312 lines
24 KiB
XML
312 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.679.13472" ID-GBIF-Dataset="dc2dd8b3-a504-463e-a832-f1e56f140b66" ID-PMC="PMC5523391" ID-Pensoft-Pub="1313-2970-679-1" ID-PubMed="28769703" ID-ZBK="8B13B2C09D7A473090579AD19400F530" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-679-1" ModsDocOrigin="ZooKeys 679" ModsDocTitle="Two new species of Fluminicola (Caenogastropoda, Lithoglyphidae) from southwest Oregon, USA, and a range extension for F.multifarius" checkinTime="1502744660791" checkinUser="pensoft" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Hubbart, Niko" docDate="2017" docId="95414B6DC719B9CC0B353788B00AEDD4" docLanguage="en" docName="ZooKeys 679: 1-20" docOrigin="ZooKeys 679" docSource="http://dx.doi.org/10.3897/zookeys.679.13472" docTitle="Fluminicola umpquaensis Hershler, Liu & Hubbart, sp. n." docType="treatment" docUuid="7EEF6D2F-5CAB-4CD2-A41C-E1EB8D556EBD" docUuidSource="ZooBank" docVersion="5" lastPageNumber="13" masterDocId="FC02FFB3FF96AB5E19083432FFC1FFD2" masterDocTitle="Two new species of Fluminicola (Caenogastropoda, Lithoglyphidae) from southwest Oregon, USA, and a range extension for F. multifarius" masterLastPageNumber="20" masterPageNumber="1" pageNumber="11" updateTime="1668164527539" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of Fluminicola (Caenogastropoda, Lithoglyphidae) from southwest Oregon, USA, and a range extension for F. multifarius</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hershler, Robert</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liu, Hsiu-Ping</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hubbart, Niko</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>679</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>20</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.679.13472</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.679.13472</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-679-1</mods:identifier>
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<mods:identifier type="ZBK">8B13B2C09D7A473090579AD19400F530</mods:identifier>
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<mods:identifier type="ZooBank">8B13B2C09D7A473090579AD19400F530</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="132649546" LSID="urn:lsid:zoobank.org:act:7EEF6D2F-5CAB-4CD2-A41C-E1EB8D556EBD" httpUri="http://treatment.plazi.org/id/95414B6DC719B9CC0B353788B00AEDD4" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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<subSubSection pageId="10" pageNumber="11" type="nomenclature">
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<paragraph pageId="10" pageNumber="11">
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<taxonomicName LSID="http://zoobank.org/7EEF6D2F-5CAB-4CD2-A41C-E1EB8D556EBD" authority="Hershler, Liu & Hubbart" class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fluminicola umpquaensis" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="umpquaensis">Fluminicola umpquaensis Hershler, Liu & Hubbart</taxonomicName>
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<taxonomicNameLabel pageId="10" pageNumber="11">sp. n.</taxonomicNameLabel>
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Figs 4
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<normalizedToken originalValue="H–I">H-I</normalizedToken>
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, 6
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="reference_group">
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<paragraph pageId="10" pageNumber="11">
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<taxonomicName class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fluminicola virens" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="virens">Fluminicola virens</taxonomicName>
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.-
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<bibRefCitation author="Pilsbry, HA" journalOrPublisher="Nautilus" pageId="14" pageNumber="15" pagination="121 - 127" title="Catalogue of the Amnicolidae of the western United States." volume="12" year="1899">Pilsbry 1899</bibRefCitation>
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: 122-123 (in part).
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<bibRefCitation author="Henderson, J" journalOrPublisher="University of Colorado Studies" pageId="13" pageNumber="14" pagination="47 - 190" title="Non-marine Mollusca of Oregon and Washington." volume="17" year="1929">Henderson 1929</bibRefCitation>
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: 169 (in part).
