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<document id="2384A4EE8E585A5FB9E840C71F685869" ID-CLB-Dataset="9693" ID-DOI="10.5852/ejt.2020.721.1127" ID-GBIF-Dataset="0cd61b23-1089-4c1b-bfc0-487887abad84" ID-Zenodo-Dep="4122118" ID-ZooBank="89E62EF8-2E45-4C59-94B7-6A5603E8939B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1603474447689" checkinUser="valdenar" docAuthor="Telnov, Dmitry" docDate="2020" docId="1A3787F69D2A926CFDD5FC60C9002D68" docLanguage="en" docName="ejt-721_telnov.pdf" docOrigin="European Journal of Taxonomy 721" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maechidius humeralis Heller 1914" docType="treatment" docVersion="10" lastPageNumber="56" masterDocId="E60EFF8E9D1C925BFF93FFBEC8562862" masterDocTitle="A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line" masterLastPageNumber="210" masterPageNumber="1" pageNumber="55" updateTime="1698846101371" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="8B626A72393C7299287083D2F13B1BA1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9784DFFB8F7AF8779CEF04E7E8B05FAB">
<mods:title id="DE2D1AA74B63A862E15A2BE270594ED0">A revision of the Maechidiini Burmeister, 1855 (Coleoptera: Scarabaeidae: Melolonthinae) from the Indo-Australian transition zone, and the first record of the tribe west of Wallace’s Line</mods:title>
</mods:titleInfo>
<mods:name id="56106131572996F48220EB18C22548AA" type="personal">
<mods:role id="F1C803BCEA9A91D0B3629BB6242D00D5">
<mods:roleTerm id="F143931B2DAEDB7CDD2A235A64160F81">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BDF48D9E8AB85D06E60197084F81A9D5">Telnov, Dmitry</mods:namePart>
</mods:name>
<mods:typeOfResource id="35DF95AE9C673092169431E87D4B4457">text</mods:typeOfResource>
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<mods:titleInfo id="E995FAEF4C8FCBF1C4375270C38D0D35">
<mods:title id="3F4FC3A68095ACF041F94290F6543200">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="EC00424169D94E353BAD603071399164">
<mods:date id="4C8B148BCF25BAF1DC5097C039C79092">2020</mods:date>
<mods:detail id="CF55035A25E6D4BFE3BC43FD9CF63750" type="pubDate">
<mods:number id="091531E14E60E7618543AEAD9F438AF1">2020-10-19</mods:number>
</mods:detail>
<mods:detail id="3DCAF9D5398496E5E9AB292918705DEB" type="volume">
<mods:number id="531B60E9F7A63D9D1476AF6855C66C68">721</mods:number>
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<mods:classification id="CC1BB480D053CABA9D34ED56390B0A64">journal article</mods:classification>
<mods:identifier id="C2D4357A577AD5B7D82CF33B2FA3D70A" type="CLB-Dataset">9693</mods:identifier>
<mods:identifier id="907619D19470090F9BD10F53F38EF9C7" type="DOI">10.5852/ejt.2020.721.1127</mods:identifier>
<mods:identifier id="6BDDF10D47F3DEA7692BD2397C57F238" type="GBIF-Dataset">0cd61b23-1089-4c1b-bfc0-487887abad84</mods:identifier>
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</mods:mods>
<treatment id="1A3787F69D2A926CFDD5FC60C9002D68" ID-DOI="http://doi.org/10.5281/zenodo.4344271" ID-GBIF-Taxon="168648137" ID-Zenodo-Dep="4344271" LSID="urn:lsid:plazi:treatment:1A3787F69D2A926CFDD5FC60C9002D68" httpUri="http://treatment.plazi.org/id/1A3787F69D2A926CFDD5FC60C9002D68" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
<subSubSection id="DA84656B9D2A926DFDD5FC60CBB82B9A" box="[582,1006,990,1017]" pageId="54" pageNumber="55" type="nomenclature">
<paragraph id="922136E09D2A926DFDD5FC60CBB82B9A" blockId="54.[427,1160,990,1051]" box="[582,1006,990,1017]" pageId="54" pageNumber="55">
<heading id="C969818C9D2A926DFDD5FC60CBB82B9A" box="[582,1006,990,1017]" centered="true" fontSize="11" level="2" pageId="54" pageNumber="55" reason="2">
<taxonomicName id="559E4D639D2A926DFDD5FC60CBB82B9A" ID-CoL="3XFJK" authority="Heller, 1914" authorityName="Heller" authorityYear="1914" box="[582,1006,990,1017]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="A0EAEAF29D2A926DFDD5FC60CB032B9B" bold="true" box="[582,853,990,1017]" italics="true" pageId="54" pageNumber="55">Maechidius humeralis</emphasis>
<bibRefCitation id="F60F4B119D2A926DFCCFFC60CBB82B9A" author="Heller K. M." box="[860,1006,990,1016]" pageId="54" pageNumber="55" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller, 1914</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2A926DFE38FBBECCDE2C79" box="[427,1160,1024,1051]" pageId="54" pageNumber="55" type="description">
<paragraph id="922136E09D2A926DFE38FBBECCDE2C79" blockId="54.[427,1160,990,1051]" box="[427,1160,1024,1051]" pageId="54" pageNumber="55">
<figureCitation id="0AA52A659D2A926DFE38FBBECA532C79" box="[427,517,1024,1051]" captionStart="Figs 42–45" captionStartId="52.[189,241,1892,1918]" captionTargetBox="[334,1252,265,1842]" captionTargetId="figure-69@52.[334,1252,265,1861]" captionTargetPageId="52" captionText="Figs 42–45. Papuan Maechidius Macleay, 1819, habitus, dorsal view. 42. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 43. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BPBM). 44. M. hirtipesArrow, 1941, lectotype, ♂ (BMNH). 45. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122144" httpUri="https://zenodo.org/record/4122144/files/figure.png" pageId="54" pageNumber="55">Figs 45</figureCitation>
,
<figureCitation id="0AA52A659D2A926DFD82FBBECA2C2C78" box="[529,634,1024,1050]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="54" pageNumber="55">127–128</figureCitation>
,
<figureCitation id="0AA52A659D2A926DFD14FBBECAE02C78" box="[647,694,1024,1050]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="54" pageNumber="55">218</figureCitation>
,
<figureCitation id="0AA52A659D2A926DFD52FBBECAA62C78" box="[705,752,1024,1050]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="54" pageNumber="55">269</figureCitation>
,
<figureCitation id="0AA52A659D2A926DFD6FFBBECB7D2C78" box="[764,811,1024,1050]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="54" pageNumber="55">308</figureCitation>
, 382, 475–476, 546, 623–625
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2A926DFF2EFBFFCBA72DDB" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="922136E09D2A926DFF2EFBFFC93A2C39" blockId="54.[189,364,1089,1115]" box="[189,364,1089,1115]" pageId="54" pageNumber="55">
<heading id="C969818C9D2A926DFF2EFBFFC93A2C39" bold="true" box="[189,364,1089,1115]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="A0EAEAF29D2A926DFF2EFBFFC93A2C39" bold="true" box="[189,364,1089,1115]" pageId="54" pageNumber="55">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2A926DFF79FBD1CA3A2CEB" blockId="54.[189,1399,1134,1263]" box="[234,620,1134,1161]" pageId="54" pageNumber="55">
<emphasis id="A0EAEAF29D2A926DFF79FBD1C9302CEB" bold="true" box="[234,358,1135,1161]" pageId="54" pageNumber="55">
<typeStatus id="4D2588429D2A926DFF79FBD1C9302CEB" box="[234,358,1135,1161]" pageId="54" pageNumber="55" type="lectotype">Lectotype</typeStatus>
</emphasis>
[herewith designated]
</paragraph>
<paragraph id="922136E09D2A926DFF2EFB2FC9E12C8D" blockId="54.[189,1399,1134,1263]" pageId="54" pageNumber="55">
<materialsCitation id="22F63CBD9D2A926DFF2EFB2FC9E12C8D" ID-GBIF-Occurrence="2898878350" collectionCode="SNSD" collectorName="Gebirge Dr. Schlaginhaufen" country="Papua New Guinea" location="PAPUA NEW GUINEA" pageId="54" pageNumber="55" specimenCount="2" typeStatus="typus">
<collectingCountry id="EA8976709D2A926DFF2EFB2FC9882CC9" box="[189,478,1169,1195]" name="Papua New Guinea" pageId="54" pageNumber="55">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D2A926DFE68FB2FCA452CC9" box="[507,531,1169,1195]" pageId="54" pageNumber="55" type="male">♂</specimenCount>
; “Kais.Wilhelmsland Toricelli [sic
<specimenCount id="8498FD699D2A926DFC2BFB2ECBEA2CC9" box="[952,956,1168,1195]" pageId="54" pageNumber="55" type="male">♂</specimenCount>
]
<collectorName id="3F6B53369D2A926DFC5DFB2FCD472CC9" box="[974,1297,1169,1195]" pageId="54" pageNumber="55">Gebirge Dr.Schlaginhaufen</collectorName>
[p, blue label]// 1910 [p] 1 [h] [blue label]//
<typeStatus id="4D2588429D2A926DFDC7FB0DCAF42CAF" box="[596,674,1203,1229]" pageId="54" pageNumber="55">Typus</typeStatus>
! [p]
<taxonomicName id="559E4D639D2A926DFD42FB0DCB102CAF" authorityName="Heller" authorityYear="1914" box="[721,838,1203,1229]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="A0EAEAF29D2A926DFD42FB0DCB102CAF" box="[721,838,1203,1229]" italics="true" pageId="54" pageNumber="55">humeralis</emphasis>
</taxonomicName>
H. [h, red label]// Staatl. Museum für Tierkunde. Dresden [p]”;
<collectionCode id="F48FAE259D2A926DFEF6FB6BC9E22C8D" box="[357,436,1237,1263]" collectionName="SNSD" pageId="54" pageNumber="55">SNSD</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="922136E09D2A926DFF79FAA8C9F42D52" blockId="54.[189,1399,1302,1465]" box="[234,418,1302,1328]" pageId="54" pageNumber="55">
<emphasis id="A0EAEAF29D2A926DFF79FAA8C9F42D52" bold="true" box="[234,418,1302,1328]" pageId="54" pageNumber="55">
<typeStatus id="4D2588429D2A926DFF79FAA8C9F42D52" box="[234,418,1302,1328]" pageId="54" pageNumber="55" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="922136E09D2A926DFF2EFA86CBA72DDB" blockId="54.[189,1399,1302,1465]" pageId="54" pageNumber="55">
<materialsCitation id="22F63CBD9D2A926DFF2EFA86CC862D16" ID-GBIF-Occurrence="2898878590" collectionCode="SNSD" collectorName="Gebirge Dr. Schlaginhaufen" country="Papua New Guinea" location="Kais. Wilhelmsland Toricelli" pageId="54" pageNumber="55" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
<collectingCountry id="EA8976709D2A926DFF2EFA86C98F2D30" box="[189,473,1336,1362]" name="Papua New Guinea" pageId="54" pageNumber="55">PAPUA NEW GUINEA</collectingCountry>
•
<specimenCount id="8498FD699D2A926DFE60FA86CA762D30" box="[499,544,1336,1362]" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
; “
<location id="9741603B9D2A926DFDAAFA86CBD12D30" LSID="urn:lsid:plazi:treatment:1A3787F69D2A926CFDD5FC60C9002D68:9741603B9D2A926DFDAAFA86CBD12D30" box="[569,903,1336,1362]" country="Papua New Guinea" name="Kais. Wilhelmsland Toricelli" pageId="54" pageNumber="55">Kais.Wilhelmsland Toricelli</location>
[sic
<specimenCount id="8498FD699D2A926DFC53FA89CB922D30" box="[960,964,1335,1362]" pageId="54" pageNumber="55" type="male">♂</specimenCount>
]
<collectorName id="3F6B53369D2A926DFC47FA86CD432D30" box="[980,1301,1336,1362]" pageId="54" pageNumber="55">Gebirge Dr.Schlaginhaufen</collectorName>
[p, blue label]// 1910 [p] 1 [h] [blue label]//
<location id="9741603B9D2A926DFDF8FAE4CC6D2D16" LSID="urn:lsid:plazi:treatment:1A3787F69D2A926CFDD5FC60C9002D68:9741603B9D2A926DFDF8FAE4CC6D2D16" box="[619,1083,1370,1397]" country="Papua New Guinea" name="Staatl. Museum fur Tierkunde. Dresden" pageId="54" pageNumber="55">Staatl. Museum für Tierkunde.Dresden</location>
[p]”;
<collectionCode id="F48FAE259D2A926DFB16FAE4CC862D16" box="[1157,1232,1370,1396]" collectionName="SNSD" pageId="54" pageNumber="55">SNSD</collectionCode>
</materialsCitation>
•
<materialsCitation id="22F63CBD9D2A926DFB7EFAE4CBBB2DDB" ID-GBIF-Occurrence="2898878399" collectionCode="SNSD" collectorName="Kais. Wilhelmsland Paup Dr. Schlaginhaufen" country="Papua New Guinea" location="Staatl. Museum fur Tierkunde. Dresden" pageId="54" pageNumber="55" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="typus">
<specimenCount id="8498FD699D2A926DFB7EFAE4CD4A2D17" box="[1261,1308,1370,1397]" pageId="54" pageNumber="55" type="female">1 ♀</specimenCount>
; “
<collectorName id="3F6B53369D2A926DFAA4FAE4CAD42DF4" pageId="54" pageNumber="55">Kais. Wilhelmsland Paup Dr.Schlaginhaufen</collectorName>
[p, blue label]// 1910 [p] 1 [h] [blue label] //
<typeStatus id="4D2588429D2A926DFB11FAC2CC862DF4" box="[1154,1232,1404,1430]" pageId="54" pageNumber="55">Typus</typeStatus>
<specimenCount id="8498FD699D2A926DFB43FAC2CC822DF4" box="[1232,1236,1404,1430]" pageId="54" pageNumber="55" type="male">♂</specimenCount>
[p]
<taxonomicName id="559E4D639D2A926DFA92FAC2CD202DF4" authorityName="Heller" authorityYear="1914" box="[1281,1398,1404,1430]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="A0EAEAF29D2A926DFA92FAC2CD202DF4" box="[1281,1398,1404,1430]" italics="true" pageId="54" pageNumber="55">humeralis</emphasis>
</taxonomicName>
H. [h, red label]//
<location id="9741603B9D2A926DFE1CFA20CB0E2DDA" LSID="urn:lsid:plazi:treatment:1A3787F69D2A926CFDD5FC60C9002D68:9741603B9D2A926DFE1CFA20CB0E2DDA" box="[399,856,1438,1465]" country="Papua New Guinea" name="Staatl. Museum fur Tierkunde. Dresden" pageId="54" pageNumber="55">Staatl. Museum für Tierkunde.Dresden</location>
[p]”;
<collectionCode id="F48FAE259D2A926DFC0DFA21CBBB2DDB" box="[926,1005,1439,1465]" collectionName="SNSD" pageId="54" pageNumber="55">SNSD</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2A926DFF2EFA61CD4C2ED2" pageId="54" pageNumber="55" type="discussion">
<paragraph id="922136E09D2A926DFF2EFA61C9792D9B" blockId="54.[189,303,1503,1529]" box="[189,303,1503,1529]" pageId="54" pageNumber="55">
<heading id="C969818C9D2A926DFF2EFA61C9792D9B" bold="true" box="[189,303,1503,1529]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="A0EAEAF29D2A926DFF2EFA61C9792D9B" bold="true" box="[189,303,1503,1529]" pageId="54" pageNumber="55">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2A926DFF2EF9B3CD4C2ED2" blockId="54.[189,1399,1549,1712]" pageId="54" pageNumber="55">
<bibRefCitation id="F60F4B119D2A926DFF2EF9B3C9F62E4A" author="Heller K. M." box="[189,416,1549,1576]" pageId="54" pageNumber="55" pagination="615 - 666" refId="ref98213" refString="Heller K. M. 1914. Coleoptera. Nova Guinea. Resultats de l'Expedition scientifique neerlandaise a la Nouvelle-Guinee en 1907 et 1909 sous les auspices du Dr. H. A. Lorentz. Zoologie 9: 615 - 666." type="journal article" year="1914">Heller (1914: 628)</bibRefCitation>
based his description of
<taxonomicName id="559E4D639D2A926DFD49F9B3CB2C2E45" authorityName="Heller" authorityYear="1914" box="[730,890,1549,1575]" class="Insecta" family="Scarabaeidae" genus="Maechidius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="humeralis">
<emphasis id="A0EAEAF29D2A926DFD49F9B3CB2C2E45" box="[730,890,1549,1575]" italics="true" pageId="54" pageNumber="55">M. humeralis</emphasis>
</taxonomicName>
on a number of specimens, although not explicitly stated, and two different localities are given. The
<typeStatus id="4D2588429D2A926DFC04F991CC552E2B" box="[919,1027,1583,1609]" pageId="54" pageNumber="55" type="lectotype">lectotype</typeStatus>
designated here is one of two
<typeStatus id="4D2588429D2A926DFF2EF9ECC9722E0E" box="[189,292,1618,1644]" pageId="54" pageNumber="55" type="syntype">syntypes</typeStatus>
with Heller’s original “typus” handwriting on a red label and is the single male
<typeStatus id="4D2588429D2A926DFA87F9ECCD252E0E" box="[1300,1395,1618,1644]" pageId="54" pageNumber="55" type="syntype">syntype</typeStatus>
.
<typeStatus id="4D2588429D2A926DFF2EF9CAC9642EEC" box="[189,306,1652,1678]" pageId="54" pageNumber="55" type="lectotype">Lectotype</typeStatus>
designation is made in order to enhance the stability of nomenclature and fix the only male specimen I based my redescription on, in case other
<typeStatus id="4D2588429D2A926DFCF7F928CB9D2ED2" box="[868,971,1686,1712]" pageId="54" pageNumber="55" type="syntype">syntypes</typeStatus>
are discovered in the future.
</paragraph>
</subSubSection>
<subSubSection id="DA84656B9D2A926CFF2EF968C9002D68" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55" type="description">
<paragraph id="922136E09D2A926DFF2EF968C9182E92" blockId="54.[189,334,1750,1776]" box="[189,334,1750,1776]" pageId="54" pageNumber="55">
<heading id="C969818C9D2A926DFF2EF968C9182E92" bold="true" box="[189,334,1750,1776]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="A0EAEAF29D2A926DFF2EF968C9182E92" bold="true" box="[189,334,1750,1776]" pageId="54" pageNumber="55">Description</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2A926DFF2EF8BACC962F7C" blockId="54.[189,1216,1796,1823]" box="[189,1216,1796,1823]" pageId="54" pageNumber="55">
MEASUREMENTS. Total body length 6.80 (smallest
<typeStatus id="4D2588429D2A926DFD6FF8BACBCA2F7D" box="[764,924,1796,1823]" pageId="54" pageNumber="55" type="paralectotype">paralectotype</typeStatus>
) to 8.10 (
<typeStatus id="4D2588429D2A926DFB9FF8BACC2D2F7D" box="[1036,1147,1796,1823]" pageId="54" pageNumber="55" type="lectotype">lectotype</typeStatus>
) mm.
</paragraph>
<paragraph id="922136E09D2A926CFF2EF8FBCA482C15" blockId="54.[189,1399,1860,2025]" lastBlockId="55.[189,1399,265,1143]" lastPageId="55" lastPageNumber="56" pageId="54" pageNumber="55">
Dorsum and venter uniformly black-brown except for brown appendages and labroclypeus. Head flattened dorsally, glossy dorsally and ventrally. Male labroclypeus (
<figureCitation id="0AA52A659D2A926DFC74F8D8CC1A2FE3" box="[999,1100,1894,1921]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="54" pageNumber="55">Fig. 127</figureCitation>
) broadly and moderately deeply emarginate anteriorly, its lateral margins strongly sinuous in both dorsal and lateral views. Anterolateral angles rounded, moderately strongly protruding, raised up at angle of nearly 80-90° to frons in lateral view. Anterior and lateral margins of labroclypeus smooth. Female labroclypeus as in
<figureCitation id="0AA52A659D2B926CFF2EFEB7C9762941" box="[189,288,265,291]" captionStart="Figs 121–132" captionStartId="139.[189,241,1716,1742]" captionTargetBox="[357,1230,276,1689]" captionTargetId="figure-149@139.[357,1230,265,1697]" captionTargetPageId="139" captionText="Figs 121–132. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, head, dorsal (121, 123–124, 126–132) and frontal (122, 125) views. 121. M. hamatus sp. nov., holotype, ♂ (IECA). 122. M. helleri (Frey, 1969) comb. nov., holotype, ♂ (photograph courtesy BPBM) (BPBM). 123. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 124. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 125. Same, ♂, holotype of M. clypeatus (Frey, 1969) comb. nov. (BMNH). 126. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 127. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 128. Same, paralectotype, ♀ (SNSD). 129. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 130. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 131. M. konjo sp. nov., holotype, ♂ (NMPC). 132. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122177" httpUri="https://zenodo.org/record/4122177/files/figure.png" pageId="55" pageNumber="56">Fig. 128</figureCitation>
. Underside of labroclypeus with sparse moderately long setae along anterior and lateral margins. Canthus broadly rounded in dorsal view. Punctures of frons irregularly shaped, annular, large and shallow. Intervening spaces subopaque, variably large. Setae inconspicuous, suberect, sparse, generally hardly surpassing length of corresponding punctures. Antenna 9-segmented, club 3-lamellate. Pronotum transverse, flattened dorsally, glossy dorsally and laterally. Anterior margin of pronotum slightly sinuous with protruding anterolateral angles, basal margin slightly sinuous. Lateral margin of pronotum sinuous in dorsal and lateral view, delicately crenulate all along (
<figureCitation id="0AA52A659D2B926CFCE3FE60CB8D299A" box="[880,987,478,504]" captionStart="Figs 208–225" captionStartId="146.[189,241,1644,1670]" captionTargetBox="[414,1187,265,1610]" captionTargetId="figure-199@146.[414,1187,265,1618]" captionTargetPageId="146" captionText="Figs 208–225. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pronotum, dorsal view. 208. M.caperatus sp. nov., holotype, ♂ (NME). 209. M.ciliatus sp. nov., holotype, ♂ (IECA). 210. M.dani sp. nov., holotype, ♂ (IECA). 211. M. deltouri sp. nov., holotype, ♂ (BMNH). 212. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 213. M. esau Heller, 1914 (lectotype, ♂ of M. setosus Moser, 1920) (ZMHB). 214. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 215. M. hamatus sp. nov., holotype, ♂ (IECA). 216. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 217. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 218. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 219. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 220. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 221. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 222. M. konjo sp. nov., holotype, ♂ (NMPC). 223. M. lapsus sp. nov., holotype, ♂ (BMNH). 224. M. legalovi sp. nov., holotype, ♂ (BMNH). 225. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122191" httpUri="https://zenodo.org/record/4122191/files/figure.png" pageId="55" pageNumber="56">Fig. 218</figureCitation>
). Long erect seta present between every two crenulae. Hypomeron separated from prosternum by moderately high emarginate carina, with moderately long setae on its anterolateral margin (
<figureCitation id="0AA52A659D2B926CFC96FD9BCB3F2A22" box="[773,873,549,576]" captionStart="Figs 266–277" captionStartId="150.[189,241,1753,1779]" captionTargetBox="[366,1222,265,1687]" captionTargetId="figure-144@150.[366,1222,265,1703]" captionTargetPageId="150" captionText="Figs 266–277. Papuan and Wallacean Maechidius Macleay, 1819, pronotal hypomeron, ventrolateral view. 266. M. aenescens Heller, 1910, lectotype, ♀ (SNSD). 267. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 268. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 269. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 270. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). 271. M. mailu sp. nov., holotype, ♂ (NHMB). 272. M. lobaticeps Frey, 1969, ♂ (BMNH). 273. M. papuanus Moser, 1926, lectotype, ♀ (ZMHB). 274. M. paupianus Heller, 1910, lectotype, ♀ (SNSD). 275. M. pauxillus Heller, 1910 comb. rest., lectotype, ♂ (SNSD). 276. M. peregrinus Lansberge, 1886, ♂ (DTC). 277. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122200" httpUri="https://zenodo.org/record/4122200/files/figure.png" pageId="55" pageNumber="56">Fig. 269</figureCitation>
) opposite to compound eye. Antennal pocket moderately deep. Punctures of pronotal disc large, variably shaped (generally ovoid), double annular, moderately deep; lateral and latero-basal punctures irregularly ovoid, oblong, larger than median ones. Intervening spaces generally smaller than punctures, glossy. Setation slightly longer than on head, appressed to suberect; seta rises from anterior margin of each puncture, not or hardly surpassing its length. Scutellar shield pointed apically. Elytron subopaque, with very vague tracks of two longitudinal carinae. Sutural carinae not present. Punctures of elytral disc sinuous, long and narrow, incisionshaped, dense (
<figureCitation id="0AA52A659D2B926CFEE6FCA3C9882B55" box="[373,478,797,823]" captionStart="Figs 302–316" captionStartId="153.[189,241,1679,1705]" captionTargetBox="[393,1194,265,1649]" captionTargetId="figure-159@153.[393,1194,265,1658]" captionTargetPageId="153" captionText="Figs 302–316. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, elytron, dorsal view. 302. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 303. M. esau Heller, 1914, lectotype, ♂ (SNSD). 304. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 305. M. hamatus sp. nov., holotype, ♂ (IECA). 306. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 307. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 308. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 309. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 310. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 311. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 312. Same, enlarged. 313. M. konjo sp. nov., holotype, ♂ (NMPC). 314. M. lapsus sp. nov., holotype, ♂ (BMNH). 315. M. legalovi sp. nov., holotype, ♂ (BMNH). 316. M. leucopsar sp. nov., holotype, ♂ (IECA). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122208" httpUri="https://zenodo.org/record/4122208/files/figure.png" pageId="55" pageNumber="56">Fig. 308</figureCitation>
). Intervening spaces wrinkled, opaque while densely transverse microstriate, generally larger than incision-shaped punctures. Intervening spaces delicately microreticulate. Setae inconspicuous, suberect, not surpassing length of punctures. Pygidium in both sexes (
<figureCitation id="0AA52A659D2B926CFB5BFCDACD392B1D" box="[1224,1391,868,895]" captionStart="Figs 469–486" captionStartId="164.[189,241,1681,1707]" captionTargetBox="[450,1137,265,1633]" captionTargetId="figure-184@164.[450,1138,265,1647]" captionTargetPageId="164" captionText="Figs 469–486. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, pygidium, dorsal view. 469. M. dendrolagus sp. nov., holotype, ♂ (NHMB). 470. M. esau Heller, 1914, ♂ (BMNH). 471. M. fraterculus Moser, 1920, lectotype, ♀ (ZMHB). 472. M. hamatus sp. nov., holotype, ♂ (IECA). 473. M. heterosquamosus Heller, 1910 comb. rest., lectotype, ♀ (SNSD). 474. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 475. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 476. Same, paralectotype, ♀ (SNSD). 477. M. interruptocarinulatus Heller, 1914, holotype, ♀ (SNSD). 478. M. jobiensis Moser, 1920, lectotype, ♂ (ZMHB). 479. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 480. M. konjo sp. nov., holotype, ♂ (NMPC). 481. M. lapsus sp. nov., holotype, ♂ (BMNH). 482. Same, paratype, ♀ (BMNH). 483. M. legalovi sp. nov., holotype, ♂ (BMNH). 484. Same, paratype, ♀ (BMNH). 485. M. leucopsar sp. nov., holotype, ♂ (IECA). 486. M. lineatopunctatus Frey, 1969, holotype, ♂ (NHMB). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122224" httpUri="https://zenodo.org/record/4122224/files/figure.png" pageId="55" pageNumber="56">Figs 475–476</figureCitation>
) flattened dorsally, with very large and shallow irregularly shaped annular punctures. Intervening spaces microreticulate, distinctly smaller than punctures. Setae of pygidium moderately long, suberect. Protibia widened distally, dorsally with delicate longitudinal carina, in both sexes with three external teeth: acute largest distal, less acute predistal and strongly obtuse small median (Fig. 382). Male and female protibial terminal spur thin and long. Male metatibial terminal spurs long, nearly equally long, female lower metatibia spur distinctly longer than upper. Tarsal claws with pulvilli. Spiculum gastrale as in
<figureCitation id="0AA52A659D2B926CFA9EFB87CD252C36" box="[1293,1395,1081,1108]" captionStart="Figs 545–569" captionStartId="169.[189,241,1541,1567]" captionTargetBox="[308,1266,258,1476]" captionTargetId="figure-249@169.[299,1280,255,1481]" captionTargetPageId="169" captionText="Figs 545–569. Papuan, Wallacean and Oriental Maechidius Macleay, 1819, spiculum gastrale. 545. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 546. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 547. M. yamdena sp. nov., paratype, ♂ (BMNH). 548. M. kazantsevi sp. nov., holotype, ♂ tBMNH). 549. M. legalovi sp. nov., holotype, ♂ (BMNH). 550. M. leucopsar sp. nov., holotype, ♂ (IECA). 551. M. longipes sp. nov., holotype, ♂ (BMNH). 552. M. luniceps Fairmaire, 1883, ♂ (BMNH). 553. M. maleo sp. nov., holotype, ♂ (IRSN). 554. M. miklouhomaclayi sp. nov., holotype, ♂ (NHMB). 555. M. muticus Arrow, 1941, syntype, ♂ (BMNH). 556. M. pedarioides Arrow, 1941, syntype, ♂ (BMNH). 557. M. perlatus (Frey, 1969) comb. nov., ♂ (SMNS). 558. M. popei (Frey, 1969) comb. nov., paratype, ♂ (BMNH). 559. M. rugicollis Moser, 1920, ♂ (BMNH). 560. M. seriegranosus Heller, 1914, lectotype, ♂ (SNSD). 561. M. esau Heller, 1914, ♂ (BMNH). 562. M. speciosus (Frey, 1969) comb. nov., ♂. 563. M. sturnus Arrow, 1941, holotype, ♂ (BMNH). 564. M. suwawa sp. nov., paratype, ♂ (BMNH). 565. M. tarsalis Arrow, 1941, lectotype, ♂ (BMNH). 566. M. trivialis sp. nov., holotype, ♂ (BMNH). 567. M. alesbezdeki sp. nov., holotype, ♂ (NME). 568. M. bintang sp. nov., holotype, ♂ (NME). 569. M. lobaticeps Frey, 1969, ♂ (DTC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122230" httpUri="https://zenodo.org/record/4122230/files/figure.png" pageId="55" pageNumber="56">Fig. 546</figureCitation>
. Aedeagus as in
<figureCitation id="0AA52A659D2B926CFEE5FBE2CA4C2C15" box="[374,538,1116,1143]" captionStart="Figs 617–631" captionStartId="173.[189,241,1892,1918]" captionTargetBox="[439,1134,274,1859]" captionTargetId="figure-71@173.[434,1154,265,1868]" captionTargetPageId="173" captionText="Figs 617–631. Papuan and Wallacean Maechidius Macleay, 1819, aedeagus, different aspects. 617– 619. M. hamatus sp. nov., holotype, ♂ (IECA). 620–622. M. hirtipes Arrow, 1941, lectotype, ♂ (BMNH). 623–625. M. humeralis Heller, 1914, lectotype, ♂ (SNSD). 626–628. M. kazantsevi sp. nov., holotype, ♂ (BMNH). 629–631. M. konjo sp. nov., holotype, ♂ (NMPC). Not to scale." figureDoi="http://doi.org/10.5281/zenodo.4122240" httpUri="https://zenodo.org/record/4122240/files/figure.png" pageId="55" pageNumber="56">Figs 623–625</figureCitation>
.
</paragraph>
<paragraph id="922136E09D2B926CFF2EFB20C9E62CDA" blockId="55.[189,432,1182,1208]" box="[189,432,1182,1208]" pageId="55" pageNumber="56">
<heading id="C969818C9D2B926CFF2EFB20C9E62CDA" bold="true" box="[189,432,1182,1208]" fontSize="11" level="3" pageId="55" pageNumber="56" reason="3">
<emphasis id="A0EAEAF29D2B926CFF2EFB20C9E62CDA" bold="true" box="[189,432,1182,1208]" pageId="55" pageNumber="56">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="922136E09D2B926CFF2EFB73C9002D68" blockId="55.[189,1398,1229,1290]" pageId="55" pageNumber="56">Lamellae of antennal club longer in male, labroclypeus stronger emarginate in female (in dorsal view) than in male.</paragraph>
</subSubSection>
</treatment>
</document>
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