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<document id="0EC758A58D0FE792109B44368F8CCCC0" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D449203FDD5FEB6C9C72F8A" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius papuanus Moser 1926" docType="treatment" docVersion="10" lastPageNumber="89" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="89" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2C9D19B2CD604F73306D71DB48557475">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
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<mods:namePart id="E8FDE3F6079054D761CA57F6A933A6F9">Telnov, Dmitry</mods:namePart>
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<mods:title id="51AB29F5AAADD0A060D83B94A133DC67">European Journal of Taxonomy</mods:title>
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<mods:date id="EE08B5CB776FF6EA261C861A01A68EC2">2020</mods:date>
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<mods:number id="FE77A0D561BF5B2B69B5B01932FE6B5A">2020-10-19</mods:number>
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<mods:number id="BD0BE98F6C64EB1B3F47DBA078E89729">721</mods:number>
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<treatment id="1A3787F69D449203FDD5FEB6C9C72F8A" ID-DOI="http://doi.org/10.5281/zenodo.4344275" ID-GBIF-Taxon="168648152" ID-Zenodo-Dep="4344275" LSID="urn:lsid:plazi:treatment:1A3787F69D449203FDD5FEB6C9C72F8A" httpUri="http://treatment.plazi.org/id/1A3787F69D449203FDD5FEB6C9C72F8A" lastPageNumber="89" pageId="88" pageNumber="89">
<subSubSection id="DA84656B9D449203FDD5FEB6CBBB2941" box="[582,1005,264,291]" pageId="88" pageNumber="89" type="nomenclature">
<paragraph id="922136E09D449203FDD5FEB6CBBB2941" blockId="88.[545,1043,264,329]" box="[582,1005,264,291]" pageId="88" pageNumber="89">
<heading id="C969818C9D449203FDD5FEB6CBBB2941" box="[582,1005,264,291]" centered="true" fontSize="11" level="2" pageId="88" pageNumber="89" reason="2">
<taxonomicName id="559E4D639D449203FDD5FEB6CBBB2941" ID-CoL="3XFKK" authority="Moser, 1926" authorityName="Moser" authorityYear="1926" box="[582,1005,264,291]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="A0EAEAF29D449203FDD5FEB6CB052941" bold="true" box="[582,851,264,291]" italics="true" pageId="88" pageNumber="89">Maechidius papuanus</emphasis>
<bibRefCitation id="F60F4B119D449203FCC9FEB7CBBB2941" author="Moser J." box="[858,1005,265,291]" pageId="88" pageNumber="89" pagination="196 - 203" refId="ref98419" refString="Moser J. 1926. Neue Melolonthiden und Cetoniden aus dem Suden von Hollandisch Neu-Guinea. Nova Guinea: a Journal of Botany, Zoology, Anthropology, Ethnography, Geology and Palaeontology of the Papuan Region. Zoology 15 (2): 196 - 203." type="journal article" year="1926">Moser, 1926</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D449203FDB2FE90CC45292B" box="[545,1043,302,329]" pageId="88" pageNumber="89" type="description">
<paragraph id="922136E09D449203FDB2FE90CC45292B" blockId="88.[545,1043,264,329]" box="[545,1043,302,329]" pageId="88" pageNumber="89">
<figureCitation id="0AA52A659D449203FDB2FE90CA2D292A" box="[545,635,302,328]" captionStart="Figs 62–65" captionStartId="81.[189,241,1926,1952]" captionTargetBox="[260,1331,260,1918]" captionTargetId="figure-59@81.[254,1332,254,1925]" captionTargetPageId="81" captionText="Figs 62–65. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 62. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 63. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 64. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 65. M. nepenthephilus sp. nov., holotype, ♂ (NME). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122154" httpUri="https://zenodo.org/record/4122154/files/figure.png" pageId="88" pageNumber="89">Figs 62</figureCitation>