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<bibRefCitation author="Burch, JB" journalOrPublisher="Walkerana" pageId="13" pageNumber="14" pagination="81 - 215" title="North American freshwater snails. Species list, ranges and illustrations." volume="3" year="1980">Burch and Tottenham 1980</bibRefCitation>
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: 102 (in part).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="types">
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<paragraph pageId="10" pageNumber="11">Types.</paragraph>
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<paragraph pageId="10" pageNumber="11">
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Holotype, USNM 1144535, Umpqua River at Bunch Bar Access (County), west of OR38 (Umpqua Highway), Douglas County, Oregon,
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.6462">43.6462°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.667">123.6670°W</geoCoordinate>
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, 9/27/1998, Terrence J. Frest and Edward J. Johannes. Paratypes, USNM 1422224 (a large series of dry shells and alcohol-preserved specimens), from same lot.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph pageId="10" pageNumber="11">Referred material.</paragraph>
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<paragraph lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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OREGON. Jackson County. USNM 1296909, Umpqua River at Bunch Bar Access, USNM 1296910, Umpqua River at Umpqua County Landing upriver of Sutherlin-Umpqua Road Bridge (Douglas County 9) on east side, north of the mouth of Calapooya Creek, opposite of the town of Umpqua, RM 102.7 (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.3661">43.3661°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.4677">123.4677°W</geoCoordinate>
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), USNM 1296911, east side of Umpqua River at Cleveland Rapids, Cleveland Rapids Park, opposite and northeast of Cleveland (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.2966">43.2966°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.4705">123.4705°W</geoCoordinate>
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), USNM 1144361, USNM 126912, east side of North Umpqua River near boat ramp on west side of Whistlers Bend County Park (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.3101">43.3101°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.2168">123.2168°W</geoCoordinate>
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), USNM 1297138, bedrock shelf and bar on west side of North Umpqua River, ca. 0.48 km southwest of mouth of Swamp Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.3015">43.3015°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-122.8692">122.8692°W</geoCoordinate>
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), USNM 1297139, Myrtle Creek just upstream of OR99 (Myrtle Creek Highway) bridge (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.0237">43.0237°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.289">123.2890°W</geoCoordinate>
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), USNM 1297140, Elk Creek just below confluence of Drew Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.8907">42.8907°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-122.9227">122.9227°W</geoCoordinate>
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), USNM 1144544, USNM 1297141, Elk Creek just south of bridge of Callahan Creek Road (FS3230), west and below Tiller Trail Highway (Douglas Co. 1, OR227) (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.8973">42.8973°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-122.9308">122.9308°W</geoCoordinate>
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), USNM 1144751, Pass Creek at site of Comstock in Pass Creek County Park, west side of I-5 off exit 164 (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.7186">43.7186°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.2076">123.2076°W</geoCoordinate>
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), USNM 1144360, USNM 1144656, south side of North Umpqua River at John P. Amacher Park, under and down river of railroad bridge west of I-5 and Winchester (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.2796">43.2796°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.3557">123.3557°W</geoCoordinate>
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), USNM 1144726, west side of North Umpqua River at and 0.32 km upstream of Chris Hestness Landing (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.2843">43.2843°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.1674">123.1674°W</geoCoordinate>
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), USNM 1144730, Umpqua River at Umpqua County Landing upriver of Sutherlin-Umpqua Road Bridge (Douglas County 9) on east side, north of the mouth of Calapooya Creek, opposite of Umpqua (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.3643">43.3643°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.4665">123.4665°W</geoCoordinate>
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), USNM
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<pageBreakToken pageId="11" pageNumber="12" start="start">1144736</pageBreakToken>
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, east side of Umpqua River at Cleveland Rapids, Cleveland Rapids Park, opposite and northeast of Cleveland (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.2972">43.2972°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.4686">123.4686°W</geoCoordinate>
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), USNM 1144748, Myrtle Creek just upstream of OR99 (Myrtle Creek Highway) bridge (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.022">43.0220°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.2871">123.2871°W</geoCoordinate>
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), USNM 1144371, USNM 1144948, west side of South Umpqua River, east of I-5 and north of OR99 (Myrtle Creek Highway) Bridge, west of town of Myrtle Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.0238">43.0238°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.2954">123.2954°W</geoCoordinate>
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), USNM 1144372, South Myrtle Creek, ca. 2.82 km from junction with North Myrtle Creek, below Days Creek Road (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.0176">43.0176°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.2666">123.2666°W</geoCoordinate>
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), USNM 1144473, north side of South Umpqua River just upstream (east) of mouth of Dumont Creek at Dumont Creek Campground (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.0337">43.0337°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-122.801">122.8010°W</geoCoordinate>
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), USNM 1144545, Umpqua River on south side, Scottsburg County Park boat ramp (upstream side), southwest of Scottsburg (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.648">43.6480°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.8377">123.8377°W</geoCoordinate>
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), USNM 1144546, USNM 1144995, Cow Creek at mouth of Salt Creek, west of Cow Creek Road (Douglas Co. 39), east of Byers (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="42.924">42.9240°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.4897">123.4897°W</geoCoordinate>
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), BellMNH uncat., North Umpqua River at fish traps above Ideyld (
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<geoCoordinate direction="north" orientation="latitude" precision="5" value="43.322">43.3220°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-123.0248">123.0248°W</geoCoordinate>
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), BellMNH uncat., North Umpqua River, Glide (
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="43.302">43.302°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-123.101">123.101°W</geoCoordinate>
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), BellMNH uncat., forks of Umpqua River below Roseburg, BellMNH uncat., North Umpqua River at Winchester (
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="43.282">43.282°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-123.355">123.355°W</geoCoordinate>
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), BellMNH uncat., North Umpqua River near Glide, BellMNH uncat., South Umpqua River near Canyonsville, BellMNH uncat., Umpqua River, Basket Point, BellMNH uncat., Umpqua River, Elkton (
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<geoCoordinate direction="north" orientation="latitude" precision="55" value="43.635">43.635°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="55" value="-123.57">123.570°W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="diagnosis">
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<paragraph pageId="11" pageNumber="12">Diagnosis.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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A large
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<taxonomicName class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fluminicola" order="Littorinimorpha" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Fluminicola</taxonomicName>
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(maximum shell height, 9.5 mm) having a conical shell with eroded spire and a broad, little tapered penis. Differs from closely related
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<taxonomicName lsidName="F. gustafsoni" pageId="11" pageNumber="12" rank="species" species="gustafsoni">F. gustafsoni</taxonomicName>
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and
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<taxonomicName lsidName="F. virens" pageId="11" pageNumber="12" rank="species" species="virens">F. virens</taxonomicName>
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in its reniform-shaped bursa copulatrix and in its mtDNA sequences. Further differentiated from
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<taxonomicName lsidName="F. gustafsoni" pageId="11" pageNumber="12" rank="species" species="gustafsoni">F. gustafsoni</taxonomicName>
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by its more elongate shell, longer outer wing of the lateral radular teeth, and smaller seminal receptacle; and from
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<taxonomicName lsidName="F. virens" pageId="11" pageNumber="12" rank="species" species="virens">F. virens</taxonomicName>
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by its more convex shell whorls and longer bursal duct.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
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<paragraph pageId="11" pageNumber="12">Description.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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Shell (Fig. 6
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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) conical, spire usually eroded, whorls>3.5. Teleoconch whorls medium convex, narrowly shouldered. Aperture ovate, slightly angled above; inner lip complete, thickened. Outer lip thin, prosocline. Umbilicus absent. Shell white, periostracum tan or olive. Shell measurements and whorl count data are summarized in Table 2.
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</paragraph>
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<caption pageId="11" pageNumber="12">
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<paragraph pageId="11" pageNumber="12">
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Figure 6. Shells, opercula and radula,
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<taxonomicName lsidName="F. umpquaensis" pageId="11" pageNumber="12" rank="species" species="umpquaensis">F. umpquaensis</taxonomicName>
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, sp. n. A Holotype, USNM 1144535 B Shell, USNM 1144736 C Shell, USNM 1144371 D, E Opercula (outer, inner sides), USNM 114535 F Portion of radular ribbon, USNM 1422224 G Central teeth, USNM 1422224 H Lateral teeth, USNM 1422224. Scale bars:
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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, 1.0 mm; F, 100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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;
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<normalizedToken originalValue="G–H">G-H</normalizedToken>
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, 10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption pageId="11" pageNumber="12">
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<paragraph pageId="11" pageNumber="12">
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Table 2. Shell parameters for
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<taxonomicName lsidName="F. umpquaensis" pageId="11" pageNumber="12" rank="species" species="umpquaensis">F. umpquaensis</taxonomicName>
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. Measurements are in mm.
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</paragraph>
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</caption>
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<paragraph pageId="11" pageNumber="12">
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<table pageId="11" pageNumber="12">
|
||
<tr pageId="11" pageNumber="12">
|
||
<th colspan="1" pageId="11" pageNumber="12" rowspan="1">
|
||
<abbrev id="ABBRID0EWOBG" pageId="11" pageNumber="12">SH</abbrev>
|
||
</th>
|
||
<th colspan="1" pageId="11" pageNumber="12" rowspan="1">
|
||
<abbrev id="ABBRID0E5OBG" pageId="11" pageNumber="12">SW</abbrev>
|
||
</th>
|
||
<th colspan="1" pageId="11" pageNumber="12" rowspan="1">
|
||
<abbrev id="ABBRID0EGPBG" pageId="11" pageNumber="12">HBW</abbrev>
|
||
</th>
|
||
<th colspan="1" pageId="11" pageNumber="12" rowspan="1">
|
||
<abbrev id="ABBRID0EOPBG" pageId="11" pageNumber="12">WBW</abbrev>
|
||
</th>
|
||
<th colspan="1" pageId="11" pageNumber="12" rowspan="1">
|
||
<abbrev id="ABBRID0EWPBG" pageId="11" pageNumber="12">AH</abbrev>
|
||
</th>
|
||
<th colspan="1" pageId="11" pageNumber="12" rowspan="1">
|
||
<abbrev id="ABBRID0E5PBG" pageId="11" pageNumber="12">AW</abbrev>
|
||
</th>
|
||
</tr>
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<tr pageId="11" pageNumber="12">
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<td colspan="7" pageId="11" pageNumber="12" rowspan="1">USNM</td>
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||
</tr>
|
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<tr pageId="11" pageNumber="12">
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<td colspan="7" pageId="11" pageNumber="12" rowspan="1">USNMN</td>
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||
</tr>
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</table>
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</paragraph>
|
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<paragraph pageId="11" pageNumber="12">
|
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Operculum (Fig. 6
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<normalizedToken originalValue="D–E">D-E</normalizedToken>
|
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) as for genus; inner side smooth. Radula (Fig. 6
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<normalizedToken originalValue="F–H">F-H</normalizedToken>
|
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) as for genus; dorsal edge of central teeth concave, lateral cusps
|
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<normalizedToken originalValue="four–five">four-five</normalizedToken>
|
||
, basal cusp
|
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<normalizedToken originalValue="two–four">two-four</normalizedToken>
|
||
. Lateral teeth having
|
||
<normalizedToken originalValue="three–six">three-six</normalizedToken>
|
||
cusps on inner side and
|
||
<normalizedToken originalValue="four–six">four-six</normalizedToken>
|
||
cusps on outer side; length of outer wing 180-200% length of cutting edge. Inner marginal teeth with 13-18 cusps, outer marginal teeth with 11-17 cusps. Radula data are from USNM 1422224.
|
||
</paragraph>
|
||
<paragraph lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
Head-foot rather lightly pigmented, cephalic tentacles with central brown longitudinal stripe along length. Pallial roof, visceral coil dark brown, almost black. Penis having dense core of internal black pigment along penial duct. Ctenidial filaments 34-36 (N = 5), lateral surfaces ridged. Glandular oviduct and associated structures shown in Figure 4
|
||
<normalizedToken originalValue="H–I">H-I</normalizedToken>
|
||
. Coiled oviduct vertical or posterior-oblique, proximal arm kinked. Bursa copulatrix medium-sized, reniform, largely overlapped by albumen gland. Bursal duct about twice as long as bursa copulatrix, very narrow. Seminal receptacle small, sac-like, completely overlapped by albumen gland. Albumen gland having short pallial component. Capsule gland slightly shorter than albumen gland, composed of two glandular
|
||
<pageBreakToken pageId="12" pageNumber="13" start="start">zones</pageBreakToken>
|
||
. Genital aperture a small, sub-terminal pore. Penis (Fig. 4J) large, straight, broad, little tapered, distally rounded, deeply folded along most of length. Penial duct near centrally positioned, rather wide, undulating along entire length, opening through small terminal papilla.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="13" type="etymology">
|
||
<paragraph pageId="12" pageNumber="13">Etymology.</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">The species name is an adjectival geographic epithet referring to the distribution of this pebblesnail in the Umpqua River basin.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph pageId="12" pageNumber="13">Distribution.</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">
|
||
<taxonomicName class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fluminicola umpquaensis" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="umpquaensis">Fluminicola umpquaensis</taxonomicName>
|
||
is widely ranging in the Umpqua River basin, and is distributed in riverine habitats as well as springs and streams.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="12" pageNumber="13" type="remarks">
|
||
<paragraph pageId="12" pageNumber="13">Remarks.</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">
|
||
As mentioned above, the smaller of the two divergent
|
||
<taxonomicName class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fluminicola" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="genus">Fluminicola</taxonomicName>
|
||
clades (containing
|
||
<taxonomicName lsidName="F. gustafsoni" pageId="12" pageNumber="13" rank="species" species="gustafsoni">F. gustafsoni</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="F. virens" pageId="12" pageNumber="13" rank="species" species="virens">F. virens</taxonomicName>
|
||
) was previously confined to the Columbia River basin (
|
||
<bibRefCitation author="Hershler, R" journalOrPublisher="Journal of Molluscan Studies" pageId="13" pageNumber="14" pagination="321 - 329" title="Molecular phylogeny of the western North American pebblesnails, genus Fluminicola (Rissooidea: Lithoglyphidae), with description of a new species." url="https://doi.org/10.1093/mollus/eys017" volume="78" year="2012">Hershler and Liu 2012</bibRefCitation>
|
||
).
|
||
<taxonomicName class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Fluminicola umpquaensis" order="Littorinimorpha" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="umpquaensis">Fluminicola umpquaensis</taxonomicName>
|
||
extends the geographic range of this lineage>200 km southward from the lower Columbia River. Eight COI haplotypes and 11 cytB haplotypes were detected in the sequenced specimens of
|
||
<taxonomicName lsidName="F. umpquaensis" pageId="12" pageNumber="13" rank="species" species="umpquaensis">F. umpquaensis</taxonomicName>
|
||
(Suppl. material 2-3, respectively).
|
||
</paragraph>
|
||
<paragraph pageId="12" pageNumber="13">
|
||
Populations identified herein as
|
||
<taxonomicName lsidName="F. umpquaensis" pageId="12" pageNumber="13" rank="species" species="umpquaensis">F. umpquaensis</taxonomicName>
|
||
were referred to as the Jade pebblesnail by
|
||
<bibRefCitation author="Frest, TJ" journalOrPublisher="Deixis Consultants, Seattle, WA" pageId="13" pageNumber="14" title="A baseline mollusk survey of southwestern Oregon, with emphasis on the Rogue and Umpqua River drainages. Final report for Oregon Natural Heritage Program, Portland, OR." year="2000">Frest and Johannes (2000</bibRefCitation>
|
||
:182). We propose that this common name continue to be applied to this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |