156 lines
20 KiB
XML
156 lines
20 KiB
XML
<document id="50BDE45A95CB735D2A53D0E509304F05" ID-CLB-Dataset="29478" ID-DOI="10.11646/zootaxa.4471.2.4" ID-GBIF-Dataset="05d2a4ba-f50f-4645-bbb3-05cd83dcbd7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1439724" ID-ZooBank="691AD8B9-27BB-40E6-A3D8-C3D17DA38B0B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538364688024" checkinUser="plazi" docAuthor="Lyle, Robin & Haddad, Charles R." docDate="2018" docId="0772878D041BFFE1FF45F8A0FB9A6E55" docLanguage="en" docName="zootaxa.4471.2.4.pdf" docOrigin="Zootaxa 4471 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Jocquestus harrisi Lyle & Haddad 2018, sp. nov." docType="treatment" docUuid="4F864F6A-8177-478E-BA13-F84566471A96" docUuidSource="ZooBank" docVersion="8" lastPageNumber="320" masterDocId="FB4BFFF50411FFEAFFD2FFBAFFFE6A45" masterDocTitle="Jocquestus, a new genus of trachelid sac spiders from the Afrotropical Region (Arachnida: Araneae)" masterLastPageNumber="333" masterPageNumber="309" pageNumber="319" updateTime="1698558233409" updateUser="ExternalLinkService">
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<mods:titleInfo id="E4D2B47925E1D9165A3ADABC0BF3B956">
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<mods:title id="33BE506BEF0809D05031664B592F7C04">Jocquestus, a new genus of trachelid sac spiders from the Afrotropical Region (Arachnida: Araneae)</mods:title>
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<mods:namePart id="1C33201C4BFD78A2C891683B64D465DC">Lyle, Robin</mods:namePart>
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<mods:namePart id="4F774FA3170691BA162ECC2884472A49">Haddad, Charles R.</mods:namePart>
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<mods:typeOfResource id="9504EAEDA6CF11B1A13E37DCBBF63DDA">text</mods:typeOfResource>
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<mods:title id="6B43017440C97A279FF2F8B62309E76C">Zootaxa</mods:title>
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<mods:part id="FB2CB14186AE580CB3D8E090EE95CF08">
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<mods:date id="293FB816DB481E8EBDE27C5F13102F0D">2018</mods:date>
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<mods:number id="D9AC7BA5B5659B0814CE61DEE82DF9B5">2018-09-05</mods:number>
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<mods:number id="B5EAB00E72DC58B8740707D520EDCAA2">4471</mods:number>
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<mods:identifier id="BDC1E118DC240C2BB5E01CDCCE8CCEBC" type="DOI">10.11646/zootaxa.4471.2.4</mods:identifier>
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<treatment id="0772878D041BFFE1FF45F8A0FB9A6E55" ID-DOI="http://doi.org/10.5281/zenodo.5949509" ID-GBIF-Taxon="148415702" ID-Zenodo-Dep="5949509" LSID="urn:lsid:zoobank.org:act:4F864F6A-8177-478E-BA13-F84566471A96" httpUri="http://treatment.plazi.org/id/0772878D041BFFE1FF45F8A0FB9A6E55" lastPageId="11" lastPageNumber="320" pageId="10" pageNumber="319">
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<subSubSection id="C7C16510041BFFE0FF45F8A0FC4A6D10" pageId="10" pageNumber="319" type="nomenclature">
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<paragraph id="8F64369B041BFFE0FF45F8A0FE276D71" blockId="10.[151,948,1818,1914]" box="[151,473,1818,1844]" pageId="10" pageNumber="319">
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<heading id="D42C81F7041BFFE0FF45F8A0FE276D71" bold="true" box="[151,473,1818,1844]" fontSize="11" level="1" pageId="10" pageNumber="319" reason="1">
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<emphasis id="BDAFEA89041BFFE0FF45F8A0FE276D71" bold="true" box="[151,473,1818,1844]" pageId="10" pageNumber="319">
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<taxonomicName id="48DB4D18041BFFE0FF45F8A0FE886D71" authority="Lyle & Haddad, 2018" authorityName="Lyle & Haddad" authorityYear="2018" box="[151,374,1818,1844]" class="Arachnida" family="Corinnidae" genus="Jocquestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="319" phylum="Arthropoda" rank="species" species="harrisi" status="sp. nov.">
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<emphasis id="BDAFEA89041BFFE0FF45F8A0FE886D71" bold="true" box="[151,374,1818,1844]" italics="true" pageId="10" pageNumber="319">Jocquestus harrisi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A69C57F2041BFFE0FEAEF8A0FE276D71" box="[380,473,1818,1844]" pageId="10" pageNumber="319" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8F64369B041BFFE0FF45F886FC4A6D10" blockId="10.[151,948,1818,1914]" box="[151,948,1852,1877]" pageId="10" pageNumber="319">urn:lsid:zoobank.org:act:4F864F6A-8177-478E-BA13-F84566471A96</paragraph>
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</subSubSection>
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<subSubSection id="C7C16510041BFFE0FF45F8DBFEC06D3F" box="[151,318,1889,1914]" pageId="10" pageNumber="319" type="description">
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<paragraph id="8F64369B041BFFE0FF45F8DBFEC06D3F" blockId="10.[151,948,1818,1914]" box="[151,318,1889,1914]" pageId="10" pageNumber="319">
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<figureCitation id="17E02A1E041BFFE0FF45F8DBFF126D3C" box="[151,236,1889,1914]" captionStart="FIGURES 9–20" captionStartId="3.[151,264,1641,1663]" captionTargetBox="[150,1436,301,1619]" captionTargetId="figure@3.[150,1436,301,1619]" captionTargetPageId="3" captionText="FIGURES 9–20. Digital microscope photographs of the dorsal habitus of Jocquestus gen. nov. species. 9. J. capensis sp. nov., male; 10. Same, female; 11. J. griswoldi sp. nov., male; 12. J. harrisi sp. nov., female; 13. J. incurvus sp. nov., male; 14. Same, female; 15. J. obliquus sp. nov., male; 16. Same, female; 17. J. roeweri (Lawrence, 1938) comb. nov., male; 18. Same, female; 19. J. schenkeli (Lessert, 1923) comb. nov., male; 20. Same, female." httpUri="https://zenodo.org/record/1439728/files/figure.png" pageId="10" pageNumber="319">Figs 12</figureCitation>
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,
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<figureCitation id="17E02A1E041BFFE0FF2AF8DBFEC06D3F" box="[248,318,1889,1914]" captionStart="FIGURES 47–51" captionStartId="10.[151,264,1690,1712]" captionTargetBox="[155,1429,201,1659]" captionTargetId="figure@10.[150,1436,195,1668]" captionTargetPageId="10" captionText="FIGURES 47–51. Jocquestus griswoldi sp. nov. male (47–49) and J. harrisi sp. nov. female (50, 51): 47. Schematic representation of cusp arrangement on legs I and II; 48. Left palp, ventral view; 49. Same, retrolateral view; 50. Epigyne, ventral view; 51. Vulva, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1439738/files/figure.png" pageId="10" pageNumber="319">50, 51</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C7C16510041BFFE0FF45F813FBC66D87" box="[151,1080,1961,1986]" pageId="10" pageNumber="319" type="etymology">
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<paragraph id="8F64369B041BFFE0FF45F813FBC66D87" blockId="10.[151,1436,1961,2021]" box="[151,1080,1961,1986]" pageId="10" pageNumber="319">
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<emphasis id="BDAFEA89041BFFE0FF45F813FEE06D87" bold="true" box="[151,286,1961,1986]" pageId="10" pageNumber="319">Etymology.</emphasis>
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This species is named after Rupert Harris, who collected the
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<typeStatus id="50608839041BFFE0FC1CF813FBCA6D87" box="[974,1076,1961,1986]" pageId="10" pageNumber="319" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C7C16510041BFFE1FF15F876FEF96ABD" lastPageId="11" lastPageNumber="320" pageId="10" pageNumber="319" type="diagnosis">
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<paragraph id="8F64369B041BFFE1FF15F876FEF96ABD" blockId="10.[151,1436,1961,2021]" lastBlockId="11.[151,1437,151,1040]" lastPageId="11" lastPageNumber="320" pageId="10" pageNumber="319">
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<emphasis id="BDAFEA89041BFFE0FF15F876FEBF6DA0" bold="true" box="[199,321,1996,2021]" pageId="10" pageNumber="319">Diagnosis.</emphasis>
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A distinctive species that can be recognised by a combination of the recurved, C-shaped epigynal ridges and the narrow ST II (
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<figureCitation id="17E02A1E041AFFE1FE21FF2DFDB26AF5" box="[499,588,151,176]" captionStart="FIGURES 47–51" captionStartId="10.[151,264,1690,1712]" captionTargetBox="[155,1429,201,1659]" captionTargetId="figure@10.[150,1436,195,1668]" captionTargetPageId="10" captionText="FIGURES 47–51. Jocquestus griswoldi sp. nov. male (47–49) and J. harrisi sp. nov. female (50, 51): 47. Schematic representation of cusp arrangement on legs I and II; 48. Left palp, ventral view; 49. Same, retrolateral view; 50. Epigyne, ventral view; 51. Vulva, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1439738/files/figure.png" pageId="11" pageNumber="320">Fig. 50</figureCitation>
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). All other species in the genus have broad ST II and obliquely aligned epigynal ridges, with the exception of
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<taxonomicName id="48DB4D18041AFFE1FD96FF07FD416A91" authority="Lyle & Haddad, 2018" authorityName="Lyle & Haddad" authorityYear="2018" box="[580,703,189,212]" class="Arachnida" family="Corinnidae" genus="Jocquestus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="320" phylum="Arthropoda" rank="species" species="incurvus" status="sp. nov.">
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<emphasis id="BDAFEA89041AFFE1FD96FF07FD416A91" box="[580,703,189,212]" italics="true" pageId="11" pageNumber="320">J. incurvus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A69C57F2041AFFE1FD17FF06FCE06A90" box="[709,798,188,213]" pageId="11" pageNumber="320" rank="species">
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<emphasis id="BDAFEA89041AFFE1FD17FF06FCE06A90" bold="true" box="[709,798,188,213]" pageId="11" pageNumber="320">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, which also has recurved epigynal ridges (
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<figureCitation id="17E02A1E041AFFE1FB29FF06FAB06A90" box="[1275,1358,188,213]" captionStart="FIGURES 52–55" captionStartId="12.[151,264,1954,1976]" captionTargetBox="[162,1419,205,1928]" captionTargetId="figure@12.[159,1429,193,1941]" captionTargetPageId="12" captionText="FIGURES 52–55. Jocquestus incurvus sp. nov. male (52–54) and female (55): 52. Schematic representation of cusp arrangement on legs I and II; 53. Left palp, ventral view; 54. Same, retrolateral view; 55. Epigyne, ventral view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1439740/files/figure.png" pageId="11" pageNumber="320">Fig. 55</figureCitation>
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). Male unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C7C16510041AFFE1FF15FEBEFD0C69C5" pageId="11" pageNumber="320" type="description">
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<paragraph id="8F64369B041AFFE1FF15FEBEFE3E6B20" blockId="11.[151,1437,151,1040]" pageId="11" pageNumber="320">
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<emphasis id="BDAFEA89041AFFE1FF15FEBEFD216B58" bold="true" box="[199,735,260,285]" pageId="11" pageNumber="320">
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Female (
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<typeStatus id="50608839041AFFE1FEE2FEBEFE646B58" box="[304,410,260,285]" pageId="11" pageNumber="320" type="holotype">holotype</typeStatus>
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, Westphalia, NCA 95/165)
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</emphasis>
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. Measurements: CL 1.40, CW 1.23, AL 1.88, AW 1.40, TL 3.28, FL 0.08, SL 0.83, SW 0.73, AME–AME 0.05, AME–ALE 0.05, ALE–ALE 0.30, PME–PME 0.10, PME– PLE 0.10, PLE–PLE 0.45.