,
<figureCitation id="0AA52A659D449203FD14FE90CAE0292B" box="[647,694,302,329]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="88" pageNumber="89">156</figureCitation>
,
<figureCitation id="0AA52A659D449203FD52FE91CAA6292B" box="[705,752,303,329]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="88" pageNumber="89">240</figureCitation>
,
<figureCitation id="0AA52A659D449203FD6FFE91CB7D292B" box="[764,811,303,329]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="88" pageNumber="89">273</figureCitation>
,
<figureCitation id="0AA52A659D449203FCA4FE91CB30292B" box="[823,870,303,329]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="88" pageNumber="89">331</figureCitation>
, 404, 501–502
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D449203FF2EFECCC94C2A4C" pageId="88" pageNumber="89" type="materials_examined">
<paragraph id="922136E09D449203FF2EFECCC93A29EE" blockId="88.[189,364,370,396]" box="[189,364,370,396]" pageId="88" pageNumber="89">
<heading id="C969818C9D449203FF2EFECCC93A29EE" bold="true" box="[189,364,370,396]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A0EAEAF29D449203FF2EFECCC93A29EE" bold="true" box="[189,364,370,396]" pageId="88" pageNumber="89">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D449203FF79FE1DCA3A29DF" blockId="88.[189,1399,418,558]" box="[234,620,418,445]" pageId="88" pageNumber="89">
<emphasis id="A0EAEAF29D449203FF79FE1DC93029DF" bold="true" box="[234,358,419,445]" pageId="88" pageNumber="89">
<typeStatus id="4D2588429D449203FF79FE1DC93029DF" box="[234,358,419,445]" pageId="88" pageNumber="89" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D449203FF2EFE77C94C2A4C" blockId="88.[189,1399,418,558]" pageId="88" pageNumber="89">
<materialsCitation id="22F63CBD9D449203FF2EFE77C94C2A4C" ID-GBIF-Occurrence="2898878303" collectionCode="ZMHB" country="Indonesia" location="INDONESIA" pageId="88" pageNumber="89" specimenCode="MFNB 2019" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<collectingCountry id="EA8976709D449203FF2EFE77C90D2981" box="[189,347,457,483]" name="Indonesia" pageId="88" pageNumber="89">INDONESIA</collectingCountry>
•
<specimenCount id="8498FD699D449203FEFCFE77C9D02981" box="[367,390,457,483]" pageId="88" pageNumber="89" type="female">♀</specimenCount>
; “Nieuw.Guinea-Expeditie 1903 [p] Etna-Bai [h] [black frame] //
<taxonomicName id="559E4D639D449203FBE5FE76C8AF2A6A" authority="Mos." authorityName="Moser" authorityYear="1926" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="A0EAEAF29D449203FBE5FE76CD202980" box="[1142,1398,456,482]" italics="true" pageId="88" pageNumber="89">Maechidius papuanus</emphasis>
Mos.
</taxonomicName>
<typeStatus id="4D2588429D449203FF6DFE50C9132A6A" box="[254,325,494,520]" pageId="88" pageNumber="89">Typus</typeStatus>
[h] //
<typeStatus id="4D2588429D449203FE16FE50CA5F2A6A" box="[389,521,494,520]" pageId="88" pageNumber="89" type="syntype">SYNTYPE</typeStatus>
<taxonomicName id="559E4D639D449203FD9CFE50CBFE2A6A" authority="Moser, 1926" authorityName="Moser" authorityYear="1926" box="[527,936,494,520]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="A0EAEAF29D449203FD9CFE50CB462A6A" box="[527,784,494,520]" italics="true" pageId="88" pageNumber="89">Maechidius papuanus</emphasis>
<bibRefCitation id="F60F4B119D449203FC85FE50CBFE2A6A" author="Moser J." box="[790,936,494,520]" pageId="88" pageNumber="89" pagination="196 - 203" refId="ref98419" refString="Moser J. 1926. Neue Melolonthiden und Cetoniden aus dem Suden von Hollandisch Neu-Guinea. Nova Guinea: a Journal of Botany, Zoology, Anthropology, Ethnography, Geology and Palaeontology of the Papuan Region. Zoology 15 (2): 196 - 203." type="journal article" year="1926">Moser, 1926</bibRefCitation>
</taxonomicName>
labelled by
<specimenCode id="C2389E9B9D449203FBA7FE50CC9E2A6A" box="[1076,1224,494,520]" collectionCode="MFNB" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="88" pageNumber="89">MFNB 2019</specimenCode>
[p, red label]”;
<collectionCode id="F48FAE259D449203FF2EFDAAC9402A4C" box="[189,278,532,558]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="88" pageNumber="89">ZMHB</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D449203FF2EFDE9C9002B70" pageId="88" pageNumber="89" type="discussion">
<paragraph id="922136E09D449203FF2EFDE9C9792A13" blockId="88.[189,303,599,625]" box="[189,303,599,625]" pageId="88" pageNumber="89">
<heading id="C969818C9D449203FF2EFDE9C9792A13" bold="true" box="[189,303,599,625]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A0EAEAF29D449203FF2EFDE9C9792A13" bold="true" box="[189,303,599,625]" pageId="88" pageNumber="89">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D449203FF2EFD37C9002B70" blockId="88.[189,1399,648,786]" pageId="88" pageNumber="89">
<bibRefCitation id="F60F4B119D449203FF2EFD37C9F62AC0" author="Moser J." box="[189,416,648,675]" pageId="88" pageNumber="89" pagination="196 - 203" refId="ref98419" refString="Moser J. 1926. Neue Melolonthiden und Cetoniden aus dem Suden von Hollandisch Neu-Guinea. Nova Guinea: a Journal of Botany, Zoology, Anthropology, Ethnography, Geology and Palaeontology of the Papuan Region. Zoology 15 (2): 196 - 203." type="journal article" year="1926">Moser (1926: 200)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D449203FD47FD37CB272AC0" authorityName="Moser" authorityYear="1926" box="[724,881,649,674]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="A0EAEAF29D449203FD47FD37CB272AC0" box="[724,881,649,674]" italics="true" pageId="88" pageNumber="89">M. papuanus</emphasis>
</taxonomicName>
on an unstated number of specimens. The
<typeStatus id="4D2588429D449203FF2EFD13C97C2AA5" box="[189,298,685,711]" pageId="88" pageNumber="89" type="lectotype">lectotype</typeStatus>
is therefore designated for the single
<typeStatus id="4D2588429D449203FD60FD10CB192AAA" box="[755,847,686,712]" pageId="88" pageNumber="89" type="syntype">syntype</typeStatus>
in ZMHB in order to enhance the stability of nomenclature and fix the specimen I based my redescription on, in case other
<typeStatus id="4D2588429D449203FBC8FD6DCC952A8F" box="[1115,1219,723,749]" pageId="88" pageNumber="89" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D449203FF2EFC82C9C72F8A" pageId="88" pageNumber="89" type="description">
<paragraph id="922136E09D449203FF2EFC82C9182B34" blockId="88.[189,334,828,854]" box="[189,334,828,854]" pageId="88" pageNumber="89">
<heading id="C969818C9D449203FF2EFC82C9182B34" bold="true" box="[189,334,828,854]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A0EAEAF29D449203FF2EFC82C9182B34" bold="true" box="[189,334,828,854]" pageId="88" pageNumber="89">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D449203FF2EFCD0CB112BEA" blockId="88.[189,839,877,904]" box="[189,839,877,904]" pageId="88" pageNumber="89">
MEASUREMENTS. Total body length
<quantity id="55669B059D449203FDC2FCD3CAEF2BEA" box="[593,697,877,904]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="88" pageNumber="89" unit="mm" value="7.9">7.90 mm</quantity>
(
<typeStatus id="4D2588429D449203FD5BFCD3CB6D2BEA" box="[712,827,877,904]" pageId="88" pageNumber="89" type="lectotype">lectotype</typeStatus>
).
</paragraph>
<paragraph id="922136E09D449203FF2EFC0FCA342F16" blockId="88.[189,1399,945,1908]" pageId="88" pageNumber="89">
Body uniformly brown, labroclypeus and legs slightly paler. Head slightly wider than long, opaque dorsally, frons flattened. Female labroclypeus (
<figureCitation id="0AA52A659D449203FD4FFC68CB692B93" box="[732,831,982,1009]" captionStart="Figs 145–156" captionStartId="141.[189,241,1787,1813]" captionTargetBox="[328,1254,265,1740]" captionTargetId="figure-124@141.[328,1260,265,1749]" captionTargetPageId="141" captionText="Figs 145–156. Papuan Maechidius Macleay, 1819, head, dorsal view. 145. M. merdeka sp. nov., holotype, ♂ (BMNH). 146. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 147. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 148. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 149. Same, ♂ (SNSD). 150. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 151. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 152. M. nepenthephilus sp. nov., holotype, ♂ (NME). 153. Same, paratype, ♀ (DTC). 154. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 155. Same, paratype, ♀ (BMNH). 156. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122181" httpUri="https://zenodo.org/record/4122181/files/figure.png" pageId="88" pageNumber="89">Fig. 156</figureCitation>
) spatulate, truncate on anterior margin, its lateral margins straight in dorsal, slightly sinuous in lateral view. Anterolateral angles of labroclypeus obtuse, on same axis as its anterior margin (not protruding). Head punctures irregularly circular, deep and dense, intervening spaces opaque, microreticulate. Head setae not present due to condition of specimen. Pronotum transverse, opaque dorsally. Anterior margin of pronotum sinuous with anterolateral angles (stronger) and median part (slightly) protruding anteriad. Basal margin of pronotum very broadly rounded. Lateral margin of pronotum in dorsal view very broadly rounded, delicately crenulate all along (
<figureCitation id="0AA52A659D449203FF55FB62C97B2C95" box="[198,301,1244,1271]" captionStart="Figs 226–241" captionStartId="147.[189,241,1681,1707]" captionTargetBox="[391,1192,268,1647]" captionTargetId="figure-170@147.[391,1196,265,1656]" captionTargetPageId="147" captionText="Figs 226–241. Papuan and Wallacean Maechidius Macleay, 1819, pronotum, dorsal view. 226. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 227. M. lobaticeps Frey, 1969, ♂ (DTC). 228. M. longipes sp. nov., holotype, ♂ (BMNH). 229. M. luniceps Fairmaire, 1883, ♂ (BMNH). 230. M. mailu sp. nov., holotype, ♂ (NHMB). 231. M. maleo sp. nov., holotype, ♂ (IRSN). 232. M. merdeka sp. nov., holotype, ♂ (BMNH). 233. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 234. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 235. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 236. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 237. M. nepenthephilus sp. nov., holotype, ♂ (NME). 238. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 239. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 240. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 241. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122193" httpUri="https://zenodo.org/record/4122193/files/figure.png" pageId="88" pageNumber="89">Fig. 240</figureCitation>
). Moderately long suberect thickened curved seta present between every two crenulae. Lateral margin of pronotum sinuous in lateral view. Hypomeron separated from prosternum by low carina, slightly arched, with long setae opposite to compound eye (
<figureCitation id="0AA52A659D449203FC0CFA99CC5C2D20" box="[927,1034,1319,1346]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="88" pageNumber="89">Fig. 273</figureCitation>
). Antennal pocket moderately deep. Pronotal punctures deep and dense, smaller and more regular circular in median third, larger and ovoid in lateral thirds. Intervening spaces generally smaller than punctures, microreticulate. Pronotal setae inconspicuous, suberect, curved; each seta rises from anterior margin of corresponding puncture, generally surpassing its length. On posterolateral and basal margins with a row of longer thicker setae. Lateral and basal margins, antero- and posterolateral angles partly covered with dense microscopical velvety pubescence. Scutellar shield narrowly rounded apically. Elytron cylindrical, opaque, with vague track of one longitudinal carina. Elytral punctures linear (elongate and narrow), incision-like, deep (
<figureCitation id="0AA52A659D449203FF55F9EDC9672E0C" box="[198,305,1619,1646]" captionStart="Figs 317–332" captionStartId="154.[189,241,1681,1707]" captionTargetBox="[401,1187,265,1655]" captionTargetId="figure-172@154.[401,1187,265,1669]" captionTargetPageId="154" captionText="Figs 317–332. Papuan and Wallacean Maechidius Macleay, 1819, elytron, dorsal view. 317. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 318. M. lobaticeps Frey, 1969, ♂ (DTC). 319. M. longipes sp. nov., holotype, ♂ (BMNH). 320. M. luniceps Fairmaire, 1883, ♂ (BMNH). 321. M. mailu sp. nov., holotype, ♂ (NHMB). 322. M. maleo sp. nov., holotype, ♂ (IRSN). 323. M. merdeka sp. nov., holotype, ♂ (BMNH). 324. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 325. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 326. M. muticusArrow, 1941, syntype, ♂ (BMNH). 327. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 328. M. nepenthephilus sp. nov., holotype, ♂ (NME). 329. M. opatroides Arrow, 1941 comb. rest., lectotype, ♀ (BMNH). 330. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 331. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 332. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122210" httpUri="https://zenodo.org/record/4122210/files/figure.png" pageId="88" pageNumber="89">Fig. 331</figureCitation>
). An appressed inconspicuous seta rises from anterior margin of each puncture, generally surpassing length of corresponding incision. Intervening spaces larger than punctures, microreticulate. Female pygidium flattened dorsally, with strong glabrous median hump (
<figureCitation id="0AA52A659D449203FBB6F920CC992EDB" box="[1061,1231,1694,1721]" captionStart="Figs487–503" captionStartId="165.[189,241,1681,1707]" captionTargetBox="[422,1160,265,1655]" captionTargetId="figure-171@165.[421,1167,265,1658]" captionTargetPageId="165" captionText="Figs487–503.Papuan andWallaceanMaechidius Macleay, 1819,pygidium, dorsalview.487.M.lobaticeps Frey, 1969, ♂ (DTC). 488. M. longipes sp. nov., holotype, ♂ (BMNH). 489. M. luniceps Fairmaire, 1883, ♂ (BMNH). 490. M. mailu sp. nov., holotype, ♂ (NHMB). 491. M. maleo sp. nov., holotype, ♂ (IRSN). 492. M. merdeka sp. nov., holotype, ♂ (BMNH). 493. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 494. M. milneanus Heller, 1914, lectotype, ♀ (SNSD). 495. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 496. M. nanus Arrow, 1941, syntype, ♂ (BMNH). 497. M. nepenthephilus sp. nov., holotype, ♂ (NME). 498. M. opatroidesArrow, 1941 comb. rest., lectotype, ♀ (BMNH). 499. M. owenstanleyi sp. nov., holotype, ♂ (BMNH). 500. Same, paratype, ♀ (BMNH). 501. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 502. Same, different aspect. 503. M. parallelicollis Moser, 1920, lectotype, ♂ (ZMHB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122226" httpUri="https://zenodo.org/record/4122226/files/figure.png" pageId="88" pageNumber="89">Figs 501–502</figureCitation>
). Area around median hump with large irregularly circular annular punctures, intervening spaces smaller than those. Setae of pygidium inconspicuous, short, suberect. Female protibia with two conspicuous distal teeth on external margin: predistal one callus-like, very inconspicuous, distal one somewhat curved inwards, obtuse (Fig. 404). Female protibial terminal spur conspicuous, very short and thick, blunt. Female metatibia widened on distal margin.
</paragraph>
<paragraph id="922136E09D449203FF2EF823C9E62FD5" blockId="88.[189,432,1949,1975]" box="[189,432,1949,1975]" pageId="88" pageNumber="89">
<heading id="C969818C9D449203FF2EF823C9E62FD5" bold="true" box="[189,432,1949,1975]" fontSize="11" level="3" pageId="88" pageNumber="89" reason="3">
<emphasis id="A0EAEAF29D449203FF2EF823C9E62FD5" bold="true" box="[189,432,1949,1975]" pageId="88" pageNumber="89">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D449203FF2EF870C9C72F8A" blockId="88.[189,401,1998,2024]" box="[189,401,1998,2024]" pageId="88" pageNumber="89">Male is unknown.</paragraph>
</subSubSection>
</treatment>
</document>
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