageId="73" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
is peculiar among all congeners primarily due to the shape of the male aedeagus, the almost straight lateral margins of the pronotum and the partly wrinkled elytral sculpture.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D559212FF2EFD99C9692AF6" pageId="73" pageNumber="74" type="etymology">
<paragraph id="922136E09D559212FF2EFD99C9122A23" blockId="73.[189,324,551,577]" box="[189,324,551,577]" pageId="73" pageNumber="74">
<heading id="C969818C9D559212FF2EFD99C9122A23" bold="true" box="[189,324,551,577]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="A0EAEAF29D559212FF2EFD99C9122A23" bold="true" box="[189,324,551,577]" pageId="73" pageNumber="74">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D559212FF2EFDE8C9692AF6" blockId="73.[189,1399,598,660]" pageId="73" pageNumber="74">
Named after the Mailu, the local tribe and language of Amazon Bay,
<collectingCountry id="EA8976709D559212FB85FDE9CD532A12" box="[1046,1285,598,625]" name="Papua New Guinea" pageId="73" pageNumber="74">Papua New Guinea</collectingCountry>
. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D559212FF2EFD02CC162BB4" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="922136E09D559212FF2EFD02C93A2AB4" blockId="73.[189,364,700,726]" box="[189,364,700,726]" pageId="73" pageNumber="74">
<heading id="C969818C9D559212FF2EFD02C93A2AB4" bold="true" box="[189,364,700,726]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="A0EAEAF29D559212FF2EFD02C93A2AB4" bold="true" box="[189,364,700,726]" pageId="73" pageNumber="74">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="22F63CBD9D559212FF79FD55C90A2B13" ID-GBIF-Occurrence="2898878351" collectingDate="1962-11-26" collectionCode="NHMB" collectorName="W. W. Brandt" country="Papua New Guinea" determinerName="G. Frey" elevation="1036" location="NEW GUINEA" pageId="73" pageNumber="74" specimenCount="1" typeStatus="holotype">
<paragraph id="922136E09D559212FF79FD55C90A2B67" blockId="73.[189,1399,747,881]" box="[234,348,747,773]" pageId="73" pageNumber="74">
<emphasis id="A0EAEAF29D559212FF79FD55C90A2B67" bold="true" box="[234,348,747,773]" pageId="73" pageNumber="74">
<typeStatus id="4D2588429D559212FF79FD55C90A2B67" box="[234,348,747,773]" pageId="73" pageNumber="74" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D559212FF2EFCB1C90A2B13" blockId="73.[189,1399,747,881]" pageId="73" pageNumber="74">
<collectingCountry id="EA8976709D559212FF2EFCB1C98E2B4B" box="[189,472,783,809]" name="Papua New Guinea" pageId="73" pageNumber="74">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D559212FE62FCB1CA5F2B4B" box="[497,521,783,809]" pageId="73" pageNumber="74" type="male">♂</specimenCount>
; “
<location id="9741603B9D559212FDB3FCB1CA8A2B4B" LSID="urn:lsid:plazi:treatment:1A3787F69D559211FDE8FEB6CBDD2B11:9741603B9D559212FDB3FCB1CA8A2B4B" box="[544,732,783,809]" country="Papua New Guinea" name="NEW GUINEA" pageId="73" pageNumber="74">NEW GUINEA</location>
<location id="9741603B9D559212FD72FCB1CBE02B4B" LSID="urn:lsid:plazi:treatment:1A3787F69D559211FDE8FEB6CBDD2B11:9741603B9D559212FD72FCB1CBE02B4B" box="[737,950,783,809]" country="Papua New Guinea" name="Amazon Bay area" pageId="73" pageNumber="74">Amazon Bay area</location>
, Komania.
<quantity id="55669B059D559212FBA0FCB1CCC72B4B" box="[1075,1169,783,809]" metricMagnitude="3" metricUnit="m" metricValue="1.03632" pageId="73" pageNumber="74" unit="ft" value="3400.0">
<elevation id="19B3D1D39D559212FBA0FCB1CCC72B4B" box="[1075,1169,783,809]" metricMagnitude="3" metricUnit="m" metricValue="1.03632" pageId="73" pageNumber="74" unit="ft" value="3400.0">3400 ft.</elevation>
</quantity>
, 11.11.–
<date id="E62010209D559212FB7CFCB1CD212B4B" box="[1263,1399,783,809]" pageId="73" pageNumber="74" value="1962-11-26">
<collectingDate id="F664E9C89D559212FB7CFCB1CD212B4B" box="[1263,1399,783,809]" pageId="73" pageNumber="74" value="1962-11-26">26.11.1962</collectingDate>
</date>
<collectorName id="3F6B53369D559212FF2EFC8DC90F2B2E" box="[189,345,818,845]" pageId="73" pageNumber="74">W. W. Brandt</collectorName>
[p]//
<typeStatus id="4D2588429D559212FE0AFC8DCA7D2B2F" box="[409,555,819,845]" pageId="73" pageNumber="74" type="paratype">PARATYPE</typeStatus>
[p, red label]//
<taxonomicName id="559E4D639D559212FD4EFC8CCC662B2E" authorityName="Frey" authorityYear="1969" box="[733,1072,818,844]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="lineatopunctatus">
<emphasis id="A0EAEAF29D559212FD4EFC8CCC662B2E" box="[733,1072,818,844]" italics="true" pageId="73" pageNumber="74">Maechidius lineatopunctatus</emphasis>
</taxonomicName>
n sp [h] det.
<determinerName id="67FDDA249D559212FB2EFC8CCD472B2E" box="[1213,1297,818,844]" pageId="73" pageNumber="74">G.Frey</determinerName>
,1967/68 [p]”;
<collectionCode id="F48FAE259D559212FF6FFCE9C90E2B13" box="[252,344,855,881]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="73" pageNumber="74">NHMB</collectionCode>
.
</paragraph>
</materialsCitation>
<paragraph id="922136E09D559212FF79FC26C9312BD0" blockId="73.[189,1088,920,982]" box="[234,359,920,946]" pageId="73" pageNumber="74">
<emphasis id="A0EAEAF29D559212FF79FC26C9312BD0" bold="true" box="[234,359,920,946]" pageId="73" pageNumber="74">
<typeStatus id="4D2588429D559212FF79FC26C9312BD0" box="[234,359,920,946]" pageId="73" pageNumber="74" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D559212FF2EFC02CC162BB4" blockId="73.[189,1088,920,982]" box="[189,1088,956,982]" pageId="73" pageNumber="74">
<materialsCitation id="22F63CBD9D559212FF2EFC02CC6A2BB4" ID-GBIF-Occurrence="2898878442" box="[189,1084,956,982]" collectingDate="1964-10-15" collectingDateMax="1964-10-24" collectingDateMin="1964-10-15" collectionCode="NHMB" collectorName="M. E. Bacchus." country="Papua New Guinea" county="Mts. Budemu" determinerName="G. Frey" elevation="1219" location="PAPUA NEW GUINEA" municipality="Finisterre Mts." pageId="73" pageNumber="74" specimenCount="3" typeStatus="paratype">
<collectingCountry id="EA8976709D559212FF2EFC02C98F2BB4" box="[189,473,956,982]" name="Papua New Guinea" pageId="73" pageNumber="74">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D559212FE61FC02CAD02BB4" box="[498,646,956,982]" pageId="73" pageNumber="74" type="generic">3 specimens</specimenCount>
; same labels as for holotype:
<collectionCode id="F48FAE259D559212FC73FC02CC6A2BB4" box="[992,1084,956,982]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="73" pageNumber="74">NHMB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D559212FF2EFC40CC312CED" pageId="73" pageNumber="74" type="discussion">
<paragraph id="922136E09D559212FF2EFC40C9792C7A" blockId="73.[189,303,1022,1048]" box="[189,303,1022,1048]" pageId="73" pageNumber="74">
<heading id="C969818C9D559212FF2EFC40C9792C7A" bold="true" box="[189,303,1022,1048]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="A0EAEAF29D559212FF2EFC40C9792C7A" bold="true" box="[189,303,1022,1048]" pageId="73" pageNumber="74">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D559212FF2EFB93CC312CED" blockId="73.[189,1398,1069,1167]" pageId="73" pageNumber="74">
According to the original description of
<taxonomicName id="559E4D639D559212FD01FB90CC1D2C2A" authority="(Frey 1969: 502)" baseAuthorityName="Frey" baseAuthorityPageNumber="502" baseAuthorityYear="1969" box="[658,1099,1069,1096]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="lineatopunctatus">
<emphasis id="A0EAEAF29D559212FD01FB90CB2A2C25" box="[658,892,1069,1095]" italics="true" pageId="73" pageNumber="74">M. lineatopunctatus</emphasis>
(
<bibRefCitation id="F60F4B119D559212FC19FB93CC142C2A" author="Frey G." box="[906,1090,1069,1096]" pageId="73" pageNumber="74" pagination="494 - 509" refId="ref98001" refString="Frey G. 1969. Revision der Maechidius - Arten Neu-Guineas (Col., Melolonth.). Entomologische Arbeiten aus dem Museum G. Frey 20: 494 - 509." type="journal article" year="1969">Frey 1969: 502</bibRefCitation>
)
</taxonomicName>
, there are
<specimenCount id="8498FD699D559212FB52FB93CD482C25" box="[1217,1310,1069,1095]" pageId="73" pageNumber="74" type="male">17 male</specimenCount>
/female
<typeStatus id="4D2588429D559212FF2EFBEFC97B2C09" box="[189,301,1105,1131]" pageId="73" pageNumber="74" type="paratype">paratypes</typeStatus>
at NHMB of which I had the opportunity to study one single male and three additional specimens that were allocated by the NHMB staff (C. Germann, personal communication).
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D559211FF2EFB08C9F22AF2" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="description">
<paragraph id="922136E09D559212FF2EFB08C9182CB2" blockId="73.[189,334,1206,1232]" box="[189,334,1206,1232]" pageId="73" pageNumber="74">
<heading id="C969818C9D559212FF2EFB08C9182CB2" bold="true" box="[189,334,1206,1232]" fontSize="11" level="3" pageId="73" pageNumber="74" reason="3">
<emphasis id="A0EAEAF29D559212FF2EFB08C9182CB2" bold="true" box="[189,334,1206,1232]" pageId="73" pageNumber="74">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D559212FF2EFB58C9212D25" blockId="73.[189,1399,1253,1351]" pageId="73" pageNumber="74">
MEASUREMENTS. Male
<typeStatus id="4D2588429D559212FE52FB5BCA7D2C9D" box="[449,555,1253,1279]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
, total body length
<quantity id="55669B059D559212FC9AFB58CB202D62" box="[777,886,1254,1280]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="73" pageNumber="74" unit="mm" value="8.2">8.20 mm</quantity>
. Head
<quantity id="55669B059D559212FC5AFB5BCC652D62" box="[969,1075,1253,1280]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="73" pageNumber="74" unit="mm" value="1.5">1.50 mm</quantity>
long, across eyes 2.00 mm wide. Pronotum
<quantity id="55669B059D559212FEEDFAB7C9B02D46" box="[382,486,1289,1316]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="73" pageNumber="74" unit="mm" value="1.7">1.70 mm</quantity>
long, maximum width
<quantity id="55669B059D559212FD66FAB7CB092D46" box="[757,863,1289,1316]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="73" pageNumber="74" unit="mm" value="2.9">2.90 mm</quantity>
. Elytral length 5.00 mm, maximum combined width 4.00 mm.
</paragraph>
<paragraph id="922136E09D559211FF2EFAD1CAE12A7D" blockId="73.[189,1399,1391,2024]" lastBlockId="74.[189,1399,265,543]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
Dorsum and venter uniformly brown, appendages and labroclypeus castaneous. Head flattened dorsally on frons, subopaque dorsally and ventrally. Male labroclypeus (
<figureCitation id="0AA52A659D559212FC72FA2DCC1A2DCF" box="[993,1100,1427,1453]" captionStart="Figs 133–144" captionStartId="140.[189,241,1822,1848]" captionTargetBox="[346,1223,273,1784]" captionTargetId="figure-120@140.[345,1234,265,1799]" captionTargetPageId="140" captionText="Figs 133–144. Papuan and Wallacean Maechidius Macleay, 1819, head, dorsal view. 133. M. lapsus sp. nov., holotype, ♂ (BMNH). 134. Same, paratype, ♀ (BMNH). 135. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 136. M. legalovi sp. nov., holotype, ♂ (BMNH). 137. Same, paratype, ♀ (BMNH). 138. M. longipes sp. nov., holotype, ♂ (BMNH). 139. M. lobaticeps Frey, 1969, ♂ (BMNH). 140. Same, ♀. 141. M. mailu sp. nov., holotype, ♂ (NHMB). 142. M. luniceps Fairmaire, 1883, ♂ (BMNH). 143. Same, ♀ (BMNH). 144. M. maleo sp. nov., holotype, ♂ (IRSN). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122179" httpUri="https://zenodo.org/record/4122179/files/figure.png" pageId="73" pageNumber="74">Fig. 141</figureCitation>
) broadly and shallowly emarginate on anterior margin, its lateral margins sinuous in both dorsal and lateral views. Anterolateral angles strongly obtuse, rounded, not protruding. Upper- and underside of labroclypeus with sparse inconspicuous setae along anterior and lateral margins. Compound eye moderately large. Canthus very broadly rounded in dorsal view. Head punctures circular, moderately deep and dense. Intervening spaces glossy to microreticulate, variably large. Head setae minute, appressed, rise from anterior margin of each puncture, not surpassing midlength of corresponding punctures. Antenna 9-segmented, club 3-lamellate. Pronotum strongly transverse, glossy dorsally and laterally.Anterior margin of pronotum slightly sinuous with anterolateral angles slightly protruding anteriad. Basal margin of pronotum slightly sinuous. Lateral margin of pronotum in dorsal view nearly straight, very slightly emarginate in basal half, delicately crenulate all along (
<figureCitation id="0AA52A659D559212FE34F946CA592F70" box="[423,527,1784,1810]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="73" pageNumber="74">Fig. 230</figureCitation>
). Minute erect seta present between every two crenulae. Lateral margin of pronotum slightly sinuous in lateral view. Hypomeron separated from prosternum by moderately high nearly straight carina which is shortly flange-like protruding in posterior half, with long setae on its anterolateral margin opposite to compound eye (
<figureCitation id="0AA52A659D559212FD68F8DDCB342F1F" box="[763,866,1891,1917]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="73" pageNumber="74">Fig. 271</figureCitation>
). Antennal pocket moderately deep. Pronotal punctures circular to ovoid, dense and moderately deep, in part annular. Intervening spaces glossy or delicately microreticulate, variably large, in part wrinkled. Setae as those on head, very inconspicuous. Scutellar shield triangular, pointed apically. Elytron subopaque, without tracks of longitudinal carinae. Punctures of elytral disc ovoid, deep, dense. Intervening spaces in part wrinkled in part glossy, variably large. Elytral setae inconspicuous, appressed; each seta rises from anterior margin of each puncture, not surpassing length of corresponding puncture (
<figureCitation id="0AA52A659D569211FC8FFEEFCBD12909" box="[796,903,337,363]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="74" pageNumber="75">Fig. 321</figureCitation>
). Male pygidium flattened dorsally, with large and shallow irregularly hexagonal punctures (
<figureCitation id="0AA52A659D569211FC8BFECBCB2929ED" box="[792,895,373,399]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="74" pageNumber="75">Fig. 490</figureCitation>
). Intervening spaces microreticulate, much smaller than punctures. Setae of pygidium very short in anterior, slightly longer in apical half. Male protibia with two inconspicuous distal teeth on external margin (Fig. 394). Male protibial terminal spur straight, pointed. Male lower mesotibial terminal spur slightly curved. Tarsal claws with pulvilli (in male only?). Aedeagus as in
<figureCitation id="0AA52A659D569211FD9DFDBACAE52A7D" box="[526,691,516,543]" captionStart="Figs 652–666" captionStartId="175.[189,241,1892,1918]" captionTargetBox="[436,1156,267,1863]" captionTargetId="figure-70@175.[426,1161,265,1868]" captionTargetPageId="175" captionText="Figs 652–666. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 652– 654. M. longipes sp. nov., holotype, ♂ (BMNH). 655–657. M. luniceps Fairmaire, 1883, ♂ (BMNH). 658–660. M. mailu sp. nov., holotype, ♂ (NHMB). 661–663. M. maleo sp. nov., holotype, ♂ (IRSN). 664–666. M. merdeka sp. nov., holotype, ♂ (BMNH). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122244" httpUri="https://zenodo.org/record/4122244/files/figure.png" pageId="74" pageNumber="75">Figs 658–660</figureCitation>
.
</paragraph>
<paragraph id="922136E09D569211FF2EFDF8C9E62A02" blockId="74.[189,432,582,608]" box="[189,432,582,608]" pageId="74" pageNumber="75">
<heading id="C969818C9D569211FF2EFDF8C9E62A02" bold="true" box="[189,432,582,608]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A0EAEAF29D569211FF2EFDF8C9E62A02" bold="true" box="[189,432,582,608]" pageId="74" pageNumber="75">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D569211FF2EFDC8C9F22AF2" blockId="74.[189,420,630,656]" box="[189,420,630,656]" pageId="74" pageNumber="75">Female not studied.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D569211FF2EFD06C96F2B63" box="[189,313,696,769]" pageId="74" pageNumber="75" type="biology_ecology">
<paragraph id="922136E09D569211FF2EFD06C9762AB0" blockId="74.[189,288,696,722]" box="[189,288,696,722]" pageId="74" pageNumber="75">
<heading id="C969818C9D569211FF2EFD06C9762AB0" bold="true" box="[189,288,696,722]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A0EAEAF29D569211FF2EFD06C9762AB0" bold="true" box="[189,288,696,722]" pageId="74" pageNumber="75">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D569211FF2EFD59C96F2B63" blockId="74.[189,313,743,769]" box="[189,313,743,769]" pageId="74" pageNumber="75">Unknown.</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D569211FF2EFC97CBDD2B11" box="[189,907,809,883]" pageId="74" pageNumber="75" type="distribution">
<paragraph id="922136E09D569211FF2EFC97C9002B21" blockId="74.[189,342,809,835]" box="[189,342,809,835]" pageId="74" pageNumber="75">
<heading id="C969818C9D569211FF2EFC97C9002B21" bold="true" box="[189,342,809,835]" fontSize="11" level="3" pageId="74" pageNumber="75" reason="3">
<emphasis id="A0EAEAF29D569211FF2EFC97C9002B21" bold="true" box="[189,342,809,835]" pageId="74" pageNumber="75">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D569211FF2EFCE7CBDD2B11" blockId="74.[189,907,857,883]" box="[189,907,857,883]" pageId="74" pageNumber="75">
Hitherto only known from Papuan Peninsula of New
<collectingCountry id="EA8976709D569211FCBCFCE7CBDE2B11" box="[815,904,857,883]" name="Guinea-Bissau" pageId="74" pageNumber="75">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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