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</paragraph>
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<paragraph id="8F64369B041AFFE1FF15FED5FEE06B95" blockId="11.[151,1437,151,1040]" pageId="11" pageNumber="320">Length of leg segments (sequence from femur to tarsus, and total): I 1.00 + 0.45 + 0.55 + 0.58 + 0.40 = 2.98; II 0.90 + 0.40 + 0.65 + 0.58 + 0.35 = 2.88; III 0.65 + 0.38 + 0.40 + 0.50 + 0.25 = 2.18; IV 0.85 + 0.38 + 0.65 + 0.50 + 0.23 = 2.61.</paragraph>
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<paragraph id="8F64369B041AFFE1FF15FE66FD0C69C5" blockId="11.[151,1437,151,1040]" pageId="11" pageNumber="320">
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Carapace reddish-brown (
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<figureCitation id="17E02A1E041AFFE1FE23FE66FDB76BB1" box="[497,585,476,501]" captionStart="FIGURES 9–20" captionStartId="3.[151,264,1641,1663]" captionTargetBox="[150,1436,301,1619]" captionTargetId="figure@3.[150,1436,301,1619]" captionTargetPageId="3" captionText="FIGURES 9–20. Digital microscope photographs of the dorsal habitus of Jocquestus gen. nov. species. 9. J. capensis sp. nov., male; 10. Same, female; 11. J. griswoldi sp. nov., male; 12. J. harrisi sp. nov., female; 13. J. incurvus sp. nov., male; 14. Same, female; 15. J. obliquus sp. nov., male; 16. Same, female; 17. J. roeweri (Lawrence, 1938) comb. nov., male; 18. Same, female; 19. J. schenkeli (Lessert, 1923) comb. nov., male; 20. Same, female." httpUri="https://zenodo.org/record/1439728/files/figure.png" pageId="11" pageNumber="320">Fig. 12</figureCitation>
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); surface finely granulate, almost smooth, covered with short fine setae; fovea distinct, at two thirds CL; ocular region brown; clypeus height approximately equal to AME diameter; ALE slightly larger than AME; AME separated by distance equal to 0.7 times their diameter; AME separated from ALE by distance equal to 0.4 AME diameter; PLE slightly larger than PME; PME separated by distance equal to twice their diameter; PME separated from PLE by distance equal to twice PME diameter. Chelicerae reddish-brown, anterior surface covered with scattered long fine setae; one distal promarginal tooth present; two retromarginal teeth, distal tooth largest. Sternum brown, darker towards borders; surface smooth, covered with scattered short fine setae. Abdomen pale yellow with grey mottling dorsally; two pairs of pale brown sigilla, one pair anterior to midpoint, second pair darker, posterior to midpoint (
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<figureCitation id="17E02A1E041AFFE1FD32FD46FCCA6951" box="[736,820,764,789]" captionStart="FIGURES 9–20" captionStartId="3.[151,264,1641,1663]" captionTargetBox="[150,1436,301,1619]" captionTargetId="figure@3.[150,1436,301,1619]" captionTargetPageId="3" captionText="FIGURES 9–20. Digital microscope photographs of the dorsal habitus of Jocquestus gen. nov. species. 9. J. capensis sp. nov., male; 10. Same, female; 11. J. griswoldi sp. nov., male; 12. J. harrisi sp. nov., female; 13. J. incurvus sp. nov., male; 14. Same, female; 15. J. obliquus sp. nov., male; 16. Same, female; 17. J. roeweri (Lawrence, 1938) comb. nov., male; 18. Same, female; 19. J. schenkeli (Lessert, 1923) comb. nov., male; 20. Same, female." httpUri="https://zenodo.org/record/1439728/files/figure.png" pageId="11" pageNumber="320">Fig. 12</figureCitation>
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). Legs I to IV uniform pale brown, anterior legs darker than posteriors; leg I with granulate texture, other legs smooth; regular leg spines, cusps absent. Epigynal ridges Cshaped, recurved, with lateral CO; ST II narrow, twisted, close to midline of epigyne anteriorly, with narrow ducts leading to oval ST I near epigastric fold (
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<figureCitation id="17E02A1E041AFFE1FDB1FCDDFD1B69C5" box="[611,741,871,896]" captionStart="FIGURES 47–51" captionStartId="10.[151,264,1690,1712]" captionTargetBox="[155,1429,201,1659]" captionTargetId="figure@10.[150,1436,195,1668]" captionTargetPageId="10" captionText="FIGURES 47–51. Jocquestus griswoldi sp. nov. male (47–49) and J. harrisi sp. nov. female (50, 51): 47. Schematic representation of cusp arrangement on legs I and II; 48. Left palp, ventral view; 49. Same, retrolateral view; 50. Epigyne, ventral view; 51. Vulva, dorsal view. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/1439738/files/figure.png" pageId="11" pageNumber="320">Figs 50, 51</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C7C16510041AFFE1FF15FC36FE44698D" pageId="11" pageNumber="320" type="materials_examined">
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<paragraph id="8F64369B041AFFE1FF15FC36FE44698D" blockId="11.[151,1437,151,1040]" pageId="11" pageNumber="320">
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<emphasis id="BDAFEA89041AFFE1FF15FC36FE8469E0" bold="true" box="[199,378,908,933]" pageId="11" pageNumber="320">
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||
<typeStatus id="50608839041AFFE1FF15FC36FEFC69E0" box="[199,258,908,933]" pageId="11" pageNumber="320">Type</typeStatus>
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material.
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||
</emphasis>
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||
<typeStatus id="50608839041AFFE1FE55FC36FE0A69E0" box="[391,500,908,933]" pageId="11" pageNumber="320" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="99DDFD12041AFFE1FDD6FC37FDE569E1" box="[516,539,909,932]" pageId="11" pageNumber="320" type="female">♀</specimenCount>
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||
:
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<collectingCountry id="F7CC760B041AFFE1FDFFFC36FCFC69E1" box="[557,770,908,932]" name="South Africa" pageId="11" pageNumber="320">
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||
<collectingRegion id="4D1FF879041AFFE1FDFFFC36FD7469E1" box="[557,650,908,932]" country="Tanzania" name="Kusini Unguja" pageId="11" pageNumber="320">SOUTH</collectingRegion>
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AFRICA
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</collectingCountry>
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:
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<collectingRegion id="4D1FF879041AFFE1FCC0FC37FC8869E1" box="[786,886,909,932]" country="South Africa" name="Limpopo" pageId="11" pageNumber="320">
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<emphasis id="BDAFEA89041AFFE1FCC0FC37FC8869E1" box="[786,886,909,932]" italics="true" pageId="11" pageNumber="320">Limpopo</emphasis>
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</collectingRegion>
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: Westphalia,
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,
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<geoCoordinate id="EAEF505C041AFFE1FB57FC36FB1E69E1" box="[1157,1248,908,932]" direction="east" orientation="longitude" pageId="11" pageNumber="320" precision="925" value="29.166666">29°10'E</geoCoordinate>
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, leg. R. Harris,
|
||
<date id="FB65105B041AFFE1FF45FC0AFEF7698D" box="[151,265,944,968]" pageId="11" pageNumber="320" value="1983-05-27">27.V.1983</date>
|
||
(NCA 95/165).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C7C16510041AFFE1FF15FC6EFB9A6E55" pageId="11" pageNumber="320" type="distribution">
|
||
<paragraph id="8F64369B041AFFE1FF15FC6EFCA269A9" blockId="11.[151,1437,151,1040]" box="[199,860,980,1005]" pageId="11" pageNumber="320">
|
||
<emphasis id="BDAFEA89041AFFE1FF15FC6EFE9E69A8" bold="true" box="[199,352,980,1005]" pageId="11" pageNumber="320">Distribution.</emphasis>
|
||
Only known from the
|
||
<typeStatus id="50608839041AFFE1FDB2FC6FFD6E69A8" box="[608,656,981,1005]" pageId="11" pageNumber="320">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="17E02A1E041AFFE1FD28FC6EFCB169A9" box="[762,847,980,1005]" captionStart="FIGURE 70" captionStartId="21.[151,250,1887,1909]" captionTargetBox="[184,1406,553,1877]" captionTargetId="figure@21.[184,1406,553,1877]" captionTargetPageId="21" captionText="FIGURE 70. Distribution of the genus Jocquestus gen. nov. in the Afrotropical Region: J. capensis sp. nov. (black circles), J. griswoldi sp. nov. male (yellow circle), J. harrisi sp. nov. (black square), J. incurvus sp. nov. (yellow squares), J. obliquus sp. nov. (black star), J. roeweri (Lawrence, 1938) comb. nov. (yellow stars) and J. schenkeli (Lessert, 1923) comb. nov. (black triangles)." httpUri="https://zenodo.org/record/1439748/files/figure.png" pageId="11" pageNumber="320">Fig. 70</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8F64369B041AFFE1FF15FC4DFB9A6E55" blockId="11.[151,1437,151,1040]" box="[199,1124,1015,1040]" pageId="11" pageNumber="320">
|
||
<emphasis id="BDAFEA89041AFFE1FF15FC4DFE446E55" bold="true" box="[199,442,1015,1040]" pageId="11" pageNumber="320">Habitat and biology.</emphasis>
|
||
Collected from the savanna biome of northern South Africa.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |